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<document id="B884762E1788009385AEBD7208CA7B40" ID-DOI="10.11646/zootaxa.5045.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5532178" ID-ZooBank="B12C62E3-70D0-4989-BB1A-F4A75C492D8F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632753269074" checkinUser="plazi" docAuthor="Ferreira, Luciane Augusto De Azevedo &amp; Anker, Arthur" docDate="2021" docId="03D62C6B770CFFB64EF95BBBFA89A998" docLanguage="en" docName="zootaxa.5045.1.1.pdf" docOrigin="Zootaxa 5045 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5045.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Petrolisthes quadratus Benedict 1901" docType="treatment" docVersion="4" lastPageNumber="114" masterDocId="FFEF54137763FFC74E6E5E39FFAFAE73" masterDocTitle="An annotated and illustrated checklist of the porcelain crabs of Panama (Decapoda: Anomura)" masterLastPageNumber="154" masterPageNumber="1" pageNumber="112" updateTime="1699415883255" updateUser="plazi">
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<mods:title id="8C880016DC2F1837BF3D7113D689D44F">An annotated and illustrated checklist of the porcelain crabs of Panama (Decapoda: Anomura)</mods:title>
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<mods:namePart id="022530EAB3BF8F48294583963EB1D400">Ferreira, Luciane Augusto De Azevedo</mods:namePart>
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<taxonomicName id="4C7FE6FE770CFFA84EF95BBBFDCAABEE" ID-CoL="e7a4e637-6abc-4aa0-9b88-06930d57a92a" authority="Benedict, 1901" authorityName="Benedict" authorityYear="1901" box="[151,613,1410,1437]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="quadratus">
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<emphasis id="B90B416F770CFFA84EF95BBBFE09ABEE" bold="true" box="[151,422,1410,1437]" italics="true" pageId="111" pageNumber="112">Petrolisthes quadratus</emphasis>
<bibRefCitation id="EFEEE08C770CFFA84FC05BBAFDCAABEE" author="Benedict, J. E." box="[430,613,1411,1437]" pageId="111" pageNumber="112" pagination="129 - 148" refId="ref70189" refString="Benedict, J. E. (1901) The anomuran collections made by the &quot; Fish Hawk &quot; Expedition to Porto Rico. Bulletin of the United States Fish Commission, 20 (2), 129 - 148." type="journal article" year="1901">Benedict, 1901</bibRefCitation>
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<paragraph id="8BC09D7D770CFFA84EF95B9EFE91ABB2" blockId="111.[151,613,1410,1473]" box="[151,318,1447,1473]" pageId="111" pageNumber="112">
(
<figureCitation id="134481F8770CFFA84EF15B9EFF52ABB2" box="[159,253,1447,1473]" captionStart="FIGURE 71" captionStartId="114.[151,250,1634,1658]" captionTargetBox="[168,1418,181,1610]" captionTargetId="figure-19@114.[168,1418,181,1610]" captionTargetPageId="114" captionText="FIGURE 71. Petrolisthes quadratus Benedict, 1901:A, B, extra-limital specimen, male (cl cw indet.) from Saint Martin (FLM- NH UF 31974), dorsal (A) and ventral (B) views [photographs by AA]." figureDoi="http://doi.org/10.5281/zenodo.5532388" httpUri="https://zenodo.org/record/5532388/files/figure.png" pageId="111" pageNumber="112">Figs. 71</figureCitation>
,
<figureCitation id="134481F8770CFFA84F675B9EFE99ABB2" box="[265,310,1447,1473]" captionStart="FIGURE 90" captionStartId="140.[151,250,1624,1648]" captionTargetBox="[164,1424,181,1599]" captionTargetId="figure-19@140.[164,1424,181,1601]" captionTargetPageId="140" captionText="FIGURE 90. Some western Atlantic Porcellanidae (all species reported from Panama or Golfo de Urabá at the Panamanian- Colombian border) photographed in situ: A, Neopisosoma angustifrons (Benedict, 1901), Golfo de Urabá, Colombia; B, Neopisosoma neglectum Werding, 1986, Golfo de Urabá, Colombia; C, Pachycheles tuerkayi Werding &amp; Hiller, 2017, Santa Marta, Colombia; D, Petrolisthes jugosus Streets, 1872, Tayrona, Colombia; E, Petrolisthes tonsorius Haig, 1960, blue colour morph, Bonaire; F, Petrolisthes quadratus Benedict, 1901, Bonaire [photographs courtesy of Juan-Felipe Lazarus-Agudelo (A–D) and Marco Faasse (E, F)]." figureDoi="http://doi.org/10.5281/zenodo.5532436" httpUri="https://zenodo.org/record/5532436/files/figure.png" pageId="111" pageNumber="112">90F</figureCitation>
)
</paragraph>
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<subSubSection id="C365CEF6770CFFA84EF95BD2FD3CA832" pageId="111" pageNumber="112" type="reference_group">
<paragraph id="8BC09D7D770CFFA84EF95BD2FD3CA832" blockId="111.[151,1436,1515,1601]" pageId="111" pageNumber="112">
<treatmentCitationGroup id="AB6FBA53770CFFA84EF95BD2FD3CA832" pageId="111" pageNumber="112">
<treatmentCitation id="0ADEBB6C770CFFA84EF95BD2FDFBA870" author="Benedict, J. E." box="[151,596,1515,1539]" page="134" pageId="111" pageNumber="112" year="1901">
<taxonomicName id="4C7FE6FE770CFFA84EF95BD2FDFBA870" ID-CoL="e7a4e637-6abc-4aa0-9b88-06930d57a92a" authority="Benedict, 1901: 134" authorityName="Benedict" authorityPageNumber="134" authorityYear="1901" box="[151,596,1515,1539]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B90B416F770CFFA84EF95BD2FED2A870" box="[151,381,1515,1539]" italics="true" pageId="111" pageNumber="112">Petrolisthes quadratus</emphasis>
<bibRefCitation id="EFEEE08C770CFFA84FEA5BD2FDFBA870" author="Benedict, J. E." box="[388,596,1515,1539]" pageId="111" pageNumber="112" pagination="129 - 148" refId="ref70189" refString="Benedict, J. E. (1901) The anomuran collections made by the &quot; Fish Hawk &quot; Expedition to Porto Rico. Bulletin of the United States Fish Commission, 20 (2), 129 - 148." type="journal article" year="1901">Benedict, 1901: 134</bibRefCitation>
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, pl. 3, fig. 4;
<treatmentCitation id="0ADEBB6C770CFFA84CB05BD2FCDCA870" author="Haig, J." box="[734,883,1515,1539]" page="18" pageId="111" pageNumber="112" year="1956">
<bibRefCitation id="EFEEE08C770CFFA84CB05BD2FCDCA870" author="Haig, J." box="[734,883,1515,1539]" pageId="111" pageNumber="112" pagination="1 - 44" refId="ref72706" refString="Haig, J. (1956) The Galatheidea (Crustacea Anomura) of the Allan Hancock Atlantic Expedition with a review of the Porcellanidae of the western North Atlantic. Allan Hancock Atlantic Expedition Report, 8, 1 - 44." type="journal article" year="1956">Haig 1956: 18</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C770CFFA84D105BD2FB8FA870" author="Haig J." box="[894,1056,1515,1539]" page="179" pageId="111" pageNumber="112" year="1962">
<bibRefCitation id="EFEEE08C770CFFA84D105BD2FB8FA870" author="Haig J." box="[894,1056,1515,1539]" pageId="111" pageNumber="112" pagination="171 - 192" refId="ref72846" refString="Haig J. (1962) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. 79. Porcellanid crabs from eastern and western America. Videnskabelige meddelelser fra Dansk naturhistorisk forening i KjObenhavn, 124, 171 - 192." type="journal article" year="1962">Haig 1962: 179</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C770CFFA84A455BD2FA91A870" author="Chace, F. A. Jr. &amp; Hobbs, H. H. Jr." box="[1067,1342,1515,1539]" page="121" pageId="111" pageNumber="112" year="1969">
<bibRefCitation id="EFEEE08C770CFFA84A455BD2FA91A870" author="Chace, F. A. Jr. &amp; Hobbs, H. H. Jr." box="[1067,1342,1515,1539]" pageId="111" pageNumber="112" pagination="1 - 258" refId="ref70605" refString="Chace, F. A. Jr. &amp; Hobbs, H. H. Jr. (1969) The freshwater and terrestrial decapod crustaceans of the West Indies with special reference to Dominica. United States National Museum Bulletin, 292, i - vi + 1 - 258, 5 pls. https: // doi. org / 10.5479 / si. 03629236.292.1" type="journal article" year="1969">Chace &amp; Hobbs 1969: 121</bibRefCitation>
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, figs. 32, 43a;
<treatmentCitation id="0ADEBB6C770CFFA84E975833FE12A851" author="Werding, B." box="[249,445,1546,1570]" page="204" pageId="111" pageNumber="112" year="1977">
<bibRefCitation id="EFEEE08C770CFFA84E975833FE12A851" author="Werding, B." box="[249,445,1546,1570]" pageId="111" pageNumber="112" pagination="173 - 214" refId="ref77184" refString="Werding, B. (1977) Los porcelanidos (Crustacea: Anomura: Porcellanidae) de la region de Santa Marta, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betin, 9, 173 - 214. https: // doi. org / 10.25268 / bimc. invemar. 1977.9.0.527" type="journal article" year="1977">Werding 1977: 204</bibRefCitation>
</treatmentCitation>
, fig. 23;
<treatmentCitation id="0ADEBB6C770CFFA84C795833FC87A851" author="Rodriguez, I. T. &amp; Hernandez, G. &amp; Felder, D. L." box="[535,808,1546,1570]" page="565" pageId="111" pageNumber="112" year="2005">
<bibRefCitation id="EFEEE08C770CFFA84C795833FC87A851" author="Rodriguez, I. T. &amp; Hernandez, G. &amp; Felder, D. L." box="[535,808,1546,1570]" pageId="111" pageNumber="112" pagination="544 - 582" refId="ref76152" refString="Rodriguez, I. T., Hernandez, G. &amp; Felder, D. L. (2005) Review of the western Atlantic Porcellanidae (Crustacea: Decapoda: Anomura) with new records, systematic observations, and comments on biogeography. Caribbean Journal of Science, 41 (3), 544 - 582." type="journal article" year="2005">
Rodríguez
<emphasis id="B90B416F770CFFA84CE85832FD15A851" box="[646,698,1546,1570]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
2005: 565
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C770CFFA84D5C5833FBBAA851" author="Hiller, A. &amp; Kraus, H. &amp; Almon, M. &amp; Werding, B." box="[818,1045,1546,1570]" page="553" pageId="111" pageNumber="112" year="2006">
<bibRefCitation id="EFEEE08C770CFFA84D5C5833FBBAA851" author="Hiller, A. &amp; Kraus, H. &amp; Almon, M. &amp; Werding, B." box="[818,1045,1546,1570]" pageId="111" pageNumber="112" pagination="547 - 569" refId="ref73577" refString="Hiller, A., Kraus, H., Almon, M. &amp; Werding, B. (2006) The Petrolisthes galathinus complex: species boundaries based on color pattern, morphology and molecules, and evolutionary interrelationships between the complex and other Porcellanidae (Crustacea: Decapoda: Anomura). Molecular Phylogenetics and Evolution, 40, 547 - 569. https: // doi. org / 10.1016 / j. ympev. 2006.03.030" type="journal article" year="2006">
Hiller
<emphasis id="B90B416F770CFFA84D1D5832FC08A851" box="[883,935,1546,1570]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
2006: 553
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C770CFFA84A715833FAECA851" author="Poupin, J. &amp; Lemaitre, R." box="[1055,1347,1546,1570]" page="21" pageId="111" pageNumber="112" year="2014">
<bibRefCitation id="EFEEE08C770CFFA84A715833FAECA851" author="Poupin, J. &amp; Lemaitre, R." box="[1055,1347,1546,1570]" pageId="111" pageNumber="112" pagination="5 - 27" refId="ref75913" refString="Poupin, J. &amp; Lemaitre, R. (2014) Porcellanid crabs from Guadeloupe Island (Crustacea, Decapoda, Anomura), with an updated list of species from the Lesser Antilles. Zoosystema, 36 (1), 5 - 27. https: // doi. org / 10.5252 / z 2014 n 1 a 1" type="journal article" year="2014">Poupin &amp; Lemaitre 2014: 21</bibRefCitation>
</treatmentCitation>
, fig. 4D, E;
<treatmentCitation id="0ADEBB6C770CFFA84E8A5810FE61A832" ID-DOI="http://doi.org/10.5281/zenodo.5633031" ID-Zenodo-Dep="5633031" author="Diez, Y. L. &amp; Lira, C." box="[228,462,1577,1601]" httpUri="http://treatment.plazi.org/id/8E572401673EF5522BC2FF04F457FEB5" page="447" pageId="111" pageNumber="112" year="2017">
<bibRefCitation id="EFEEE08C770CFFA84E8A5810FE61A832" author="Diez, Y. L. &amp; Lira, C." box="[228,462,1577,1601]" pageId="111" pageNumber="112" pagination="441 - 456" refId="ref70973" refString="Diez, Y. L. &amp; Lira, C. (2017) Systematics and biogeography of Cuban porcelain crabs (Decapoda: Anomura: Porcellanidae). Zootaxa, 4216 (5), 441 - 456. https: // doi. org / 10.11646 / zootaxa. 4216.5.2" type="journal article" year="2017">Diez &amp; Lira 2017: 447</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADEBB6C770CFFA84FB65810FD20A832" author="Poupin, J." box="[472,655,1577,1601]" page="151" pageId="111" pageNumber="112" year="2018">
<bibRefCitation id="EFEEE08C770CFFA84FB65810FD20A832" author="Poupin, J." box="[472,655,1577,1601]" pageId="111" pageNumber="112" refId="ref75876" refString="Poupin, J. (2018) Les crustaces decapodes des Petites Antilles: avec de nouvelles observations pour Saint-Martin, la Guadeloupe et la Martinique. Museum National d'Histoire Naturelle, Paris, 264 pp." type="book" year="2018">Poupin 2018: 151</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C365CEF6770CFFA84EF95856FD77A88A" pageId="111" pageNumber="112" type="materials_examined">
<paragraph id="8BC09D7D770CFFA84EF95856FE40A8A0" blockId="111.[151,1436,1647,2005]" pageId="111" pageNumber="112">
<emphasis id="B90B416F770CFFA84EF95856FED3A8FA" bold="true" box="[151,380,1647,1673]" pageId="111" pageNumber="112">Material examined</emphasis>
.
<materialsCitation id="3B179720770CFFA84FE35849FE44A8A0" collectingDate="2012-04-13" collectionCode="FLMNH" collectorName="Sta. &amp; M. Bemis" country="Panama" county="Pinel Island" latitude="18.107416" location="None" longLatPrecision="1" longitude="-63.017277" municipality="French Antilles" pageId="111" pageNumber="112" specimenCode="UF 31974" specimenCount="1" specimenCount-male="1">
<location id="8EA0CBA6770CFFA84FE35849FE65A8F9" LSID="urn:lsid:plazi:treatment:03D62C6B770CFFB64EF95BBBFA89A998:8EA0CBA6770CFFA84FE35849FE65A8F9" box="[397,458,1648,1674]" country="Panama" county="Pinel Island" latitude="18.107416" longLatPrecision="1" longitude="-63.017277" municipality="French Antilles" name="None" pageId="111" pageNumber="112">None</location>
from
<collectingCountry id="F368DDED770CFFA84C7D5849FDC0A8F9" box="[531,623,1648,1674]" name="Panama" pageId="111" pageNumber="112">Panama</collectingCountry>
.
<location id="8EA0CBA6770CFFA84C125849FCA4A8F9" LSID="urn:lsid:plazi:treatment:03D62C6B770CFFB64EF95BBBFA89A998:8EA0CBA6770CFFA84C125849FCA4A8F9" box="[636,779,1648,1674]" country="Panama" county="Pinel Island" latitude="18.107416" longLatPrecision="1" longitude="-63.017277" municipality="French Antilles" name="Extra-limital" pageId="111" pageNumber="112">Extra-limital</location>
material.
<collectingMunicipality id="6BA40707770CFFA84DEF5849FB9BA8F9" box="[897,1076,1648,1674]" pageId="111" pageNumber="112">French Antilles</collectingMunicipality>
:
<specimenCount id="9D7956F4770CFFA84A2F5849FB3DA8F9" box="[1089,1170,1648,1674]" pageId="111" pageNumber="112" type="male">1 male</specimenCount>
, cl cw indet. (
<collectionCode id="ED6E05B8770CFFA84B565849FA33A8F9" box="[1336,1436,1648,1674]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="111" pageNumber="112" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBD93506770CFFA84EF958ACFEA0A8DC" box="[151,271,1685,1711]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="111" pageNumber="112" type="Museum">UF 31974</specimenCode>
), Saint Martin,
<collectorName id="268AF8AB770CFFA84FA658ACFE5CA8DC" box="[456,499,1685,1711]" pageId="111" pageNumber="112">Sta.</collectorName>
STM-024,
<collectingCounty id="62A1E5F1770CFFA84C1258ADFCA8A8DD" box="[636,775,1684,1710]" pageId="111" pageNumber="112">Pinel Island</collectingCounty>
,
<geoCoordinate id="EE4BFBBA770CFFA84D7858ADFC18A8DD" box="[790,951,1684,1710]" degrees="18" direction="north" minutes="06" orientation="latitude" pageId="111" pageNumber="112" precision="1" seconds="26.7" value="18.107416">18°06’26.7”N</geoCoordinate>
,
<geoCoordinate id="EE4BFBBA770CFFA84DA858ADFBC5A8DD" box="[966,1130,1684,1710]" degrees="63" direction="west" minutes="01" orientation="longitude" pageId="111" pageNumber="112" precision="1" seconds="02.2" value="-63.017277">63°01’02.2”W</geoCoordinate>
, intertidal, leg.
<collectorName id="268AF8AB770CFFA84B4D58ACFA37A8DD" box="[1315,1432,1684,1711]" pageId="111" pageNumber="112">M. Bemis</collectorName>
,
<date id="FFC1BBBD770CFFA84EF95880FEBAA8A0" box="[151,277,1721,1747]" pageId="111" pageNumber="112" value="2012-04-13">
<collectingDate id="EF854255770CFFA84EF95880FEBAA8A0" box="[151,277,1721,1747]" pageId="111" pageNumber="112" value="2012-04-13">13.04.2012</collectingDate>
</date>
(fcn BSTM-0492)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC09D7D770CFFA84EA958E7FD77A88A" blockId="111.[151,1436,1647,2005]" box="[199,728,1758,1785]" pageId="111" pageNumber="112">
<emphasis id="B90B416F770CFFA84EA958E7FD95A88B" bold="true" box="[199,570,1758,1784]" pageId="111" pageNumber="112">
Previous records from
<collectingCountry id="F368DDED770CFFA84FB658E7FD95A88B" box="[472,570,1758,1784]" name="Panama" pageId="111" pageNumber="112">Panama</collectingCountry>
</emphasis>
.
<bibRefCitation id="EFEEE08C770CFFA84C2658E7FD7CA88A" author="Haig, J." box="[584,723,1758,1785]" pageId="111" pageNumber="112" pagination="1 - 44" refId="ref72706" refString="Haig, J. (1956) The Galatheidea (Crustacea Anomura) of the Allan Hancock Atlantic Expedition with a review of the Porcellanidae of the western North Atlantic. Allan Hancock Atlantic Expedition Report, 8, 1 - 44." type="journal article" year="1956">Haig (1956)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C365CEF6770CFFA84EA9593AFC97A914" pageId="111" pageNumber="112" type="distribution">
<paragraph id="8BC09D7D770CFFA84EA9593AFC97A914" blockId="111.[151,1436,1647,2005]" pageId="111" pageNumber="112">
<emphasis id="B90B416F770CFFA84EA9593AFEF6A96E" bold="true" box="[199,345,1795,1821]" pageId="111" pageNumber="112">Distribution</emphasis>
. West Atlantic:
<collectingCountry id="F368DDED770CFFA84C7C593AFDE7A96E" box="[530,584,1795,1821]" name="United States of America" pageId="111" pageNumber="112">USA</collectingCountry>
(Florida),
<collectingCountry id="F368DDED770CFFA84CAB593AFCB2A96E" box="[709,797,1795,1821]" name="Mexico" pageId="111" pageNumber="112">Mexico</collectingCountry>
,
<collectingCountry id="F368DDED770CFFA84D44593AFCDBA96E" box="[810,884,1795,1821]" name="Belize" pageId="111" pageNumber="112">Belize</collectingCountry>
,
<collectingCountry id="F368DDED770CFFA84DEF593AFC10A96E" box="[897,959,1795,1821]" name="Cuba" pageId="111" pageNumber="112">Cuba</collectingCountry>
,
<collectingCountry id="F368DDED770CFFA84DA2593AFBF9A96E" box="[972,1110,1795,1821]" name="Puerto Rico" pageId="111" pageNumber="112">Puerto Rico</collectingCountry>
,
<collectingCountry id="F368DDED770CFFA84A0C593AFB64A96E" box="[1122,1227,1795,1821]" name="Bahamas" pageId="111" pageNumber="112">Bahamas</collectingCountry>
, N, E and S Less- er Antilles,
<collectingCountry id="F368DDED770CFFA84F705911FED8A931" box="[286,375,1832,1858]" name="Panama" pageId="111" pageNumber="112">Panama</collectingCountry>
(Bahía de Caledonia),
<collectingCountry id="F368DDED770CFFA84CEC5911FD5FA931" box="[642,752,1832,1858]" name="Colombia" pageId="111" pageNumber="112">Colombia</collectingCountry>
and
<collectingCountry id="F368DDED770CFFA84D455911FC31A931" box="[811,926,1832,1858]" name="Venezuela" pageId="111" pageNumber="112">Venezuela</collectingCountry>
(e.g.
<bibRefCitation id="EFEEE08C770CFFA84D8C5911FB29A931" author="Werding, B." box="[994,1158,1832,1858]" pageId="111" pageNumber="112" pagination="173 - 214" refId="ref77184" refString="Werding, B. (1977) Los porcelanidos (Crustacea: Anomura: Porcellanidae) de la region de Santa Marta, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betin, 9, 173 - 214. https: // doi. org / 10.25268 / bimc. invemar. 1977.9.0.527" type="journal article" year="1977">Werding 1977</bibRefCitation>
;
<bibRefCitation id="EFEEE08C770CFFA84AFA5911FA38A931" author="Rodriguez, I. T. &amp; Hernandez, G. &amp; Felder, D. L." box="[1172,1431,1831,1858]" pageId="111" pageNumber="112" pagination="544 - 582" refId="ref76152" refString="Rodriguez, I. T., Hernandez, G. &amp; Felder, D. L. (2005) Review of the western Atlantic Porcellanidae (Crustacea: Decapoda: Anomura) with new records, systematic observations, and comments on biogeography. Caribbean Journal of Science, 41 (3), 544 - 582." type="journal article" year="2005">
Rodríguez
<emphasis id="B90B416F770CFFA84B7C5911FAEAA931" box="[1298,1349,1832,1858]" italics="true" pageId="111" pageNumber="112">et al</emphasis>
., 2005
</bibRefCitation>
;
<bibRefCitation id="EFEEE08C770CFFA84EF95975FE1DA914" author="Poupin, J. &amp; Lemaitre, R." box="[151,434,1868,1895]" pageId="111" pageNumber="112" pagination="5 - 27" refId="ref75913" refString="Poupin, J. &amp; Lemaitre, R. (2014) Porcellanid crabs from Guadeloupe Island (Crustacea, Decapoda, Anomura), with an updated list of species from the Lesser Antilles. Zoosystema, 36 (1), 5 - 27. https: // doi. org / 10.5252 / z 2014 n 1 a 1" type="journal article" year="2014">Poupin &amp; Lemaitre 2014</bibRefCitation>
;
<bibRefCitation id="EFEEE08C770CFFA84FD35975FD22A914" author="Diez, Y. L. &amp; Lira, C." box="[445,653,1868,1895]" pageId="111" pageNumber="112" pagination="441 - 456" refId="ref70973" refString="Diez, Y. L. &amp; Lira, C. (2017) Systematics and biogeography of Cuban porcelain crabs (Decapoda: Anomura: Porcellanidae). Zootaxa, 4216 (5), 441 - 456. https: // doi. org / 10.11646 / zootaxa. 4216.5.2" type="journal article" year="2017">Diez &amp; Lira, 2017</bibRefCitation>
;
<bibRefCitation id="EFEEE08C770CFFA84CF75975FC82A914" author="Poupin, J." box="[665,813,1868,1895]" pageId="111" pageNumber="112" refId="ref75876" refString="Poupin, J. (2018) Les crustaces decapodes des Petites Antilles: avec de nouvelles observations pour Saint-Martin, la Guadeloupe et la Martinique. Museum National d'Histoire Naturelle, Paris, 264 pp." type="book" year="2018">Poupin 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C365CEF6770CFFA84EA95948FDA9A9A6" pageId="111" pageNumber="112" type="biology_ecology">
<paragraph id="8BC09D7D770CFFA84EA95948FDA9A9A6" blockId="111.[151,1436,1647,2005]" pageId="111" pageNumber="112">
<emphasis id="B90B416F770CFFA84EA95948FE89A9F8" bold="true" box="[199,294,1905,1931]" pageId="111" pageNumber="112">Ecology</emphasis>
. Largely intertidal, known depth range
<quantity id="4C873098770CFFA84C8E5948FC85A9FF" box="[736,810,1905,1932]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="111" pageNumber="112" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</quantity>
; on coral substrate, in tide pools, among rocks and coral rubble, especially common at splash zone on sand near piles of rocks (
<bibRefCitation id="EFEEE08C770CFFA84DCC59AFFB39A9C3" author="Chace, F. A. Jr. &amp; Hobbs, H. H. Jr." box="[930,1174,1942,1968]" pageId="111" pageNumber="112" pagination="1 - 258" refId="ref70605" refString="Chace, F. A. Jr. &amp; Hobbs, H. H. Jr. (1969) The freshwater and terrestrial decapod crustaceans of the West Indies with special reference to Dominica. United States National Museum Bulletin, 292, i - vi + 1 - 258, 5 pls. https: // doi. org / 10.5479 / si. 03629236.292.1" type="journal article" year="1969">Chace &amp; Hobbs 1969</bibRefCitation>
;
<bibRefCitation id="EFEEE08C770CFFA84ACF59AFFAEDA9C3" author="Werding, B." box="[1185,1346,1942,1968]" pageId="111" pageNumber="112" pagination="173 - 214" refId="ref77184" refString="Werding, B. (1977) Los porcelanidos (Crustacea: Anomura: Porcellanidae) de la region de Santa Marta, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betin, 9, 173 - 214. https: // doi. org / 10.25268 / bimc. invemar. 1977.9.0.527" type="journal article" year="1977">Werding 1977</bibRefCitation>
;
<bibRefCitation id="EFEEE08C770CFFA84B2359AFFEF3A9A6" author="Poupin, J. &amp; Lemaitre, R." pageId="111" pageNumber="112" pagination="5 - 27" refId="ref75913" refString="Poupin, J. &amp; Lemaitre, R. (2014) Porcellanid crabs from Guadeloupe Island (Crustacea, Decapoda, Anomura), with an updated list of species from the Lesser Antilles. Zoosystema, 36 (1), 5 - 27. https: // doi. org / 10.5252 / z 2014 n 1 a 1" type="journal article" year="2014">Poupin &amp; Lemaitre 2014</bibRefCitation>
;
<bibRefCitation id="EFEEE08C770CFFA84F095982FE55A9A6" author="Poupin, J." box="[359,506,1979,2005]" pageId="111" pageNumber="112" refId="ref75876" refString="Poupin, J. (2018) Les crustaces decapodes des Petites Antilles: avec de nouvelles observations pour Saint-Martin, la Guadeloupe et la Martinique. Museum National d'Histoire Naturelle, Paris, 264 pp." type="book" year="2018">Poupin 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF00CDF57713FFB74EF959A1FC7DA9A7" ID-DOI="http://doi.org/10.5281/zenodo.5532381" ID-Zenodo-Dep="5532381" httpUri="https://zenodo.org/record/5532381/files/figure.png" pageId="112" pageNumber="113" startId="112.[151,250,1944,1968]" targetBox="[179,1407,179,1918]" targetPageId="112">
<paragraph id="8BC09D7D7713FFB74EF959A1FC7DA9A7" blockId="112.[151,1436,1944,2004]" pageId="112" pageNumber="113">
<emphasis id="B90B416F7713FFB74EF959A1FEB5A9C3" bold="true" box="[151,282,1944,1968]" pageId="112" pageNumber="113">FIGURE 69</emphasis>
.
<taxonomicName id="4C7FE6FE7713FFB74F4659A1FD09A9C3" authority="Glassell, 1937" authorityName="Glassell" authorityYear="1937" box="[296,678,1944,1968]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="polymitus">
<emphasis id="B90B416F7713FFB74F4659A1FDA5A9C3" box="[296,522,1944,1968]" italics="true" pageId="112" pageNumber="113">Petrolisthes polymitus</emphasis>
<bibRefCitation id="EFEEE08C7713FFB74C7F59A1FD09A9C3" author="Glassell, S. A." box="[529,678,1944,1968]" pageId="112" pageNumber="113" pagination="79 - 88" refId="ref71907" refString="Glassell, S. A. (1937) The Templeton Crocker Expedition. IV. Porcellanid crabs from the Gulf of California. Zoologica, 22 (4), 79 - 88. https: // doi. org / 10.2307 / 1436949" type="journal article" year="1937">Glassell, 1937</bibRefCitation>
</taxonomicName>
: A, B, ovigerous female (cl 4.0 mm, cw 4.1 mm) from Las Perlas Islands, Panama (MZUSP 33956), dorsal (A) and ventral (B) views [photographs by AA].
</paragraph>
</caption>
<caption id="DF00CDF57712FFB64EF95AF9FD83AB53" ID-DOI="http://doi.org/10.5281/zenodo.5532383" ID-Zenodo-Dep="5532383" httpUri="https://zenodo.org/record/5532383/files/figure.png" pageId="113" pageNumber="114" startId="113.[151,250,1216,1240]" targetBox="[201,1386,181,1192]" targetPageId="113">
<paragraph id="8BC09D7D7712FFB64EF95AF9FD83AB53" blockId="113.[151,1437,1216,1312]" pageId="113" pageNumber="114">
<emphasis id="B90B416F7712FFB64EF95AF9FEB5AAAB" bold="true" box="[151,282,1216,1240]" pageId="113" pageNumber="114">FIGURE 70</emphasis>
.
<taxonomicName id="4C7FE6FE7712FFB64F475AF9FD07AAAB" authority="Glassell, 1937" authorityName="Glassell" authorityYear="1937" box="[297,680,1216,1240]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="polymitus">
<emphasis id="B90B416F7712FFB64F475AF9FDA3AAAB" box="[297,524,1216,1240]" italics="true" pageId="113" pageNumber="114">Petrolisthes polymitus</emphasis>
<bibRefCitation id="EFEEE08C7712FFB64C7D5AF9FD07AAAB" author="Glassell, S. A." box="[531,680,1216,1240]" pageId="113" pageNumber="114" pagination="79 - 88" refId="ref71907" refString="Glassell, S. A. (1937) The Templeton Crocker Expedition. IV. Porcellanid crabs from the Gulf of California. Zoologica, 22 (4), 79 - 88. https: // doi. org / 10.2307 / 1436949" type="journal article" year="1937">Glassell, 1937</bibRefCitation>
</taxonomicName>
: A, male (cl cw indet.) from Ciudad de Panamá, Panama (not deposited), dorsal view; B, C, female (cl 4.5, cw 4.7) from Isla Afuerita, Coiba Archipelago, Panama (FLMNH UF 57718), dorsal (B) and ventral (C) views [photographs by AA].
</paragraph>
</caption>
<subSubSection id="C365CEF67712FFB64EA95B70FA89A998" pageId="113" pageNumber="114" type="discussion">
<paragraph id="8BC09D7D7712FFB64EA95B70FA32A82C" blockId="113.[151,1437,1353,2028]" pageId="113" pageNumber="114">
<emphasis id="B90B416F7712FFB64EA95B70FE9BAB10" bold="true" box="[199,308,1353,1379]" pageId="113" pageNumber="114">Remarks</emphasis>
.
<taxonomicName id="4C7FE6FE7712FFB64F2B5B70FDEAAB10" authorityName="Benedict" authorityYear="1901" box="[325,581,1353,1379]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B90B416F7712FFB64F2B5B70FDEAAB10" box="[325,581,1353,1379]" italics="true" pageId="113" pageNumber="114">Petrolisthes quadratus</emphasis>
</taxonomicName>
belongs to an exclusively American group of species of
<taxonomicName id="4C7FE6FE7712FFB64AB25B70FACEAB10" authorityName="Stimpson" authorityYear="1858" box="[1244,1377,1353,1379]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B90B416F7712FFB64AB25B70FACEAB10" box="[1244,1377,1353,1379]" italics="true" pageId="113" pageNumber="114">Petrolisthes</emphasis>
</taxonomicName>
with an unarmed carapace, relatively smooth, subrectangular P1 carpi and short, stubby ambulatory legs (P2–P4), which also includes the transisthmian
<taxonomicName id="4C7FE6FE7712FFB64C395BABFCC4ABD8" authority="Haig, 1960" authorityName="Haig" authorityYear="1960" box="[599,875,1425,1451]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="tonsorius">
<emphasis id="B90B416F7712FFB64C395BABFD70ABD8" box="[599,735,1426,1451]" italics="true" pageId="113" pageNumber="114">P. tonsorius</emphasis>
<bibRefCitation id="EFEEE08C7712FFB64C845BA8FCC4ABD8" author="Haig, J." box="[746,875,1425,1451]" pageId="113" pageNumber="114" pagination="1 - 440" refId="ref72816" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the Eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig, 1960</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7FE6FE7712FFB64DC45BABFAAAABDF" authority="Stimpson, 1895" authorityName="Stimpson" authorityYear="1895" box="[938,1285,1425,1452]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="tridentatus">
<emphasis id="B90B416F7712FFB64DC45BABFBECABD8" box="[938,1091,1425,1451]" italics="true" pageId="113" pageNumber="114">P. tridentatus</emphasis>
Stimpson, 1895
</taxonomicName>
, the recently described western Atlantic
<taxonomicName id="4C7FE6FE7712FFB64FB75B8FFC2FABBC" authority="Hiller &amp; Werding, 2019" authorityName="Hiller &amp; Werding" authorityYear="2019" box="[473,896,1461,1487]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="virgilius">
<emphasis id="B90B416F7712FFB64FB75B8FFDF9ABBC" box="[473,598,1461,1487]" italics="true" pageId="113" pageNumber="114">P. virgilius</emphasis>
<bibRefCitation id="EFEEE08C7712FFB64C0A5B8CFC2FABBC" author="Hiller, A. &amp; Werding, B." box="[612,896,1461,1487]" pageId="113" pageNumber="114" pagination="143 - 151" refId="ref73734" refString="Hiller, A. &amp; Werding, B. (2019) A new species of Petrolisthes (Crustacea, Anomura, Porcellanidae) inhabiting vermetid formations (Mollusca, Gastropoda, Vermetidae) in the southern Caribbean Sea. ZooKeys, 876, 143 - 151. https: // doi. org / 10.3897 / zookeys. 876.37244" type="journal article" year="2019">Hiller &amp; Werding, 2019</bibRefCitation>
</taxonomicName>
(see below and
<figureCitation id="134481F87712FFB64A235B8CFB15ABBC" box="[1101,1210,1461,1487]" captionStart="FIGURE 88" captionStartId="138.[151,250,1251,1275]" captionTargetBox="[151,1435,181,1226]" captionTargetId="figure-19@138.[151,1435,181,1226]" captionTargetPageId="138" captionText="FIGURE 88. Some Caribbean Porcellanidae reported from Colombia, in all likelihood also occurring in Panama: A, Madarateuchus vanderhorsti (Schmitt, 1924), male from Santa Marta, Colombia (CRBM-UV), dorsal view [photograph courtesy of Juan Felipe Lazarus-Agudelo]; B, Parapetrolisthes tortugensis (Glassell, 1945), ovigerous female from Utila, Honduras (not deposited), dorsal view; C, Petrolisthes amoenus (Guérin-Méneville, 1855), male from Saint Martin (FLMNH UF 32408), dorsal view, right P1 missing [photographs in B and C by AA]; D, Petrolisthes virgilius Hiller &amp; Werding, 2019, male from Golfo de Urabá, Colombia (CRBM-UV), dorsal view [photograph courtesy of Juan Felipe Lazarus-Agudelo]." figureDoi="http://doi.org/10.5281/zenodo.5532430" httpUri="https://zenodo.org/record/5532430/files/figure.png" pageId="113" pageNumber="114">Fig. 88D</figureCitation>
), and a number of eastern Pacific taxa (see also
<bibRefCitation id="EFEEE08C7712FFB64F855BE0FD1AAB80" author="Hiller, A. &amp; Kraus, H. &amp; Almon, M. &amp; Werding, B." box="[491,693,1497,1523]" pageId="113" pageNumber="114" pagination="547 - 569" refId="ref73577" refString="Hiller, A., Kraus, H., Almon, M. &amp; Werding, B. (2006) The Petrolisthes galathinus complex: species boundaries based on color pattern, morphology and molecules, and evolutionary interrelationships between the complex and other Porcellanidae (Crustacea: Decapoda: Anomura). Molecular Phylogenetics and Evolution, 40, 547 - 569. https: // doi. org / 10.1016 / j. ympev. 2006.03.030" type="journal article" year="2006">
Hiller
<emphasis id="B90B416F7712FFB64C5B5BE3FDDFAB80" box="[565,624,1497,1523]" italics="true" pageId="113" pageNumber="114">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="EFEEE08C7712FFB64CAC5BE0FC7EAB80" author="Hiller, A. &amp; Werding, B." box="[706,977,1497,1523]" pageId="113" pageNumber="114" pagination="143 - 151" refId="ref73734" refString="Hiller, A. &amp; Werding, B. (2019) A new species of Petrolisthes (Crustacea, Anomura, Porcellanidae) inhabiting vermetid formations (Mollusca, Gastropoda, Vermetidae) in the southern Caribbean Sea. ZooKeys, 876, 143 - 151. https: // doi. org / 10.3897 / zookeys. 876.37244" type="journal article" year="2019">Hiller &amp; Werding 2019</bibRefCitation>
). The morphological distinctiveness and validity of
<taxonomicName id="4C7FE6FE7712FFB64F775BC7FE05A864" authorityName="Benedict" authorityYear="1901" box="[281,426,1533,1559]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B90B416F7712FFB64F775BC7FE05A864" box="[281,426,1533,1559]" italics="true" pageId="113" pageNumber="114">P. quadratus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7FE6FE7712FFB64F865BC7FDC0A864" authorityName="Haig" authorityYear="1960" box="[488,623,1534,1559]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="tonsorius">
<emphasis id="B90B416F7712FFB64F865BC7FDC0A864" box="[488,623,1534,1559]" italics="true" pageId="113" pageNumber="114">P. tonsorius</emphasis>
</taxonomicName>
, presumably the most closely related species based on morphological grounds, were discussed by
<bibRefCitation id="EFEEE08C7712FFB64F955818FD16A848" author="Werding, B." box="[507,697,1569,1595]" pageId="113" pageNumber="114" pagination="173 - 214" refId="ref77184" refString="Werding, B. (1977) Los porcelanidos (Crustacea: Anomura: Porcellanidae) de la region de Santa Marta, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betin, 9, 173 - 214. https: // doi. org / 10.25268 / bimc. invemar. 1977.9.0.527" type="journal article" year="1977">Werding (1977)</bibRefCitation>
. The two species can be most easily separated from each other by the shape of the P1 carpus and P3 merus, and the proportions of the P2–P4 meri (
<bibRefCitation id="EFEEE08C7712FFB64A88587CFA3FA82C" author="Werding, B." box="[1254,1424,1605,1631]" pageId="113" pageNumber="114" pagination="173 - 214" refId="ref77184" refString="Werding, B. (1977) Los porcelanidos (Crustacea: Anomura: Porcellanidae) de la region de Santa Marta, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betin, 9, 173 - 214. https: // doi. org / 10.25268 / bimc. invemar. 1977.9.0.527" type="journal article" year="1977">Werding 1977</bibRefCitation>
).
</paragraph>
<paragraph id="8BC09D7D7712FFB64EA95850FA89A998" blockId="113.[151,1437,1353,2028]" pageId="113" pageNumber="114">
<bibRefCitation id="EFEEE08C7712FFB64EA95850FE7FA8F7" author="Chace, F. A. Jr. &amp; Hobbs, H. H. Jr." box="[199,464,1641,1668]" pageId="113" pageNumber="114" pagination="1 - 258" refId="ref70605" refString="Chace, F. A. Jr. &amp; Hobbs, H. H. Jr. (1969) The freshwater and terrestrial decapod crustaceans of the West Indies with special reference to Dominica. United States National Museum Bulletin, 292, i - vi + 1 - 258, 5 pls. https: // doi. org / 10.5479 / si. 03629236.292.1" type="journal article" year="1969">Chace &amp; Hobbs (1969)</bibRefCitation>
provided a detailed description of the colour pattern of
<taxonomicName id="4C7FE6FE7712FFB64A3F5853FB4FA8F0" authorityName="Benedict" authorityYear="1901" box="[1105,1248,1641,1667]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B90B416F7712FFB64A3F5853FB4FA8F0" box="[1105,1248,1641,1667]" italics="true" pageId="113" pageNumber="114">P. quadratus</emphasis>
</taxonomicName>
from
<collectingCountry id="F368DDED7712FFB64B495850FA37A8F0" box="[1319,1432,1641,1667]" name="Dominica" pageId="113" pageNumber="114">Dominica</collectingCountry>
, whereas
<bibRefCitation id="EFEEE08C7712FFB64E9658B4FE07A8D4" author="Werding, B." box="[248,424,1677,1703]" pageId="113" pageNumber="114" pagination="173 - 214" refId="ref77184" refString="Werding, B. (1977) Los porcelanidos (Crustacea: Anomura: Porcellanidae) de la region de Santa Marta, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betin, 9, 173 - 214. https: // doi. org / 10.25268 / bimc. invemar. 1977.9.0.527" type="journal article" year="1977">Werding (1977)</bibRefCitation>
and
<bibRefCitation id="EFEEE08C7712FFB64FB258B4FCACA8D4" author="Poupin, J. &amp; Lemaitre, R." box="[476,771,1677,1703]" pageId="113" pageNumber="114" pagination="5 - 27" refId="ref75913" refString="Poupin, J. &amp; Lemaitre, R. (2014) Porcellanid crabs from Guadeloupe Island (Crustacea, Decapoda, Anomura), with an updated list of species from the Lesser Antilles. Zoosystema, 36 (1), 5 - 27. https: // doi. org / 10.5252 / z 2014 n 1 a 1" type="journal article" year="2014">Poupin &amp; Lemaitre (2014)</bibRefCitation>
found
<taxonomicName id="4C7FE6FE7712FFB64D3E58B7FC73A8D4" authorityName="Benedict" authorityYear="1901" box="[848,988,1677,1703]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="B90B416F7712FFB64D3E58B7FC73A8D4" box="[848,988,1677,1703]" italics="true" pageId="113" pageNumber="114">P. quadratus</emphasis>
</taxonomicName>
to be somewhat variable in colour. Most
<bibRefCitation id="EFEEE08C7712FFB64EF95888FEF4A8B8" author="Werding, B." box="[151,347,1713,1739]" pageId="113" pageNumber="114" pagination="173 - 214" refId="ref77184" refString="Werding, B. (1977) Los porcelanidos (Crustacea: Anomura: Porcellanidae) de la region de Santa Marta, Colombia. Anales del Instituto de Investigaciones Marinas de Punta Betin, 9, 173 - 214. https: // doi. org / 10.25268 / bimc. invemar. 1977.9.0.527" type="journal article" year="1977">Werding’s (1977)</bibRefCitation>
specimens from Santa Marta,
<collectingCountry id="F368DDED7712FFB64CC05888FCB0A8B8" box="[686,799,1713,1739]" name="Colombia" pageId="113" pageNumber="114">Colombia</collectingCountry>
, were sandy yellowish, with the P1 darker and often with an olive tinge; however, in some individuals, the carapace was speckled with brown spots. Similarly,
<bibRefCitation id="EFEEE08C7712FFB64B4358ECFEE1A960" author="Poupin, J. &amp; Lemaitre, R." pageId="113" pageNumber="114" pagination="5 - 27" refId="ref75913" refString="Poupin, J. &amp; Lemaitre, R. (2014) Porcellanid crabs from Guadeloupe Island (Crustacea, Decapoda, Anomura), with an updated list of species from the Lesser Antilles. Zoosystema, 36 (1), 5 - 27. https: // doi. org / 10.5252 / z 2014 n 1 a 1" type="journal article" year="2014">Poupin &amp; Lemaitre (2014)</bibRefCitation>
stated that in the specimens from
<collectingCountry id="F368DDED7712FFB64CA158C0FCF6A960" box="[719,857,1785,1811]" name="Guadeloupe" pageId="113" pageNumber="114">Guadeloupe</collectingCountry>
, the carapace and P1 may be either brown speckled with white or uniform creamy white sandy pale; the P2–P4 are transversely banded with white and pale brown (see their fig. 4D). The herein illustrated specimens from
<collectingCountry id="F368DDED7712FFB64C855978FCD7A928" box="[747,888,1857,1883]" name="Sint Martin" pageId="113" pageNumber="114">Saint Martin</collectingCountry>
and
<collectingCountry id="F368DDED7712FFB64DDE5978FBA6A928" box="[944,1033,1857,1883]" name="Bonaire, Sint Eustatius and Saba" pageId="113" pageNumber="114">Bonaire</collectingCountry>
(
<figureCitation id="134481F87712FFB64A765978FBD9A928" box="[1048,1142,1857,1883]" captionStart="FIGURE 71" captionStartId="114.[151,250,1634,1658]" captionTargetBox="[168,1418,181,1610]" captionTargetId="figure-19@114.[168,1418,181,1610]" captionTargetPageId="114" captionText="FIGURE 71. Petrolisthes quadratus Benedict, 1901:A, B, extra-limital specimen, male (cl cw indet.) from Saint Martin (FLM- NH UF 31974), dorsal (A) and ventral (B) views [photographs by AA]." figureDoi="http://doi.org/10.5281/zenodo.5532388" httpUri="https://zenodo.org/record/5532388/files/figure.png" pageId="113" pageNumber="114">Figs. 71</figureCitation>
,
<figureCitation id="134481F87712FFB64AEC5978FB00A928" box="[1154,1199,1857,1883]" captionStart="FIGURE 90" captionStartId="140.[151,250,1624,1648]" captionTargetBox="[164,1424,181,1599]" captionTargetId="figure-19@140.[164,1424,181,1601]" captionTargetPageId="140" captionText="FIGURE 90. Some western Atlantic Porcellanidae (all species reported from Panama or Golfo de Urabá at the Panamanian- Colombian border) photographed in situ: A, Neopisosoma angustifrons (Benedict, 1901), Golfo de Urabá, Colombia; B, Neopisosoma neglectum Werding, 1986, Golfo de Urabá, Colombia; C, Pachycheles tuerkayi Werding &amp; Hiller, 2017, Santa Marta, Colombia; D, Petrolisthes jugosus Streets, 1872, Tayrona, Colombia; E, Petrolisthes tonsorius Haig, 1960, blue colour morph, Bonaire; F, Petrolisthes quadratus Benedict, 1901, Bonaire [photographs courtesy of Juan-Felipe Lazarus-Agudelo (A–D) and Marco Faasse (E, F)]." figureDoi="http://doi.org/10.5281/zenodo.5532436" httpUri="https://zenodo.org/record/5532436/files/figure.png" pageId="113" pageNumber="114">90F</figureCitation>
) conform well to the previously described colour patterns (see
<bibRefCitation id="EFEEE08C7712FFB64C0C595CFCD5A90C" author="Poupin, J. &amp; Lemaitre, R." box="[610,890,1893,1919]" pageId="113" pageNumber="114" pagination="5 - 27" refId="ref75913" refString="Poupin, J. &amp; Lemaitre, R. (2014) Porcellanid crabs from Guadeloupe Island (Crustacea, Decapoda, Anomura), with an updated list of species from the Lesser Antilles. Zoosystema, 36 (1), 5 - 27. https: // doi. org / 10.5252 / z 2014 n 1 a 1" type="journal article" year="2014">Poupin &amp; Lemaitre 2014</bibRefCitation>
: fig. 4D for the other colour variant).
<bibRefCitation id="EFEEE08C7712FFB64B7B595CFA33A9F3" author="Haig, J." box="[1301,1436,1893,1920]" pageId="113" pageNumber="114" pagination="1 - 44" refId="ref72706" refString="Haig, J. (1956) The Galatheidea (Crustacea Anomura) of the Allan Hancock Atlantic Expedition with a review of the Porcellanidae of the western North Atlantic. Allan Hancock Atlantic Expedition Report, 8, 1 - 44." type="journal article" year="1956">Haig (1956)</bibRefCitation>
did not report on the colour pattern of her single female specimen from Bahía de Caledonia, presently the only known record from
<collectingCountry id="F368DDED7712FFB64F195997FE7CA9BB" box="[375,467,1966,1992]" name="Panama" pageId="113" pageNumber="114">Panama</collectingCountry>
. However, she noticed that it “differs from the rest of the material examined in having a brush of pubescence on the outer surface of the left hand”, thus leaving some doubts about this record.
</paragraph>
</subSubSection>
</treatment>
</document>
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