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<document id="82068342A23175598A0877C686D84B3C" ID-CLB-Dataset="5365" ID-DOI="10.11646/zootaxa.3812.1.1" ID-GBIF-Dataset="fd5ba21e-09ce-4ac6-b84f-56632ed93917" ID-ISSN="1175-5326" ID-Zenodo-Dep="4919079" ID-ZooBank="7847D731-9F66-4856-A79F-9435FED25B1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623268636583" checkinUser="felipe" docAuthor="Murányi, Dávid, Gamboa, Maribet &amp; Orci, Kirill Márk" docDate="2014" docId="03F99336FF3AFFDB1BE45FC1D6FEFCDA" docLanguage="en" docName="zt03812p082.pdf" docOrigin="Zootaxa 3812 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Zwicknia tuberculata Murányi &amp; Gamboa &amp; Orci 2014, comb. n." docType="treatment" docVersion="6" lastPageNumber="53" masterDocId="FFC0EB4EFF08FFEF1B735B43D539FFDD" masterDocTitle="Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)" masterLastPageNumber="82" masterPageNumber="1" pageNumber="51" updateTime="1699007786927" updateUser="ExternalLinkService">
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<mods:title id="EE914DCD9070A16E1ABF8A0C417F94CB">Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)</mods:title>
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<mods:namePart id="24670438037C1472E7415FFC8E93C7F4">Murányi, Dávid</mods:namePart>
<mods:nameIdentifier id="C863E7817035D54754EE976D50229892" type="email">muranyi@zool.nhmus.hu</mods:nameIdentifier>
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<mods:namePart id="EC818B80EA4FCA69515B143AF3239879">Gamboa, Maribet</mods:namePart>
<mods:nameIdentifier id="D4A1B9CF4AE47ACEFA2FF698621B273C" type="email">maribetg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C6BA67C1763EDBB34DE26B75F394019D">Orci, Kirill Márk</mods:namePart>
<mods:nameIdentifier id="140E195FFC08EBFB50C0C5F5010A8331" type="email">muranyi@zool.nhmus.hu</mods:nameIdentifier>
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<mods:title id="CBDDFA0E19CC818D1E7E87BB2579A113">Zootaxa</mods:title>
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<mods:date id="693C5EA7CF4B008DF47FEDAB36620C60">2014</mods:date>
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<mods:number id="BB4082FC438E069138B141D985A2717B">2014-06-06</mods:number>
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<mods:number id="23E47A4F1D56D440684A42DE9183E744">3812</mods:number>
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<mods:classification id="FF85FF5A2CD7DCB0B998F5C7B1D4BE39">journal article</mods:classification>
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<mods:identifier id="5C5A8D9E06A4A0F208FDB7F28C4C67D6" type="DOI">10.11646/zootaxa.3812.1.1</mods:identifier>
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<treatment id="03F99336FF3AFFDB1BE45FC1D6FEFCDA" ID-DOI="http://doi.org/10.5281/zenodo.4919261" ID-GBIF-Taxon="183709170" ID-Zenodo-Dep="4919261" LSID="urn:lsid:plazi:treatment:03F99336FF3AFFDB1BE45FC1D6FEFCDA" httpUri="http://treatment.plazi.org/id/03F99336FF3AFFDB1BE45FC1D6FEFCDA" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
<subSubSection id="C34A71ABFF3AFFDD1BE45FC1D7DFFB40" box="[151,742,1154,1181]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="8BEF2220FF3AFFDD1BE45FC1D7DFFB40" blockId="50.[151,742,1154,1214]" box="[151,742,1154,1181]" pageId="50" pageNumber="51">
<heading id="D0A7954CFF3AFFDD1BE45FC1D7DFFB40" bold="true" box="[151,742,1154,1181]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="4">
<emphasis id="B924FE32FF3AFFDD1BE45FC1D7DFFB40" bold="true" box="[151,742,1154,1181]" pageId="50" pageNumber="51">
<taxonomicName id="4C5059A3FF3AFFDD1BE45FC1D74AFB40" ID-CoL="7GGKH" authority="(Zhiltzova, 1964)" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[151,627,1154,1181]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="tuberculata" status="comb. nov.">
<emphasis id="B924FE32FF3AFFDD1BE45FC1D4A1FB41" bold="true" box="[151,408,1154,1180]" italics="true" pageId="50" pageNumber="51">Zwicknia tuberculata</emphasis>
(
<bibRefCitation id="EFC15FD1FF3AFFDD1ADB5FC0D752FB40" author="Zhiltzova, L. A." box="[424,619,1155,1181]" pageId="50" pageNumber="51" pagination="347 - 362" refId="ref46420" refString="Zhiltzova, L. A. (1964) K poznaniu vesnianok (Plecoptera) Kavkaza. VI. Novie vidi Taeniopterygidae, Nemouridae i Capniidae. Revue d'Entomologie de l'USSR, 43 (2), 347 - 362." type="journal article" year="1964">Zhiltzova, 1964</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2174349FF3AFFDD19085FC0D7DFFB40" box="[635,742,1155,1181]" pageId="50" pageNumber="51" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF3AFFDD1BE45FE6D745FB63" box="[151,636,1189,1214]" pageId="50" pageNumber="51" type="description">
<paragraph id="8BEF2220FF3AFFDD1BE45FE6D745FB63" blockId="50.[151,742,1154,1214]" box="[151,636,1189,1214]" pageId="50" pageNumber="51">
(
<figureCitation id="136B3EA5FF3AFFDD1BEC5FE6D5C5FB63" box="[159,252,1189,1214]" captionStart="FIGURES 56–57" captionStartId="30.[151,264,1771,1794]" captionTargetBox="[217,1393,202,1724]" captionTargetId="figure-84@30.[176,1420,165,1752]" captionTargetPageId="30" captionText="FIGURES 56–57. Macropterous wings of Zwicknia Murányi, gen. n.—56: Female Z. bifrons (Newman, 1838); 57: male Z. tuberculata (Zhiltzova, 1964) (A: anal vein; a: anal crossvein; arc: arculus; C: costa; Cu: cubital vein; cu-a: cubitoanal crossvein; h: humeral crossvein; M: medial vein; m-cu: mediocubital crossvein; R: radial vein; r: radial crossvein; r-m: radiomedial crossvein; Sc: subcosta)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919106" httpUri="https://zenodo.org/record/4919106/files/figure.png" pageId="50" pageNumber="51">Figs. 57</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD1A7B5FE6D466FB63" box="[264,351,1189,1214]" captionStart="FIGURES 98–101" captionStartId="51.[151,264,1972,1995]" captionTargetBox="[242,1328,192,1954]" captionTargetId="figure-50@51.[235,1352,173,1972]" captionTargetPageId="51" captionText="FIGURES 98–101. Male terminalia of Zwicknia tuberculata (Zhiltzova, 1964); Kazakhstan, Jambyl Province, Karatau Mts., Bajaldyrtal—98: dorsal view; 99: ventral view; 100: lateral view; 101: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919128" httpUri="https://zenodo.org/record/4919128/files/figure.png" pageId="50" pageNumber="51">98–101</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD1A195FE6D4AFFB63" box="[362,406,1189,1214]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="50" pageNumber="51">112</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD1AD25FE6D4F4FB63" box="[417,461,1189,1214]" captionStart="FIGURES 124–126" captionStartId="58.[151,264,1502,1525]" captionTargetBox="[166,1399,232,1399]" captionTargetId="figure-159@58.[151,1436,223,1410]" captionTargetPageId="58" captionText="FIGURES 124–126. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—124: Z. sevanica (Zhiltzova, 1964); 125: Z. tuberculata (Zhiltzova, 1964); 126: Z. turkestanica turkestanica (Kimmins, 1950)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919144" httpUri="https://zenodo.org/record/4919144/files/figure.png" pageId="50" pageNumber="51">125</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD1AAA5FE6D73CFB63" box="[473,517,1189,1214]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="50" pageNumber="51">146</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD19625FE6D704FB63" box="[529,573,1189,1214]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="50" pageNumber="51">163</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD193A5FE6D74DFB63" box="[585,628,1189,1214]" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="50" pageNumber="51">196</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF3AFFDD1BE45FAED5D6FAFF" pageId="50" pageNumber="51" type="reference_group">
<paragraph id="8BEF2220FF3AFFDD1BE45FAED5D6FAFF" blockId="50.[151,1435,1261,1314]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB40050EFF3AFFDD1BE45FAED5D6FAFF" pageId="50" pageNumber="51">
<taxonomicName id="4C5059A3FF3AFFDD1BE45FAED736FADE" ID-CoL="QQTZ" authority="Zhiltzova, 1964" authorityName="Zhiltzova" authorityYear="1964" box="[151,527,1261,1283]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B924FE32FF3AFFDD1BE45FAED459FADE" box="[151,352,1261,1283]" italics="true" pageId="50" pageNumber="51">Capnia tuberculata</emphasis>
<bibRefCitation id="EFC15FD1FF3AFFDD1A1F5FAED736FADE" author="Zhiltzova, L. A." box="[364,527,1261,1283]" pageId="50" pageNumber="51" pagination="347 - 362" refId="ref46420" refString="Zhiltzova, L. A. (1964) K poznaniu vesnianok (Plecoptera) Kavkaza. VI. Novie vidi Taeniopterygidae, Nemouridae i Capniidae. Revue d'Entomologie de l'USSR, 43 (2), 347 - 362." type="journal article" year="1964">Zhiltzova, 1964</bibRefCitation>
</taxonomicName>
—
<treatmentCitation id="0AF10431FF3AFFDD195A5FAED63DFADE" author="Zhiltzova, L. A." box="[553,772,1261,1283]" page="359" pageId="50" pageNumber="51" year="1964">
<bibRefCitation id="EFC15FD1FF3AFFDD195A5FAED63DFADE" author="Zhiltzova, L. A." box="[553,772,1261,1283]" pageId="50" pageNumber="51" pagination="347 - 362" refId="ref46420" refString="Zhiltzova, L. A. (1964) K poznaniu vesnianok (Plecoptera) Kavkaza. VI. Novie vidi Taeniopterygidae, Nemouridae i Capniidae. Revue d'Entomologie de l'USSR, 43 (2), 347 - 362." type="journal article" year="1964">Zhiltzova 1964: 359</bibRefCitation>
</treatmentCitation>
. (description of the adult);
<treatmentCitation id="0AF10431FF3AFFDD1F5D5FAED1D1FADE" author="Zwick, P." box="[1070,1256,1261,1283]" page="391" pageId="50" pageNumber="51" year="1975">
<bibRefCitation id="EFC15FD1FF3AFFDD1F5D5FAED1D1FADE" author="Zwick, P." box="[1070,1256,1261,1283]" pageId="50" pageNumber="51" pagination="387 - 396" refId="ref46733" refString="Zwick, P. (1975) Weitere Plecoptera aus Anatolien. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 48 (3 / 4), 387 - 396." type="journal article" year="1975">Zwick 1975: 391</bibRefCitation>
</treatmentCitation>
. (first record for
<collectingCountry id="F34762B0FF3AFFDD1BE45E4FD5DDFAFF" box="[151,228,1292,1314]" name="Turkey" pageId="50" pageNumber="51">Turkey</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF3AFFDD1BE45E0AD585FA2F" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="8BEF2220FF3AFFDD1BE45E0AD585FA2F" blockId="50.[151,1437,1353,2026]" pageId="50" pageNumber="51">
<emphasis id="B924FE32FF3AFFDD1BE45E0AD428FABF" bold="true" box="[151,273,1353,1378]" pageId="50" pageNumber="51">Diagnosis.</emphasis>
Male epiproct: Ep-scl narrow and pointed in dorsal view, long, tip slightly downcurved in lateral view, membranous part long; ventral membranous section reach the base in lateral view, apical spines long, a few spines distributed also on the apex of Ep-scl. Process of male Tg 9: high, slightly folded backwards, &lt;2× wider than Epscl, rounded and with constantly narrow sides caudally. Female Sg: vaginal complex dark, transparent through the Sg.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF3AFFDD1BB45EBED4ECF983" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="8BEF2220FF3AFFDD1BB45EBED4ECF983" blockId="50.[151,1437,1353,2026]" pageId="50" pageNumber="51">
<emphasis id="B924FE32FF3AFFDD1BB45EBED48CF9CB" bold="true" box="[199,437,1533,1558]" pageId="50" pageNumber="51">Material examined.</emphasis>
<materialsCitation id="3B38287DFF3AFFDD1ACC5EBED4E9F983" ID-GBIF-Occurrence="3325453306" collectingDate="1970-04-28" collectionCode="PZC" collectorName="F. Klima" country="Kazakhstan" county="Bajaldyrtal" determinerName="P. Zwick" elevation="2" location="Jambyl Province" municipality="Karatau Mts." pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F34762B0FF3AFFDD1ACC5EBED7B8F9CB" box="[447,641,1533,1558]" name="Kazakhstan" pageId="50" pageNumber="51">KAZAKHSTAN</collectingCountry>
:
<location id="8E8F74FBFF3AFFDD19FC5EBED66AF9CB" LSID="urn:lsid:plazi:treatment:03F99336FF3AFFDB1BE45FC1D6FEFCDA:8E8F74FBFF3AFFDD19FC5EBED66AF9CB" box="[655,851,1533,1558]" country="Kazakhstan" county="Bajaldyrtal" municipality="Karatau Mts." name="Jambyl Province" pageId="50" pageNumber="51">Jambyl Province</location>
,
<collectingMunicipality id="6B8BB85AFF3AFFDD18125EBDD6C2F9C8" box="[865,1019,1534,1557]" pageId="50" pageNumber="51">Karatau Mts.</collectingMunicipality>
,
<collectingCounty id="628E5AACFF3AFFDD1F795EBED1B0F9CB" box="[1034,1161,1533,1558]" pageId="50" pageNumber="51">Bajaldyrtal</collectingCounty>
, 22–
<date id="FFEE04E0FF3AFFDD1FB05EBED07DF9CB" box="[1219,1348,1533,1558]" pageId="50" pageNumber="51" value="1970-04-28">
<collectingDate id="EFAAFD08FF3AFFDD1FB05EBED07DF9CB" box="[1219,1348,1533,1558]" pageId="50" pageNumber="51" value="1970-04-28">28.04.1970</collectingDate>
</date>
, leg.
<collectorName id="26A547F6FF3AFFDD1EFB5EBDD5D9F9E7" pageId="50" pageNumber="51">F. Klima</collectorName>
, det.
<determinerName id="7E33CEE4FF3AFFDD1A565D61D4A8F9E7" box="[293,401,1569,1594]" pageId="50" pageNumber="51">P. Zwick</determinerName>
:
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<elevation id="007DC513FF3AFFDD1AD35D62D4FAF9E7" box="[416,451,1569,1594]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="m" value="2.0">2m</elevation>
</quantity>
5f (
<collectionCode id="ED41BAE5FF3AFFDD1A885D62D70AF9E7" box="[507,563,1569,1594]" pageId="50" pageNumber="51">PZC</collectionCode>
; used for drawings
<figureCitation id="136B3EA5FF3AFFDD18515D62D6BCF9E7" box="[802,901,1569,1594]" captionStart="FIGURES 56–57" captionStartId="30.[151,264,1771,1794]" captionTargetBox="[217,1393,202,1724]" captionTargetId="figure-84@30.[176,1420,165,1752]" captionTargetPageId="30" captionText="FIGURES 56–57. Macropterous wings of Zwicknia Murányi, gen. n.—56: Female Z. bifrons (Newman, 1838); 57: male Z. tuberculata (Zhiltzova, 1964) (A: anal vein; a: anal crossvein; arc: arculus; C: costa; Cu: cubital vein; cu-a: cubitoanal crossvein; h: humeral crossvein; M: medial vein; m-cu: mediocubital crossvein; R: radial vein; r: radial crossvein; r-m: radiomedial crossvein; Sc: subcosta)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919106" httpUri="https://zenodo.org/record/4919106/files/figure.png" pageId="50" pageNumber="51">Figs. 57</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD18E65D62D6D2F9E7" box="[917,1003,1569,1594]" captionStart="FIGURES 98–101" captionStartId="51.[151,264,1972,1995]" captionTargetBox="[242,1328,192,1954]" captionTargetId="figure-50@51.[235,1352,173,1972]" captionTargetPageId="51" captionText="FIGURES 98–101. Male terminalia of Zwicknia tuberculata (Zhiltzova, 1964); Kazakhstan, Jambyl Province, Karatau Mts., Bajaldyrtal—98: dorsal view; 99: ventral view; 100: lateral view; 101: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919128" httpUri="https://zenodo.org/record/4919128/files/figure.png" pageId="50" pageNumber="51">98–101</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD188F5D62D111F9E7" box="[1020,1064,1569,1594]" captionStart="FIGURES 124–126" captionStartId="58.[151,264,1502,1525]" captionTargetBox="[166,1399,232,1399]" captionTargetId="figure-159@58.[151,1436,223,1410]" captionTargetPageId="58" captionText="FIGURES 124–126. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—124: Z. sevanica (Zhiltzova, 1964); 125: Z. tuberculata (Zhiltzova, 1964); 126: Z. turkestanica turkestanica (Kimmins, 1950)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919144" httpUri="https://zenodo.org/record/4919144/files/figure.png" pageId="50" pageNumber="51">125</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD1F4A5D62D15CF9E7" box="[1081,1125,1569,1594]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="50" pageNumber="51">146</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD1F055D62D19AF9E7" box="[1142,1187,1569,1594]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="50" pageNumber="51">163</figureCitation>
;
<specimenCount id="9D56E9A9FF3AFFDD1FC75D61D027F9E7" box="[1204,1310,1569,1594]" pageId="50" pageNumber="51" type="male">one male</specimenCount>
terminalia prepared for SEM
<figureCitation id="136B3EA5FF3AFFDD1A1B5D06D4F1F983" box="[360,456,1605,1630]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="50" pageNumber="51">Fig. 112</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF3AFFDB1BB45D2AD068FE17" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51" type="description">
<paragraph id="8BEF2220FF3AFFDD1BB45D2AD618F914" blockId="50.[151,1437,1353,2026]" pageId="50" pageNumber="51">
<emphasis id="B924FE32FF3AFFDD1BB45D2AD461F95F" bold="true" box="[199,344,1641,1666]" pageId="50" pageNumber="51">Description.</emphasis>
Head, thorax, appendages and basal segments of the abdomen generotypic. Males and females macropterous. Dimensions of the presently examined specimens: body length: males 6.0–6.5, females 7.5–9.0 mm; forewing length: males 5.5–6.0 mm, females 7.0–8.5 mm.
</paragraph>
<paragraph id="8BEF2220FF3AFFDD1BB45D96D058F837" blockId="50.[151,1437,1353,2026]" pageId="50" pageNumber="51">
Male terminalia (
<figureCitation id="136B3EA5FF3AFFDD1AFC5D96D711F933" box="[399,552,1749,1774]" captionStart="FIGURES 98–101" captionStartId="51.[151,264,1972,1995]" captionTargetBox="[242,1328,192,1954]" captionTargetId="figure-50@51.[235,1352,173,1972]" captionTargetPageId="51" captionText="FIGURES 98–101. Male terminalia of Zwicknia tuberculata (Zhiltzova, 1964); Kazakhstan, Jambyl Province, Karatau Mts., Bajaldyrtal—98: dorsal view; 99: ventral view; 100: lateral view; 101: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919128" httpUri="https://zenodo.org/record/4919128/files/figure.png" pageId="50" pageNumber="51">Figs. 98–101</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD19455D96D75DF933" box="[566,612,1749,1774]" captionStart="FIGURES 124–126" captionStartId="58.[151,264,1502,1525]" captionTargetBox="[166,1399,232,1399]" captionTargetId="figure-159@58.[151,1436,223,1410]" captionTargetPageId="58" captionText="FIGURES 124–126. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—124: Z. sevanica (Zhiltzova, 1964); 125: Z. tuberculata (Zhiltzova, 1964); 126: Z. turkestanica turkestanica (Kimmins, 1950)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919144" httpUri="https://zenodo.org/record/4919144/files/figure.png" pageId="50" pageNumber="51">125</figureCitation>
): Process of Tg 9 high, slightly fold backwards, its apex is about 1½× wider than the medial section of Ep-scl; rounded in shape, apex smooth; sides continuously narrow in caudal view, membranous portion narrowest apically (
<figureCitation id="136B3EA5FF3AFFDD191B5C5ED7F5F8EB" box="[616,716,1821,1846]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="50" pageNumber="51">Fig. 163</figureCitation>
). Tg 10, B-scl and Lb-scl generotypic. Ep-scl narrow and acute in dorsal view, medially not swollen, width medially &lt;½ of basal width; tip slightly downcurved in lateral view, divided section long. Ventral membranous part between the division of Ep-scl reaching base in lateral view; apical spines long, distributed not only on the membranous portion but spines extending to the Ep-scl (
<figureCitation id="136B3EA5FF3AFFDD1FA25CCAD004F87F" box="[1233,1341,1929,1954]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="50" pageNumber="51">Figs. 112</figureCitation>
,
<figureCitation id="136B3EA5FF3AFFDD1E3B5CCAD04FF87F" box="[1352,1398,1929,1954]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="50" pageNumber="51">146</figureCitation>
). Iscl generotypic, Ec long and contorted on the available specimens. St 9 not projecting medially, vesicle vestigial. Sg wide and rounded with not so pronounced triangular shape, tip rounded. Pp, Fp, Rp and cerci generotypic.
</paragraph>
<caption id="DF2F72A8FF3BFFDC1BE45CF7D14DF833" ID-DOI="http://doi.org/10.5281/zenodo.4919128" ID-Zenodo-Dep="4919128" httpUri="https://zenodo.org/record/4919128/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,264,1972,1995]" targetBox="[242,1328,192,1954]" targetPageId="51">
<paragraph id="8BEF2220FF3BFFDC1BE45CF7D14DF833" blockId="51.[151,1435,1972,2030]" pageId="51" pageNumber="52">
<emphasis id="B924FE32FF3BFFDC1BE45CF7D45BF817" bold="true" box="[151,354,1972,1995]" pageId="51" pageNumber="52">FIGURES 98–101.</emphasis>
Male terminalia of
<taxonomicName id="4C5059A3FF3BFFDC19425CF6D6FFF816" authority="(Zhiltzova, 1964)" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[561,966,1973,1995]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B924FE32FF3BFFDC19425CF6D630F816" box="[561,777,1973,1995]" italics="true" pageId="51" pageNumber="52">Zwicknia tuberculata</emphasis>
(
<bibRefCitation id="EFC15FD1FF3BFFDC186B5CF6D686F816" author="Zhiltzova, L. A." box="[792,959,1973,1995]" pageId="51" pageNumber="52" pagination="347 - 362" refId="ref46420" refString="Zhiltzova, L. A. (1964) K poznaniu vesnianok (Plecoptera) Kavkaza. VI. Novie vidi Taeniopterygidae, Nemouridae i Capniidae. Revue d'Entomologie de l'USSR, 43 (2), 347 - 362." type="journal article" year="1964">Zhiltzova, 1964</bibRefCitation>
)
</taxonomicName>
; Kazakhstan, Jambyl Province, Karatau Mts., Bajaldyrtal—98: dorsal view; 99: ventral view; 100: lateral view; 101: caudal view—scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BEF2220FF3CFFDB1BB45BD4D78CFEC7" blockId="52.[151,1437,151,775]" pageId="52" pageNumber="53">
Female subgenital plate: Rectangular, posterior margin nearly linear and sometimes slightly incised, equal to the segment`s posterior margin. Antero-lateral recess indistinct, the plate brown with darker longitudinal median lines caused by transparency of the unusually dark vaginal complex; lateral sclerites small (see
<bibRefCitation id="EFC15FD1FF3CFFDB1F9B5B9DD0A1FF2A" author="Zhiltzova, L. A." box="[1256,1432,222,247]" pageId="52" pageNumber="53" pagination="347 - 362" refId="ref46420" refString="Zhiltzova, L. A. (1964) K poznaniu vesnianok (Plecoptera) Kavkaza. VI. Novie vidi Taeniopterygidae, Nemouridae i Capniidae. Revue d'Entomologie de l'USSR, 43 (2), 347 - 362." type="journal article" year="1964">Zhiltzova 1964</bibRefCitation>
:
<figureCitation id="136B3EA5FF3CFFDB1BE45A42D5D1FEC7" box="[151,232,257,282]" captionStart="FIGURES 23–31" captionStartId="20.[151,264,1415,1438]" captionTargetBox="[201,1406,206,1387]" captionTargetId="figure-243@20.[177,1421,185,1401]" captionTargetPageId="20" captionText="FIGURES 23–31. Male paraprocts and fusion plate of Capniidae, dorso-lateral view—23: Zwicknia bifrons (Newman); 24: Capnia s.s. nigra (Pictet); 25: Capnia s.l. vidua Klapálek; 26: Capnia s.l. prolongata Zhiltzova; 27: Capnia s.l. pedestris Kimmins; 28: Capnia s.l. arensi Zhiltzova; 29: Bolshecapnia gregsoni (Ricker); 30: Isocapnia grandis (Banks); 31: Eucapnopsis brevicauda (Claassen) (Fp: fusion plate; Pp: paraproct; Rp: retractoral plate)—not to scale." figureDoi="http://doi.org/10.5281/zenodo.4919091" httpUri="https://zenodo.org/record/4919091/files/figure.png" pageId="52" pageNumber="53">Fig. 28</figureCitation>
, repeated as Fig.
<quantity id="4CA88FC5FF3CFFDB1ADF5A42D4CAFEC7" box="[428,499,257,282]" metricMagnitude="1" metricUnit="m" metricValue="1.47574" pageId="52" pageNumber="53" unit="in" value="581.0">581 in</quantity>
<bibRefCitation id="EFC15FD1FF3CFFDB1A8A5A42D790FEC7" author="Zhiltzova, L. A." box="[505,681,257,282]" pageId="52" pageNumber="53" pagination="1 - 538" refId="ref46559" refString="Zhiltzova, L. A. (2003) Plecoptera, Gruppe Euholognatha. Fauna of Russia and Neighbouring Countries, New Series, 145, 1 - 538." type="journal article" year="2003">Zhiltzova 2003</bibRefCitation>
).
</paragraph>
<paragraph id="8BEF2220FF3CFFDB1BB45A67D068FE17" blockId="52.[151,1437,151,775]" pageId="52" pageNumber="53">
<emphasis id="B924FE32FF3CFFDB1BB45A67D405FEE1" bold="true" box="[199,316,292,317]" pageId="52" pageNumber="53">Affinities.</emphasis>
The relationship of
<taxonomicName id="4C5059A3FF3CFFDB195B5A66D7F1FEE0" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[552,712,292,317]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B924FE32FF3CFFDB195B5A66D7F1FEE0" box="[552,712,292,317]" italics="true" pageId="52" pageNumber="53">Z. tuberculata</emphasis>
</taxonomicName>
with
<taxonomicName id="4C5059A3FF3CFFDB187F5A66D6B7FEE1" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[780,910,293,316]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sevanica">
<emphasis id="B924FE32FF3CFFDB187F5A66D6B7FEE1" box="[780,910,293,316]" italics="true" pageId="52" pageNumber="53">Z. sevanica</emphasis>
</taxonomicName>
is discussed above. The species is apparently nearer to
<taxonomicName id="4C5059A3FF3CFFDB1A725A0AD490FEBD" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Kimmins" baseAuthorityYear="1950" box="[257,425,327,352]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="turkestanica">
<emphasis id="B924FE32FF3CFFDB1A725A0AD490FEBD" box="[257,425,327,352]" italics="true" pageId="52" pageNumber="53">Z. turkestanica</emphasis>
</taxonomicName>
, but the males differ by having a more robust Ep-scl and Tg 9 process that is slightly fold backwards instead of perpendicularly elevated. In addition to the above species, males of
<taxonomicName id="4C5059A3FF3CFFDB1FF85A2FD010FE59" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[1163,1321,363,388]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B924FE32FF3CFFDB1FF85A2FD010FE59" box="[1163,1321,363,388]" italics="true" pageId="52" pageNumber="53">Z. tuberculata</emphasis>
</taxonomicName>
are easily distinguishable from other
<taxonomicName id="4C5059A3FF3CFFDB1ABB5ACDD717FE7A" authorityName="Muranyi" authorityYear="2014" box="[456,558,398,423]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF3CFFDB1ABB5ACDD717FE7A" box="[456,558,398,423]" italics="true" pageId="52" pageNumber="53">Zwicknia</emphasis>
</taxonomicName>
by the high, but rounded process of Tg 9. Unlike other
<taxonomicName id="4C5059A3FF3CFFDB1FD85ACDD057FE7A" box="[1195,1390,398,423]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B924FE32FF3CFFDB1FD85ACDD02BFE7A" box="[1195,1298,398,423]" italics="true" pageId="52" pageNumber="53">Zwicknia</emphasis>
species
</taxonomicName>
, the females are also distinct with their dark vaginal complex, transparent through the Sg. The larva is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF3CFFDB1BB45A97D4F4FD8A" pageId="52" pageNumber="53" type="distribution">
<paragraph id="8BEF2220FF3CFFDB1BB45A97D4F4FD8A" blockId="52.[151,1437,151,775]" pageId="52" pageNumber="53">
<emphasis id="B924FE32FF3CFFDB1BB45A97D4CBFE30" bold="true" box="[199,498,468,493]" pageId="52" pageNumber="53">Distribution and ecology.</emphasis>
The species was described from
<collectingCountry id="F34762B0FF3CFFDB181F5A97D6E8FE30" box="[876,977,468,493]" name="Armenia" pageId="52" pageNumber="53">Armenia</collectingCountry>
, known also from the Russian part of the Caucasus,
<collectingCountry id="F34762B0FF3CFFDB1A605AB4D454FDCD" box="[275,365,503,528]" name="Georgia" pageId="52" pageNumber="53">Georgia</collectingCountry>
and the Pontic Mts. of northern
<collectingCountry id="F34762B0FF3CFFDB19875AB4D66EFDCD" box="[756,855,503,528]" name="Turkey" pageId="52" pageNumber="53">Anatolia</collectingCountry>
, and herein reported from
<collectingCountry id="F34762B0FF3CFFDB1FE75AB4D02EFDCD" box="[1172,1303,503,528]" name="Kazakhstan" pageId="52" pageNumber="53">Kazakhstan</collectingCountry>
(
<figureCitation id="136B3EA5FF3CFFDB1E5A5AB4D0A9FDCD" box="[1321,1424,503,528]" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="52" pageNumber="53">Fig. 196</figureCitation>
). Adults are active from April to August but mostly in May near brooks to small rivers of mountainous areas between 1,000 and
<quantity id="4CA88FC5FF3CFFDB1A02597DD4F0FD8A" box="[369,457,574,599]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="52" pageNumber="53" unit="m" value="2.0">2,000m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF3CFFDB1BB45922D6FEFCDA" pageId="52" pageNumber="53" type="discussion">
<paragraph id="8BEF2220FF3CFFDB1BB45922D6FEFCDA" blockId="52.[151,1437,151,775]" pageId="52" pageNumber="53">
<emphasis id="B924FE32FF3CFFDB1BB45922D402FDA7" bold="true" box="[199,315,609,634]" pageId="52" pageNumber="53">Remarks.</emphasis>
The
<specimenCount id="9D56E9A9FF3CFFDB1A005921D4DCFDA7" box="[371,485,609,634]" pageId="52" pageNumber="53" type="male">two males</specimenCount>
and
<specimenCount id="9D56E9A9FF3CFFDB19695922D79AFDA7" box="[538,675,609,634]" pageId="52" pageNumber="53" type="female">five females</specimenCount>
listed above are morphologically agree with the original description of
<taxonomicName id="4C5059A3FF3CFFDB1BCA59C6D463FD40" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Zhiltzova" baseAuthorityYear="1964" box="[185,346,644,669]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B924FE32FF3CFFDB1BCA59C6D463FD40" box="[185,346,644,669]" italics="true" pageId="52" pageNumber="53">Z. tuberculata</emphasis>
</taxonomicName>
, especially in regard to the distinctive female, despite that this material is geographically far outside the known range of this species. However, drumming calls of the Asian
<taxonomicName id="4C5059A3FF3CFFDB1F6559E4D1EDFD1D" box="[1046,1236,679,704]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B924FE32FF3CFFDB1F6559E4D144FD1D" box="[1046,1149,679,704]" italics="true" pageId="52" pageNumber="53">Zwicknia</emphasis>
species
</taxonomicName>
are unknown and no fresh material is available for molecular studies. Their identification is presently based only on morphological characters and was not confirmed with drumming and molecular studies.
</paragraph>
</subSubSection>
</treatment>
</document>
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