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<document ID-DOI="10.5252/geodiversitas2019v41a15" ID-GBIF-Dataset="1e89d723-7f54-4d6b-85dd-6bcd573eca32" ID-ISSN="1638-9395" ID-Zenodo-Dep="3694209" ID-ZooBank="urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C" checkinTime="1583172268754" checkinUser="valdenar" docAuthor="Bonis, Louis de, Gardin, Axelle &amp; Blondel, Cécile" docDate="2019" docId="03A487991A65FF9BFF52F9396B5AFB9E" docLanguage="en" docName="Geodiversitas.41.15.601-621.pdf.imf" docOrigin="Geodiversitas 41 (15)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pachycynodon Schlosser 1888" docType="treatment" docVersion="7" lastPageNumber="604" masterDocId="FF9DFFE11A60FF9EFFADFF8D6E62FF8E" masterDocTitle="Carnivora from the early Oligocene of the ‘ Phosphorites du Quercy’ in southwestern France" masterLastPageNumber="621" masterPageNumber="601" pageNumber="604" updateTime="1675702286038" updateUser="tatiana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Carnivora from the early Oligocene of the ‘ Phosphorites du Quercy’ in southwestern France</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bonis, Louis de</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">louis.debonis@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gardin, Axelle</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">axelle.gardin@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Blondel, Cécile</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">cecile.blondel@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-09-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>41</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>15</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>601</mods:start>
<mods:end>621</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2019v41a15</mods:identifier>
<mods:identifier type="GBIF-Dataset">1e89d723-7f54-4d6b-85dd-6bcd573eca32</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="Zenodo-Dep">3694209</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3703507" ID-GBIF-Taxon="162367536" ID-Zenodo-Dep="3703507" LSID="urn:lsid:plazi:treatment:03A487991A65FF9BFF52F9396B5AFB9E" httpUri="http://treatment.plazi.org/id/03A487991A65FF9BFF52F9396B5AFB9E" lastPageNumber="604" pageId="5" pageNumber="604">
<subSubSection box="[255,652,1716,1742]" pageId="5" pageNumber="604" type="nomenclature">
<paragraph blockId="5.[255,652,1716,1742]" box="[255,652,1716,1742]" pageId="5" pageNumber="604">
<heading box="[255,652,1716,1742]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="604" reason="2">
Genus
<taxonomicName authority="Schlosser, 1888" authorityName="Schlosser" authorityYear="1888" box="[332,652,1716,1742]" class="Mammalia" family="Ursidae" genus="Pachycynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis box="[332,477,1716,1742]" italics="true" pageId="5" pageNumber="604">Pachycynodon</emphasis>
<bibRefCitation author="Schlosser" box="[484,652,1716,1742]" firstAuthor="Schlosser" pageId="5" pageNumber="604" pagination="1 - 162" refId="ref19017" refString="SCHLOSSER M. 1888. - Die Affen, Lemuren, Chiropteren, Insectivoren, Marsupialier, Creodonten und Carnivoren des europaischen Tertiars und deren Beziehungen zu ihren lebenden und fossilen aussereuropaischen Verwandten. Beitrage zur Palaontologie Osterreich Ungarns und des Orients I. Theil 6: 1 - 224. II Theil 7: 1 - 162." type="journal article" year="1888">Schlosser, 1888</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="604" type="materials_examined">
<paragraph blockId="5.[132,775,1772,1823]" pageId="5" pageNumber="604">
<typeStatus box="[132,181,1772,1796]" pageId="5" pageNumber="604">
<collectionCode box="[132,181,1772,1796]" pageId="5" pageNumber="604">TYPE</collectionCode>
</typeStatus>
<collectionCode box="[186,257,1777,1794]" pageId="5" pageNumber="604">SPECIES</collectionCode>
. —
<taxonomicName authority="Schlosser, 1888" authorityName="Schlosser" authorityYear="1888" box="[298,689,1773,1797]" class="Mammalia" family="Ursidae" genus="Pachycynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="species" species="crassirostris">
<emphasis box="[298,535,1773,1797]" italics="true" pageId="5" pageNumber="604">Pachycynodon crassirostris</emphasis>
<bibRefCitation author="Schlosser" box="[539,689,1773,1797]" firstAuthor="Schlosser" pageId="5" pageNumber="604" pagination="1 - 162" refId="ref19017" refString="SCHLOSSER M. 1888. - Die Affen, Lemuren, Chiropteren, Insectivoren, Marsupialier, Creodonten und Carnivoren des europaischen Tertiars und deren Beziehungen zu ihren lebenden und fossilen aussereuropaischen Verwandten. Beitrage zur Palaontologie Osterreich Ungarns und des Orients I. Theil 6: 1 - 224. II Theil 7: 1 - 162." type="journal article" year="1888">Schlosser, 1888</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="604" type="description">
<paragraph blockId="5.[132,775,1875,2030]" box="[132,277,1875,1901]" pageId="5" pageNumber="604">DESCRIPTION</paragraph>
<paragraph blockId="5.[132,775,1875,2030]" lastBlockId="5.[813,1456,536,1041]" pageId="5" pageNumber="604">
Schlosser created the genus
<taxonomicName authorityName="Schlosser" authorityYear="1888" box="[437,580,1907,1933]" class="Mammalia" family="Ursidae" genus="Pachycynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="610" phylum="Chordata" rank="genus">
<emphasis box="[437,580,1907,1933]" italics="true" pageId="5" pageNumber="604">Pachycynodon</emphasis>
</taxonomicName>
for a P4 and M1 from the Quercy phosphorites (Schlosser 1988: pl. IX, fig. 2, 7, 9 and pl. IX, fig. 4 respectively) without any biostratigraphic context. He thought that these specimens were conspecific with
<emphasis box="[871,979,536,562]" italics="true" pageId="5" pageNumber="604">Cynodictis</emphasis>
<taxonomicName authority="Filhol, 1876" authorityName="Filhol" authorityYear="1876" box="[987,1247,536,562]" class="Mammalia" family="Ursidae" genus="Pachycynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="species" species="crassirostris">
<emphasis box="[987,1102,537,562]" italics="true" pageId="5" pageNumber="604">crassirostris</emphasis>
<bibRefCitation author="Filhol" box="[1111,1247,536,562]" firstAuthor="Filhol" pageId="5" pageNumber="604" pagination="1 - 338" refId="ref17470" refString="FILHOL H. 1876 - 1877. - Recherches sur les phosphorites du Quercy. Etudes des fossiles qu'on y rencontre et specialement des mammiferes. Bibliotheque de l'Ecole des Hautes Etudes, Section des Sciences naturelles. Premiere partie, 15 (4): 1 - 220. Deuxieme partie 16 (1): 1 - 338." type="journal article" year="1876">Filhol, 1876</bibRefCitation>
</taxonomicName>
(
<bibRefCitation author="Filhol" box="[1264,1325,536,562]" firstAuthor="Filhol" pageId="5" pageNumber="604" pagination="1 - 338" refId="ref17470" refString="FILHOL H. 1876 - 1877. - Recherches sur les phosphorites du Quercy. Etudes des fossiles qu'on y rencontre et specialement des mammiferes. Bibliotheque de l'Ecole des Hautes Etudes, Section des Sciences naturelles. Premiere partie, 15 (4): 1 - 220. Deuxieme partie 16 (1): 1 - 338." type="journal article" year="1876">1876</bibRefCitation>
: figs 67, 69 and 72) but, in fact the Filhol’s specimen, a mandible with c, p3, m1-m2, belongs to the genus
<taxonomicName authorityName="Filhol" authorityYear="1881" box="[1222,1378,600,626]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis box="[1222,1378,600,626]" italics="true" pageId="5" pageNumber="604">Amphicynodon</emphasis>
</taxonomicName>
. Thus, the name
<emphasis box="[918,1032,632,657]" italics="true" pageId="5" pageNumber="604">crassirostris</emphasis>
being conserved, the
<typeStatus box="[1262,1308,632,658]" pageId="5" pageNumber="604">type</typeStatus>
species of the genus is
<taxonomicName authorityName="Filhol" authorityYear="1876" box="[904,1168,664,690]" class="Mammalia" family="Ursidae" genus="Pachycynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="species" species="crassirostris">
<emphasis box="[904,1168,664,690]" italics="true" pageId="5" pageNumber="604">Pachycynodon crassirostris</emphasis>
</taxonomicName>
. Later, several species were added to the genus and to the
<taxonomicName authorityName="Simpson" authorityYear="1945" box="[1155,1381,695,721]" class="Mammalia" family="Amphicynodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="family">Amphicynodontidae</taxonomicName>
, forming a group with robust dentition (more robust than that of
<taxonomicName authorityName="Filhol" authorityYear="1881" box="[813,967,759,785]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis box="[813,967,759,785]" italics="true" pageId="5" pageNumber="604">Amphicynodon</emphasis>
</taxonomicName>
), mandible often deep, p4 as high or higher than the protoconid of m1, massive cuspids of the trigonid of m1, postmetaconid cristid sloping very obliquely toward the preentocristid, paraconid often present in m2. The robust P4 has a short metastyle and large plateau-like protocone. There is a trend in M1 toward an almost quadratic outline, without paraconule and disto-lingual metaconule and with a large cingulum around surrounding the protocone. The M2 is more or less reduced. The muzzle is elongated.
</paragraph>
</subSubSection>
</treatment>
</document>
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