165 lines
19 KiB
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165 lines
19 KiB
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<document id="8AA55BAD0AF2B06648D5257962E0A344" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629657024485" checkinUser="felipe" docAuthor="Daugeron, Christophe" docDate="2009" docId="A83687902D482060FF323D3EC5FAFA79" docLanguage="en" docName="zt02318p544.pdf" docOrigin="Zootaxa 2318" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empis (Kritempis) macropalpa Egger 1860" docType="treatment" docVersion="2" lastPageNumber="538" masterDocId="540FFFE82D4E2067FFA53B42C75CFFC8" masterDocTitle="Systematics of the Euro-Mediterranean Empis (Kritempis) (Diptera: Empididae: Empidinae) *" masterLastPageNumber="544" masterPageNumber="531" pageNumber="537" updateTime="1699090386252" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="1D0AFFE8011A487A62CCB60946CCD655">
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<mods:title id="D7BAAB88478602321B5AB64E4FBFF96B">Systematics of the Euro-Mediterranean Empis (Kritempis) (Diptera: Empididae: Empidinae) *</mods:title>
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<mods:namePart id="5C5C6782692C1E8EF599942836AA42B5">Daugeron, Christophe</mods:namePart>
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<mods:typeOfResource id="4B431E6C748A07A98E99DF76B0636EDD">text</mods:typeOfResource>
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<mods:title id="1C6750D1B82CD71894F9E28BBBEDA39E">Zootaxa</mods:title>
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<mods:part id="377DB736767579E58CFD9100510792C9">
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<mods:date id="B165A020E65DC3CA437EBCBB0AB89DD4">2009</mods:date>
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<mods:detail id="40F711E6E464FF82308CEC8374AC4272" type="pubDate">
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<mods:number id="4037A70B2CD76F875F52857BD1F754CD">2009-12-22</mods:number>
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<mods:detail id="0C2D615CE61759CD6CB38729C2A53036" type="volume">
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<mods:number id="31EA429650D82FA7C17F9886F5703FC5">2318</mods:number>
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<mods:start id="ED45AF0B12AC85B71AC37B2D1B9EA3A7">531</mods:start>
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<mods:classification id="5E54DB7456250C5183B4B5422D52A541">journal article</mods:classification>
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<mods:identifier id="62555CB9BC11D6F6552EA4B0C521A4F7" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="A83687902D482060FF323D3EC5FAFA79" LSID="urn:lsid:plazi:treatment:A83687902D482060FF323D3EC5FAFA79" httpUri="http://treatment.plazi.org/id/A83687902D482060FF323D3EC5FAFA79" lastPageId="7" lastPageNumber="538" pageId="6" pageNumber="537">
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<subSubSection id="6885650D2D482061FF323D3EC5FFF95D" box="[151,675,1659,1686]" pageId="6" pageNumber="537" type="nomenclature">
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<paragraph id="202036862D482061FF323D3EC5FFF95D" blockId="6.[151,675,1659,1720]" box="[151,675,1659,1686]" pageId="6" pageNumber="537">
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<heading id="7B6881EA2D482061FF323D3EC5FFF95D" bold="true" box="[151,675,1659,1686]" fontSize="11" level="1" pageId="6" pageNumber="537" reason="1">
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<taxonomicName id="E79F4D052D482061FF323D3EC5FFF95D" ID-CoL="8BZJ3" authority="Egger, 1860" authorityName="Egger" authorityYear="1860" box="[151,675,1659,1686]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="macropalpa" subGenus="Kritempis">
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<emphasis id="12EBEA942D482061FF323D3EC5FFF95D" bold="true" box="[151,675,1659,1686]" pageId="6" pageNumber="537">
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<emphasis id="12EBEA942D482061FF323D3EC7BFF95E" bold="true" box="[151,227,1660,1686]" italics="true" pageId="6" pageNumber="537">Empis</emphasis>
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(
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<emphasis id="12EBEA942D482061FF513D3EC635F95E" bold="true" box="[244,361,1660,1686]" italics="true" pageId="6" pageNumber="537">Kritempis</emphasis>
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)
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<emphasis id="12EBEA942D482061FEDF3D3EC555F95E" bold="true" box="[378,521,1660,1686]" italics="true" pageId="6" pageNumber="537">macropalpa</emphasis>
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<bibRefCitation id="440E4B772D482061FDB43D39C5FFF95D" author="Egger, J." box="[529,675,1659,1685]" pageId="6" pageNumber="537" refId="ref9132" refString="Egger, J. (1860) Dipterologische Beitrage. Verhandlungen des Zoologisch-Botanischen Gesellschaft in Wien, 10," type="book" year="1860">Egger, 1860</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6885650D2D482061FF323DDCC7B3F970" box="[151,239,1694,1720]" pageId="6" pageNumber="537" type="description">
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<paragraph id="202036862D482061FF323DDCC7B3F970" blockId="6.[151,675,1659,1720]" box="[151,239,1694,1720]" pageId="6" pageNumber="537">
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(
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<figureCitation id="B8A42A032D482061FF3A3DDCC7B9F970" box="[159,229,1694,1720]" captionStart="FIGURES 1–4" captionStartId="4.[151,269,1861,1885]" captionTargetBox="[156,1435,591,1835]" captionTargetId="figure-238@4.[151,1436,586,1835]" captionTargetPageId="4" captionText="FIGURES 1–4. Male hypopygium of Empis (Kritempis) spp. in lateral view, scale bars: 0.1 mm. Abbreviations: cerc = cercus; ej ap = ejaculatory apodeme; epn = epandrium; hyp = lateral arm of hypandrium; ph =phallus. 1. Empis (K.) algira Macquart (Algeria, Rouiba). 2. Empis (K) macquarti Becker (lectotype, Algeria). 3. Empis (K.) macropalpa Egger (lectotype, Sicily). 4. Empis (K.) nigrimana Becker (holotype, Algeria)." pageId="6" pageNumber="537">Fig. 3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="6885650D2D482061FF543DB0C436F8C1" box="[241,874,1778,1801]" pageId="6" pageNumber="537" type="reference_group">
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<paragraph id="202036862D482061FF543DB0C436F8C1" blockId="6.[241,874,1778,1801]" box="[241,874,1778,1801]" pageId="6" pageNumber="537">
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<treatmentCitationGroup id="008F11A82D482061FF543DB0C436F8C1" box="[241,874,1778,1801]" pageId="6" pageNumber="537">
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<treatmentCitation id="A13E10972D482061FF543DB0C527F8C1" author="Egger, J." box="[241,635,1778,1801]" page="344" pageId="6" pageNumber="537" year="1860">
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<taxonomicName id="E79F4D052D482061FF543DB0C527F8C1" ID-CoL="8BZJ3" authority="Egger, 1860: 344" authorityName="Egger" authorityPageNumber="344" authorityYear="1860" box="[241,635,1778,1801]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="species" species="macropalpa">
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<emphasis id="12EBEA942D482061FF543DB0C6E0F8C1" box="[241,444,1778,1801]" italics="true" pageId="6" pageNumber="537">Empis macropalpa</emphasis>
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<bibRefCitation id="440E4B772D482061FE673DB0C527F8C1" author="Egger, J." box="[450,635,1778,1801]" pageId="6" pageNumber="537" refId="ref9132" refString="Egger, J. (1860) Dipterologische Beitrage. Verhandlungen des Zoologisch-Botanischen Gesellschaft in Wien, 10," type="book" year="1860">Egger, 1860: 344</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="6885650D2D482060FF323C77C5FAFA79" lastPageId="7" lastPageNumber="538" pageId="6" pageNumber="537" type="diagnosis">
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<paragraph id="202036862D482061FF323C77C20BF857" blockId="6.[151,1436,1845,2029]" pageId="6" pageNumber="537">
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<emphasis id="12EBEA942D482061FF323C77C64BF887" bold="true" box="[151,279,1845,1871]" pageId="6" pageNumber="537">Diagnosis.</emphasis>
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Greyish species with brownish legs, labrum about as long as head height, presutural supraalar seta fine, M
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<subScript id="BC1B34C32D482061FF553C2EC7A4F8B2" attach="left" box="[240,248,1900,1914]" fontSize="6" pageId="6" pageNumber="537">2</subScript>
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abbreviated; male with scape and palpus with numerous rather long setae; female with mid femur with a few short posteroventral pennate setae apically, hind femur with ventral pennation on apical half.
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</paragraph>
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<paragraph id="202036862D482061FF633CEEC2C0F825" blockId="6.[151,1436,1845,2029]" pageId="6" pageNumber="537">
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<materialsCitation id="90F73CDB2D482061FF633CEEC2C0F825" location="In" municipality="The" pageId="6" pageNumber="537" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sicilia" typeStatus="lectotype">
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<emphasis id="12EBEA942D482061FF633CEEC687F80E" bold="true" box="[198,475,1964,1990]" pageId="6" pageNumber="537">
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<typeStatus id="FF2488242D482061FF633CEEC61EF80E" box="[198,322,1964,1990]" pageId="6" pageNumber="537" type="lectotype">Lectotype</typeStatus>
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designation
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</emphasis>
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.
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<collectingMunicipality id="C044ACFC2D482061FE483CEEC547F80E" box="[493,539,1964,1990]" pageId="6" pageNumber="537">The</collectingMunicipality>
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original description is based on both sexes from
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<collectingRegion id="E25BF8642D482061FBD13CEEC3E4F80E" box="[1140,1208,1964,1990]" country="Italy" name="Sicilia" pageId="6" pageNumber="537">Sicily</collectingRegion>
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but the number of specimens examined was not given by
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<bibRefCitation id="440E4B772D482061FDF83C91C5A0F825" author="Egger, J." box="[605,764,2003,2029]" pageId="6" pageNumber="537" refId="ref9132" refString="Egger, J. (1860) Dipterologische Beitrage. Verhandlungen des Zoologisch-Botanischen Gesellschaft in Wien, 10," type="book" year="1860">Egger (1860)</bibRefCitation>
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.
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<location id="2540605D2D482061FCA33C91C442F825" LSID="urn:lsid:plazi:treatment:A83687902D482060FF323D3EC5FAFA79:2540605D2D482061FCA33C91C442F825" box="[774,798,2003,2029]" municipality="The" name="In" pageId="6" pageNumber="537" stateProvince="Sicilia">In</location>
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the NMW collection,
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<specimenCount id="3699FD0F2D482061FB813C91C3C0F825" box="[1060,1180,2003,2029]" pageId="6" pageNumber="537" type="male">five males</specimenCount>
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and
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<specimenCount id="3699FD0F2D482061FB703C91C22DF825" box="[1237,1393,2003,2029]" pageId="6" pageNumber="537" type="female">three females</specimenCount>
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are
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</materialsCitation>
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</paragraph>
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<paragraph id="202036862D482061FF30337EC2FCF786" blockId="6.[149,1440,2098,2132]" box="[149,1440,2098,2132]" pageId="6" pageNumber="537">
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EURO-MEDITERRANEAN
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<taxonomicName id="E79F4D052D482061FE46337EC56AF79C" authorityName="Linnaeus" authorityYear="1758" box="[483,566,2108,2132]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="537" phylum="Arthropoda" rank="genus">EMPIS</taxonomicName>
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(KRITEMPIS)
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<emphasis id="12EBEA942D482061FC7A337AC372F785" box="[991,1070,2104,2125]" italics="true" pageId="6" pageNumber="537">Zootaxa</emphasis>
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2318 © 2009 Magnolia Press · 537
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</paragraph>
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<paragraph id="202036862D492060FF323BDAC378FEEE" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">
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<materialsCitation id="90F73CDB2D492060FF323BDAC3F5FF10" location="Specimens" pageId="7" pageNumber="538" specimenCount="1" stateProvince="Sicilia" typeStatus="syntype">
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present and all labelled in the same way: “Mann, 1858,
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<collectingRegion id="E25BF8642D492060FCEE3BDAC4C2FF7A" box="[843,926,152,178]" country="Italy" name="Sicilia" pageId="7" pageNumber="538">Sicilia</collectingRegion>
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;
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<taxonomicName id="E79F4D052D492060FC093BDAC38EFF7A" authority=", Alte Sammlung" authorityName="Alte Sammlung" box="[940,1234,152,178]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="genus">Empis, Alte Sammlung</taxonomicName>
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”.
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<location id="2540605D2D492060FB4F3BDAC232FF7A" LSID="urn:lsid:plazi:treatment:A83687902D482060FF323D3EC5FAFA79:2540605D2D492060FB4F3BDAC232FF7A" box="[1258,1390,152,178]" name="Specimens" pageId="7" pageNumber="538" stateProvince="Sicilia">Specimens</location>
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are conspecific and correspond to the original description and thus here considered
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<typeStatus id="FF2488242D492060FB9A3BFCC3F5FF10" box="[1087,1193,190,216]" pageId="7" pageNumber="538" type="syntype">syntypes</typeStatus>
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</materialsCitation>
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. A male specimen is herewith designated as
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<typeStatus id="FF2488242D492060FE1E3BA7C570FF37" box="[443,556,229,255]" pageId="7" pageNumber="538" type="lectotype">lectotype</typeStatus>
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<materialsCitation id="90F73CDB2D492060FD903BA7C37CFEEE" location="The" pageId="7" pageNumber="538" specimenCount="1" stateProvince="Sicilia" typeStatus="paralectotype">
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of the species and labelled accordingly to fix and stabilize the current concept of the name.
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<location id="2540605D2D492060FE373A4EC6E3FEEE" LSID="urn:lsid:plazi:treatment:A83687902D482060FF323D3EC5FAFA79:2540605D2D492060FE373A4EC6E3FEEE" box="[402,447,268,294]" name="The" pageId="7" pageNumber="538" stateProvince="Sicilia">The</location>
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remaining
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<specimenCount id="3699FD0F2D492060FDE03A4EC455FEEE" box="[581,777,268,294]" pageId="7" pageNumber="538" type="generic">seven specimens</specimenCount>
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become
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<typeStatus id="FF2488242D492060FCD63A4EC37CFEEE" box="[883,1056,268,294]" pageId="7" pageNumber="538" type="paralectotype">paralectotypes</typeStatus>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="202036862D492060FF633A70C6B1FE08" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">
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<emphasis id="12EBEA942D492060FF633A70C622FE84" bold="true" box="[198,382,306,332]" pageId="7" pageNumber="538">Re-description</emphasis>
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. Male. Head. Occiput dusty greyish, covered with fine, rather long black setae. Ocellar triangle with pair of fine setae. Scape dark brown with numerous long setae, pedicel dark brown, postpedicel black. Labrum dark brown, slightly longer than head height; labium blackish; palpus blackish, long, projected, with numerous distinct setae.
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</paragraph>
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<paragraph id="202036862D492060FF633A8FC5A3FD60" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">Thorax. Dusty greyish. Antepronotum with a few black setae. Antepronotal lobes with 1 rather fine, long black basal bristle, many other finer, short brownish to pale yellow anteriors. Proepisternum, prosternum with fan of pale yellow setae. Scutum with 3 blackish stripes on acrostichals and dorsocentrals; acrostichals quadrito biserial, fine. Dorsocentrals quadri- to biserial, fine, except for 3-4 strong, long prescutellars. Other strong, long setae as follows: 1 presutural, 1 postsutural supraalars; 3 notopleurals. Laterotergite with fan of long pale yellow setae. Anterior, posterior spiracles yellowish.</paragraph>
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<paragraph id="202036862D492060FF6339F7C3FDFD07" blockId="7.[151,1436,152,1457]" box="[198,1185,693,723]" pageId="7" pageNumber="538">
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Wing. Length about 6.7 mm. Feebly tinted of brown. M
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<subScript id="BC1B34C32D492060FCF13987C400FD1B" attach="left" box="[852,860,709,723]" fontSize="6" pageId="7" pageNumber="538">2</subScript>
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abbreviated. Halter yellow.
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</paragraph>
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<paragraph id="202036862D492060FF63399DC6F0FBCF" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">Legs. Coxae greyish with many pale yellow setae. Fore femur brownish; fore tibia brownish to yellowish basally with anterodorsal row of rather strong setae about as long as tibia depth, finer posterodorsals; fore tarsus dark brown, fore tarsomeres 1–2 with strong apical setae. Mid femur brownish; mid tibia brownish to yellowish basally, with antero- and posterodorsal rows of 3-5 strong, rather long setae, antero- and posteroventral rows of strong, shorter setae on apical half; mid tarsus dark brown with tarsomeres 1–4 with 2 ventral rows of spine-like setae. Hind femur brownish to yellowish apically; hind tibia brownish to yellowish basally, with antero- and posterodorsal rows of setae as long as tibia depth; hind tarsomeres dark brown with 2 ventral spine-like setae.</paragraph>
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<paragraph id="202036862D492060FF633F56C785FB9C" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">Abdomen. Tergites brownish to greyish at margins, sternites greyish, covered with rather long pale yellow setae.</paragraph>
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<paragraph id="202036862D492060FF633F23C33AFB6A" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">
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Hypopygium (
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<figureCitation id="B8A42A032D492060FED33F23C69EFBB3" box="[374,450,1121,1147]" captionStart="FIGURES 1–4" captionStartId="4.[151,269,1861,1885]" captionTargetBox="[156,1435,591,1835]" captionTargetId="figure-238@4.[151,1436,586,1835]" captionTargetPageId="4" captionText="FIGURES 1–4. Male hypopygium of Empis (Kritempis) spp. in lateral view, scale bars: 0.1 mm. Abbreviations: cerc = cercus; ej ap = ejaculatory apodeme; epn = epandrium; hyp = lateral arm of hypandrium; ph =phallus. 1. Empis (K.) algira Macquart (Algeria, Rouiba). 2. Empis (K) macquarti Becker (lectotype, Algeria). 3. Empis (K.) macropalpa Egger (lectotype, Sicily). 4. Empis (K.) nigrimana Becker (holotype, Algeria)." pageId="7" pageNumber="538">Fig. 3</figureCitation>
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). Cercus broad, somewhat square. Epandrial lamella somewhat lengthened with many strong, long apical setae. Phallus rather simple, gently curved, not undulated.
|
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|
</paragraph>
|
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|
<paragraph id="202036862D492060FF633FEDC6FAFAAB" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">
|
|||
|
Female. Similar to male except for following characters: scape, palpus with shorter setae. All setae on legs short; fore tibia yellowish to brownish apically; mid femur with a few posteroventral short pennate setae apically; mid tibia yellowish to brownish apically, with posteroventral group of
|
|||
|
<emphasis id="12EBEA942D492060FBFE3FBEC322FADD" box="[1115,1150,1276,1301]" italics="true" pageId="7" pageNumber="538">ca.</emphasis>
|
|||
|
10 very short spine-like setae; hind femur with numerous fine pale yellow ventral setae, ventral pennation on apical half. Wing clear. Abdominal setae short.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="202036862D492060FF633E32C5FAFA79" blockId="7.[151,1436,152,1457]" pageId="7" pageNumber="538">
|
|||
|
<emphasis id="12EBEA942D492060FF633E32C617FA42" bold="true" box="[198,331,1392,1418]" pageId="7" pageNumber="538">Chorotype</emphasis>
|
|||
|
. W-Mediterranean.
|
|||
|
<taxonomicName id="E79F4D052D492060FD9D3E32C449FA41" authorityName="Egger" authorityYear="1860" box="[568,789,1392,1417]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="538" phylum="Arthropoda" rank="species" species="macropalpa">
|
|||
|
<emphasis id="12EBEA942D492060FD9D3E32C449FA41" box="[568,789,1392,1417]" italics="true" pageId="7" pageNumber="538">Empis macropalpa</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
is known from
|
|||
|
<collectingRegion id="E25BF8642D492060FC753E32C349FA42" box="[976,1045,1392,1418]" country="Italy" name="Sicilia" pageId="7" pageNumber="538">Sicily</collectingRegion>
|
|||
|
, but not mentioned in the on-line checklist of the Italian fauna (
|
|||
|
<bibRefCitation id="440E4B772D492060FE563ED5C5C5FA79" author="Raffone, G." box="[499,665,1431,1457]" pageId="7" pageNumber="538" refId="ref9587" refString="Raffone, G. (2003) Diptera Microphoridae, Hybotidae, Empididae. In: Stoch, S. (Ed.) Checklist of the species of the Italian fauna on-line version 2.0 [Available at http: // www. faunaitalia. it / checklist / index. html] [accessed September 2009]." type="url" year="2003">Raffone 2003</bibRefCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|