167 lines
18 KiB
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167 lines
18 KiB
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<document id="DD1F29F3D1338ECCFFA37A36083B7B4B" ID-CLB-Dataset="187956" ID-DOI="10.11646/zootaxa.5205.6.1" ID-GBIF-Dataset="4b6eb79c-018f-470b-ac46-6646270ec7ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="7318605" ID-ZooBank="9D1DEC25-5FA2-4D64-807E-F103C3FCB5CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668422045351" checkinUser="plazi" docAuthor="Shear, William A. & Marek, Paul E." docDate="2022" docId="03F89303FFBFFFB29AF92FC4E02CFC18" docLanguage="en" docName="zootaxa.5205.6.1.pdf" docOrigin="Zootaxa 5205 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nototrisaria Shear & Marek 2022, new genus" docType="treatment" docVersion="7" lastPageNumber="504" masterDocId="FFC1EB7BFFBCFFB19A6E2F53E208FFDC" masterDocTitle="The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="531" masterPageNumber="501" pageNumber="504" updateTime="1699411384392" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="C3EABC5CBCD12CDCA967A93F99834FD9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9A516589DC3182827821AB80B42663DD">
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<mods:title id="BF32B1237BF471FD23E1A4905C713CE3">The millipede family Striariidae Bollman, 1893. VI. Six new genera and thirteen new species from western North America (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="24AD1130BB86936A808468A6C41C95A9" type="personal">
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<mods:roleTerm id="9932FD7643D129503D7569A1A31CC4D0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CAD0B0BF4819FE30414454924269F693">Shear, William A.</mods:namePart>
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<mods:nameIdentifier id="B783B03AD749A424EDABCCFACA716910" type="ORCID">0000-0002-5887-7003</mods:nameIdentifier>
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<mods:affiliation id="E47C7D3FB5D35D0AE16BF2556C538F63">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville VA 23901 USA. wshear @ hsc. edu; https: // orcid. org / 0000 - 0002 - 5887 - 7003</mods:affiliation>
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<mods:nameIdentifier id="ED2F4A2E987585EFED3CE28C5D674FC5" type="email">wshear@hsc.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="AF81AEB3F68A8620748ACA152C3BD849" type="personal">
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<mods:roleTerm id="D3B3B1830B533690095E5412E43D627E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7F5BA74E13C8A9798EDEF55721264D42">Marek, Paul E.</mods:namePart>
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<mods:affiliation id="79ED759DA95AADB48DA9EF2080C60138">Department of Entomology, Virginia Tech, Blacksburg VA 24061 USA.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="88C98B0F28C6E0C21862529DF2F9176A">text</mods:typeOfResource>
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<mods:titleInfo id="636E13F98270A2A9AC0D3A3A077FB226">
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<mods:title id="C0905AE5B23460143811F382FBD66E32">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="90AC325A6EFE1673DAE11E78AD72239B">
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<mods:date id="1E7E541A6C630A6EC61C08C0A7210AF8">2022</mods:date>
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<mods:detail id="E7CDCFBAE8DC57411EE95304E48CE5AD" type="pubDate">
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<mods:number id="F1F6DCD73F69AC53AB0F728CDA4EB6F5">2022-11-11</mods:number>
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<mods:detail id="B6C342277146BEB04FC5F640C2CD7763" type="volume">
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<mods:number id="9AFEF1C9CA1AB756A9DA9B00E5317064">5205</mods:number>
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<mods:detail id="CE896C58F9195A759920009A08525480" type="issue">
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<mods:number id="EC0155B9FA033F183332FDC807633871">6</mods:number>
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<mods:start id="878E9742AC8DC8A4ACC019043ABA737C">501</mods:start>
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<mods:classification id="5C2B43EA14D9F94C14A88CB4CAC62C17">journal article</mods:classification>
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<mods:identifier id="8FFE3052EDA549D579AA0671F7E88655" type="CLB-Dataset">187956</mods:identifier>
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<mods:identifier id="0F5C45889F6FE9EFE27F35BB02504DBA" type="DOI">10.11646/zootaxa.5205.6.1</mods:identifier>
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<mods:identifier id="68C5A6FB4C9F23BE46A0517C77DC2C20" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03F89303FFBFFFB29AF92FC4E02CFC18" ID-DOI="http://doi.org/10.5281/zenodo.7327157" ID-GBIF-Taxon="205076356" ID-Zenodo-Dep="7327157" LSID="urn:lsid:plazi:treatment:03F89303FFBFFFB29AF92FC4E02CFC18" httpUri="http://treatment.plazi.org/id/03F89303FFBFFFB29AF92FC4E02CFC18" lastPageNumber="504" pageId="3" pageNumber="504">
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<subSubSection id="C34B719EFFBFFFB29AF92FC4E0D5FF6D" box="[151,733,151,177]" pageId="3" pageNumber="504" type="nomenclature">
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<paragraph id="8BEE2215FFBFFFB29AF92FC4E0D5FF6D" blockId="3.[151,733,151,177]" box="[151,733,151,177]" pageId="3" pageNumber="504">
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<heading id="D0A69579FFBFFFB29AF92FC4E0D5FF6D" bold="true" box="[151,733,151,177]" fontSize="11" level="1" pageId="3" pageNumber="504" reason="1">
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<emphasis id="B925FE07FFBFFFB29AF92FC4E0D5FF6D" bold="true" box="[151,733,151,177]" pageId="3" pageNumber="504">
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Genus
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<taxonomicName id="4C515996FFBFFFB29A802FC4E046FF6D" authority="Shear & Marek" authorityName="Shear & Marek" authorityYear="2022" box="[238,590,151,177]" class="Diplopoda" family="Striariidae" genus="Nototrisaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B925FE07FFBFFFB29A802FC4E389FF6D" bold="true" box="[238,385,151,177]" italics="true" pageId="3" pageNumber="504">Nototrisaria</emphasis>
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Shear & Marek
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</taxonomicName>
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,
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<taxonomicNameLabel id="A216437CFFBFFFB298352FC4E0D5FF6D" box="[603,733,151,177]" pageId="3" pageNumber="504" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B719EFFBFFFB29AF92F8FE112FF28" box="[151,794,219,244]" pageId="3" pageNumber="504" type="type_taxon">
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<paragraph id="8BEE2215FFBFFFB29AF92F8FE112FF28" blockId="3.[151,794,219,244]" box="[151,794,219,244]" pageId="3" pageNumber="504">
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<emphasis id="B925FE07FFBFFFB29AF92F8FE316FF2F" bold="true" box="[151,286,219,244]" pageId="3" pageNumber="504">
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<typeStatus id="54EA9CB7FFBFFFB29AF92F8FE2C4FF28" box="[151,204,220,244]" pageId="3" pageNumber="504">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C515996FFBFFFB29B442F8FE09FFF28" authority="Shear & Marek" authorityName="Shear & Marek" authorityYear="2022" box="[298,663,220,244]" class="Diplopoda" family="Striariidae" genus="Nototrisaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
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<emphasis id="B925FE07FFBFFFB29B442F8FE3FAFF2F" box="[298,498,220,243]" italics="true" pageId="3" pageNumber="504">Nototrisaria ornata</emphasis>
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Shear & Marek
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</taxonomicName>
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,
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<taxonomicNameLabel id="A216437CFFBFFFB298CF2F8FE112FF28" box="[673,794,220,244]" pageId="3" pageNumber="504" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B719EFFBFFFB29AF92E71E757FEBC" pageId="3" pageNumber="504" type="etymology">
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<paragraph id="8BEE2215FFBFFFB29AF92E71E757FEBC" blockId="3.[151,1437,290,964]" pageId="3" pageNumber="504">
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<emphasis id="B925FE07FFBFFFB29AF92E71E32AFEE0" bold="true" box="[151,290,290,316]" pageId="3" pageNumber="504">Etymology:</emphasis>
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The name of the genus combines
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<emphasis id="B925FE07FFBFFFB298CB2E70E0E8FEE0" box="[677,736,291,316]" italics="true" pageId="3" pageNumber="504">noto-</emphasis>
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in reference to the enlarged, paranotum-like sixth crests (C6), and
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<emphasis id="B925FE07FFBFFFB29AA92E14E320FEBC" box="[199,296,327,352]" italics="true" pageId="3" pageNumber="504">
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<taxonomicName id="4C515996FFBFFFB29AA92E14E32CFEBC" authorityName="Shear" authorityYear="2020" box="[199,292,327,352]" class="Diplopoda" family="Striariidae" genus="Trisaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">Trisaria</taxonomicName>
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,
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</emphasis>
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the name of a related genus in the subfamily. The name should be treated as feminine in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B719EFFBFFFB29AA92E3AE721FE10" pageId="3" pageNumber="504" type="diagnosis">
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<paragraph id="8BEE2215FFBFFFB29AA92E3AE721FE10" blockId="3.[151,1437,290,964]" pageId="3" pageNumber="504">
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<emphasis id="B925FE07FFBFFFB29AA92E3AE34CFE5F" bold="true" box="[199,324,361,387]" pageId="3" pageNumber="504">Diagnosis:</emphasis>
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A genus of
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<taxonomicName id="4C515996FFBFFFB29BBD2E39E043FE58" authorityName="Shear" authorityYear="2020" box="[467,587,362,388]" class="Diplopoda" family="Striariidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="531" phylum="Arthropoda" rank="subFamily" subFamily="Trisariinae">Trisariinae</taxonomicName>
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composed of a single small species, distinct from other small striariids in the highly ornate crests of the metazonites, with C1 on anterior rings consisting of anterior and posterior knobs but disappearing on posterior rings, C2 strongly exaggerated, and C6 expanded into paranota (see
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<figureCitation id="136A3E90FFBFFFB29ED62EE1E714FE10" box="[1208,1308,434,460]" captionStart="FIGURES 2–6" captionStartId="4.[152,270,1887,1911]" captionTargetBox="[200,1388,182,1863]" captionTargetId="figure-25@4.[199,1388,181,1863]" captionTargetPageId="4" captionText="FIGURES 2–6. Nototrisaria ornata, n. gen., n. sp. 2. Anterior end of male, ventral view. 3. Rings 2–4 of male, dorsal view. 4. Posterior end of male, oblique view. 5. Right gonopod, lateral view. 6. Legpair 9 of male, anterior view. Abbreviations: aac, anterior angiocoxite; col, collum; cc, colpodoxite; c1, crest 1; c6, crest 6; cxs, coxosternite; cx10, coxa of leg 10; L1, leg 1; L2, leg 2; L3, leg 3; m, mandible; numerals, ring numbers; pac, posterior angiocoxite; t9, telopodite of leg 9." figureDoi="http://doi.org/10.5281/zenodo.7318609" httpUri="https://zenodo.org/record/7318609/files/figure.png" pageId="3" pageNumber="504">Figs 3, 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B719EFFBFFFB29AA92E86E13FFE2C" box="[199,823,469,496]" pageId="3" pageNumber="504" type="description">
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<paragraph id="8BEE2215FFBFFFB29AA92E86E13FFE2C" blockId="3.[151,1437,290,964]" box="[199,823,469,496]" pageId="3" pageNumber="504">
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<emphasis id="B925FE07FFBFFFB29AA92E86E353FE33" bold="true" box="[199,347,469,495]" pageId="3" pageNumber="504">Description:</emphasis>
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As for the single species described below.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B719EFFBFFFB29AA92EAAE02CFC18" pageId="3" pageNumber="504" type="distribution">
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<paragraph id="8BEE2215FFBFFFB29AA92EAAE06AFDE4" blockId="3.[151,1437,290,964]" pageId="3" pageNumber="504">
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<emphasis id="B925FE07FFBFFFB29AA92EAAE36BFDCF" bold="true" box="[199,355,505,531]" pageId="3" pageNumber="504">Distribution:</emphasis>
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Widely distributed in
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<collectingRegion id="4995ECF7FFBFFFB298372EA9E114FDC8" box="[601,796,506,532]" country="United States of America" name="Washington" pageId="3" pageNumber="504">Washington State</collectingRegion>
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west of the Cascade Ranges, including Pacific, Jefferson, Whakiakum and Grays Harbor Counties.
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</paragraph>
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<paragraph id="8BEE2215FFBFFFB29AA92D11E6ADFCE8" blockId="3.[151,1437,290,964]" pageId="3" pageNumber="504">
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<emphasis id="B925FE07FFBFFFB29AA92D11E2F7FD80" bold="true" box="[199,255,578,604]" pageId="3" pageNumber="504">Note</emphasis>
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: The genus has similarities to the previously described
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<emphasis id="B925FE07FFBFFFB299E92D11E611FD80" box="[903,1049,578,604]" italics="true" pageId="3" pageNumber="504">Stegostriaria</emphasis>
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<bibRefCitation id="EFC05FE4FFBFFFB29E4D2D11E716FD80" author="Shear, W. A. & Marek, P. E." box="[1059,1310,578,604]" pageId="3" pageNumber="504" pagination="461 - 472" refId="ref17383" refString="Shear, W. A. & Marek, P. E. (2022) The millipede family Striariidae Bollman, 1893. V. Stegostriaria dulcidormus, n. gen., n. sp., Kentrostriaria ohara, n. gen., n. sp., and the convergent evolution of exaggerated metazonital crests (Diplopoda, Chordeumatida, Striarioidea). Zootaxa, 5094 (3), 461 - 472. https: // doi. org / 10.11646 / zootaxa. 5094.3.5" type="journal article" year="2022">
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Shear & Marek,
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<quantity id="4CA98FF0FFBFFFB29E882D11E716FD80" box="[1254,1310,578,604]" metricMagnitude="1" metricUnit="m" metricValue="5.135879999999999" pageId="3" pageNumber="504" unit="in" value="2022.0">2022</quantity>
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</bibRefCitation>
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in that C2 is elevated and larger than all the other crests. However,
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<taxonomicName id="4C515996FFBFFFB2994D2D34E609FD5C" authority="Shear & Marek, 2022" authorityName="Shear & Marek" authorityYear="2022" box="[803,1025,615,640]" class="Diplopoda" family="Striariidae" genus="Nototrisaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="ornata" status="n. gen., n. sp.">
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<emphasis id="B925FE07FFBFFFB2994D2D34E609FD5C" box="[803,1025,615,640]" italics="true" pageId="3" pageNumber="504">Nototrisaria ornata</emphasis>
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</taxonomicName>
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<emphasis id="B925FE07FFBFFFB29E672D35E6A1FD5C" bold="true" box="[1033,1193,614,640]" pageId="3" pageNumber="504">
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<taxonomicNameLabel id="A216437CFFBFFFB29E672D35E6A1FD5C" box="[1033,1193,614,640]" pageId="3" pageNumber="504" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</emphasis>
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, also has C6 broadly extended, forming paranota, C1 is suppressed and missing posterior of the midbody rings, and there are no intercalary crests. The gonopods of
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<taxonomicName id="4C515996FFBFFFB29BDE2DFCE032FD14" authority="Shear & Marek, 2022" authorityName="Shear & Marek" authorityYear="2022" box="[432,570,687,712]" class="Diplopoda" family="Striariidae" genus="Nototrisaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B925FE07FFBFFFB29BDE2DFCE032FD14" box="[432,570,687,712]" italics="true" pageId="3" pageNumber="504">Nototrisaria</emphasis>
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</taxonomicName>
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<emphasis id="B925FE07FFBFFFB2982A2DFDE09CFD14" bold="true" box="[580,660,686,712]" pageId="3" pageNumber="504">
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<taxonomicNameLabel id="A216437CFFBFFFB2982A2DFDE09CFD14" box="[580,660,686,712]" pageId="3" pageNumber="504" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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are very different from those of
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<emphasis id="B925FE07FFBFFFB29E602DFDE73DFD14" box="[1038,1333,686,712]" italics="true" pageId="3" pageNumber="504">Stegostriaria dulcidormus</emphasis>
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<bibRefCitation id="EFC05FE4FFBFFFB29F502DFDE324FD30" author="Shear, W. A. & Marek, P. E." pageId="3" pageNumber="504" pagination="461 - 472" refId="ref17383" refString="Shear, W. A. & Marek, P. E. (2022) The millipede family Striariidae Bollman, 1893. V. Stegostriaria dulcidormus, n. gen., n. sp., Kentrostriaria ohara, n. gen., n. sp., and the convergent evolution of exaggerated metazonital crests (Diplopoda, Chordeumatida, Striarioidea). Zootaxa, 5094 (3), 461 - 472. https: // doi. org / 10.11646 / zootaxa. 5094.3.5" type="journal article" year="2022">Shear & Marek, 2022</bibRefCitation>
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, and the complicated, highly modified ninth legs of that species are unique and unlike the simpler ninth legs of
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<taxonomicName id="4C515996FFBFFFB29B442DA4E00EFCCC" authority="Shear & Marek, 2022" authorityName="Shear & Marek" authorityYear="2022" box="[298,518,759,784]" class="Diplopoda" family="Striariidae" genus="Nototrisaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="ornata" status="n. gen., n. sp.">
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<emphasis id="B925FE07FFBFFFB29B442DA4E00EFCCC" box="[298,518,759,784]" italics="true" pageId="3" pageNumber="504">Nototrisaria ornata</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="B925FE07FFBFFFB298632DA5E0A0FCCC" bold="true" box="[525,680,758,784]" pageId="3" pageNumber="504">
|
|||
|
<taxonomicNameLabel id="A216437CFFBFFFB298632DA5E0A0FCCC" box="[525,680,758,784]" pageId="3" pageNumber="504" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
|||
|
</emphasis>
|
|||
|
<emphasis id="B925FE07FFBFFFB298C12DA5E1DCFCCC" box="[687,980,758,784]" italics="true" pageId="3" pageNumber="504">Stegostriaria dulcidormus</emphasis>
|
|||
|
is known only from Linn and Tillamook Counties in
|
|||
|
<collectingRegion id="4995ECF7FFBFFFB29B712C49E37EFCE8" box="[287,374,794,820]" country="United States of America" name="Oregon" pageId="3" pageNumber="504">Oregon</collectingRegion>
|
|||
|
, well to the south, although the intervening forests are not well-sampled.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8BEE2215FFBFFFB29AA92C6DE02CFC18" blockId="3.[151,1437,290,964]" pageId="3" pageNumber="504">
|
|||
|
Despite the differences in the gonopods and ninth legs from species of
|
|||
|
<taxonomicName id="4C515996FFBFFFB29E692C6CE669FC84" authorityName="Shear" authorityYear="2020" box="[1031,1121,831,856]" class="Diplopoda" family="Striariidae" genus="Trisaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B925FE07FFBFFFB29E692C6CE669FC84" box="[1031,1121,831,856]" italics="true" pageId="3" pageNumber="504">Trisaria</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, the absence of a gonopod flagellocoxite and the fusion of all elements of the ninth legs, as well as an apparent comblike branch evidently fused to the anterior angiocoxite and a strongly fimbriate colpocoxite indicate that this genus and species are best placed in the subfamily
|
|||
|
<taxonomicName id="4C515996FFBFFFB29BCA2CF9E028FC18" authorityName="Shear" authorityYear="2020" box="[420,544,938,964]" class="Diplopoda" family="Striariidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="504" phylum="Arthropoda" rank="subFamily" subFamily="Trisariinae">Trisariinae</taxonomicName>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|