148 lines
17 KiB
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148 lines
17 KiB
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<document id="C837D74F3FE8552F45FCAD97C3BC13C9" ID-DOI="10.5962/bhl.part.117802" ID-ISSN="0035-418X" ID-Zenodo-Dep="6311706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645021050174" checkinUser="carolina" docAuthor="Cuccodoro, Giulio" docDate="2011" docId="03F7D871AF7B5D51FF0FAD7175B6D9A7" docLanguage="en" docName="RevSuisseZool.118.107-148.pdf.imf" docOrigin="Revue suisse de Zoologie 118 (1)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Megarthrus flavosignatus Bierig 1940" docType="treatment" docVersion="4" lastPageNumber="113" masterDocId="FFCEA009AF7E5D57FF98AE43766CDA37" masterDocTitle="Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)" masterLastPageNumber="147" masterPageNumber="107" pageNumber="112" updateTime="1699296264638" updateUser="plazi">
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<mods:title id="EBA3C28970FD02197626BE3CB34B6C9D">Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)</mods:title>
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<mods:namePart id="4ABADF516486CBFB2D0479D0BD6483F1">Cuccodoro, Giulio</mods:namePart>
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<mods:typeOfResource id="03730577C473C4E1F51186D3B2E950CE">text</mods:typeOfResource>
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<mods:title id="91A5DDC12D769AE5262A77BB0488876A">Revue suisse de Zoologie</mods:title>
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<mods:part id="E2E4E3F84986AAE5A62A266453D7950E">
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<mods:date id="2C4BCE291CE1781D692BC8BC0B3A0014">2011</mods:date>
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<mods:number id="F2746C625BA159D1DCE81878E2CA2C66">2011-03-31</mods:number>
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<mods:number id="6CAF64F9439BE5502474F8813B65BB60">118</mods:number>
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<mods:detail id="7D8DB307A6A9A2B16E4A5C97ECC73620" type="issue">
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<mods:number id="11044C77BC6B5732225FE9AC8B35FA58">1</mods:number>
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<mods:start id="6CC64E846D847641F7556E26B43F43B4">107</mods:start>
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<mods:classification id="9E031DD4029A656426A215A61CC4C7D1">journal article</mods:classification>
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<mods:identifier id="2A52D6F1B231DA231FB44C2388CEF2C8" type="DOI">10.5962/bhl.part.117802</mods:identifier>
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<mods:identifier id="4D9801EB9B4FF976219830ADE324A523" type="ISSN">0035-418X</mods:identifier>
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<mods:identifier id="D2A82D5DFC28A42E5041636C22295C0A" type="Zenodo-Dep">6311706</mods:identifier>
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<treatment id="03F7D871AF7B5D51FF0FAD7175B6D9A7" ID-DOI="http://doi.org/10.5281/zenodo.6311365" ID-Zenodo-Dep="6311365" LSID="urn:lsid:plazi:treatment:03F7D871AF7B5D51FF0FAD7175B6D9A7" httpUri="http://treatment.plazi.org/id/03F7D871AF7B5D51FF0FAD7175B6D9A7" lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112">
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<subSubSection id="C3443AECAF7B5D52FF0FAD71725DD97D" box="[151,1073,818,842]" pageId="5" pageNumber="112" type="nomenclature">
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<paragraph id="8BE16967AF7B5D52FF0FAD71725DD97D" blockId="5.[151,1073,818,842]" box="[151,1073,818,842]" pageId="5" pageNumber="112">
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<heading id="D0A9DE0BAF7B5D52FF0FAD71725DD97D" box="[151,1073,818,842]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="112" reason="2">
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<taxonomicName id="4C5E12E4AF7B5D52FF0FAD71742DD97D" ID-CoL="3YX9R" authority="Bierig, 1940" authorityName="Bierig" authorityYear="1940" box="[151,577,818,842]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="flavosignatus">
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<emphasis id="B92AB575AF7B5D52FF0FAD7177D8D97D" bold="true" box="[151,436,818,842]" italics="true" pageId="5" pageNumber="112">Megarthrus flavosignatus</emphasis>
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<bibRefCitation id="EFCF1496AF7B5D52FE23AD70742DD97D" author="BIERIG, A." box="[443,577,819,842]" pageId="5" pageNumber="112" pagination="373 - 380" refId="ref12543" refString="BIERIG, A. 1940. Proteinini (Col. Staph.) Costaricenses. Revista de Entomologia 11: 373 - 380." type="journal article" year="1940">Bierig, 1940</bibRefCitation>
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</taxonomicName>
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Figs 5, 57-68
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3443AECAF7B5D52FF0FAD15742BD95C" box="[151,583,854,875]" pageId="5" pageNumber="112" type="reference_group">
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<paragraph id="8BE16967AF7B5D52FF0FAD15742BD95C" blockId="5.[151,583,854,875]" box="[151,583,854,875]" pageId="5" pageNumber="112">
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<treatmentCitationGroup id="AB4E4E49AF7B5D52FF0FAD15742BD95C" box="[151,583,854,875]" pageId="5" pageNumber="112">
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<treatmentCitation id="0AFF4F76AF7B5D52FF0FAD15742FD95C" author="BIERIG, A." box="[151,579,854,875]" page="378" pageId="5" pageNumber="112" year="1940">
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<taxonomicName id="4C5E12E4AF7B5D52FF0FAD15742FD95C" ID-CoL="3YX9R" authority="Bierig, 1940: 378" authorityName="Bierig" authorityPageNumber="378" authorityYear="1940" box="[151,579,854,875]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="flavosignatus">
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<emphasis id="B92AB575AF7B5D52FF0FAD1577E3D95C" box="[151,399,854,875]" italics="true" pageId="5" pageNumber="112">Megarthrus flavosignatus</emphasis>
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<bibRefCitation id="EFCF1496AF7B5D52FE0EAD15742FD95C" author="BIERIG, A." box="[406,579,854,875]" pageId="5" pageNumber="112" pagination="373 - 380" refId="ref12543" refString="BIERIG, A. 1940. Proteinini (Col. Staph.) Costaricenses. Revista de Entomologia 11: 373 - 380." type="journal article" year="1940">Bierig, 1940: 378</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3443AECAF7B5D52FF7BAD3475B6D9FD" pageId="5" pageNumber="112" type="materials_examined">
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<paragraph id="8BE16967AF7B5D52FF7BAD34772DD991" blockId="5.[151,1073,886,934]" pageId="5" pageNumber="112">
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<materialsCitation id="3B36633AAF7B5D52FF7BAD34772DD991" collectionCode="TYPE, MATERIAL, FMNH" country="Costa Rica" location="Carpintera" municipality="Typus" pageId="5" pageNumber="112" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED4FF1A2AF7B5D52FF7BAD347715D9BC" box="[227,377,887,908]" pageId="5" pageNumber="112">TYPE MATERIAL</collectionCode>
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:
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<typeStatus id="54E5D7C5AF7B5D52FE13AD347788D9BB" box="[395,484,887,908]" pageId="5" pageNumber="112" type="holotype">Holotype</typeStatus>
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(Ƌ, in
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<collectionCode id="ED4FF1A2AF7B5D52FDA2AD3474E8D9BB" box="[570,644,887,908]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="112" type="Museum">FMNH</collectionCode>
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): “
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<location id="8E813FBCAF7B5D52FD3AAD347567D9BB" LSID="urn:lsid:plazi:treatment:03F7D871AF7B5D51FF0FAD7175B6D9A7:8E813FBCAF7B5D52FD3AAD347567D9BB" box="[674,779,887,908]" country="Costa Rica" municipality="Typus" name="Carpintera" pageId="5" pageNumber="112">Carpintera</location>
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, 6.viii.39,
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<collectingCountry id="F34929F7AF7B5D52FCE2AD347584D9BB" box="[890,1000,887,908]" name="Costa Rica" pageId="5" pageNumber="112">Costa Rica</collectingCountry>
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[handwritten] |
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<collectingMunicipality id="6B85F31DAF7B5D52FF6DADD27758D991" box="[245,308,913,934]" pageId="5" pageNumber="112">Typus</collectingMunicipality>
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”.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE16967AF7B5D52FF7BADF675B6D9FD" blockId="5.[227,986,947,970]" box="[227,986,947,970]" pageId="5" pageNumber="112">
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<materialsCitation id="3B36633AAF7B5D52FF7BADF675E8D9FD" box="[227,900,947,970]" collectingDate="1940-06-16" collectionCode="MATERIAL" country="Costa Rica" location="Carpintera" municipality="Typus" pageId="5" pageNumber="112" specimenCount="1">
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ADDITIONAL MATERIAL: Same data as holotype, but
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<date id="FFE04FA7AF7B5D52FD48ADF67555D9FD" box="[720,825,949,970]" pageId="5" pageNumber="112" value="1940-06-16">
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<collectingDate id="EFA4B64FAF7B5D52FD48ADF67555D9FD" box="[720,825,949,970]" pageId="5" pageNumber="112" value="1940-06-16">16.vi.1940</collectingDate>
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</date>
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, 1 Ƌ in
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</materialsCitation>
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<collectionCode id="ED4FF1A2AF7B5D52FC12ADF675BBD9FD" box="[906,983,949,970]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="112" type="Museum">MHNG</collectionCode>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3443AECAF7B5D51FF7BAD9477CED8AD" lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112" type="description">
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<paragraph id="8BE16967AF7B5D52FF7BAD9474C4DF71" blockId="5.[151,1074,983,1662]" pageId="5" pageNumber="112">
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DESCRIPTION: Habitus as in Fig. 5. Combined length of pronotum and elytra =
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<quantity id="4CA6C482AF7B5D52FF0FADB4777ADE39" box="[151,278,1015,1038]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="5" pageNumber="112" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3-1.4 mm</quantity>
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; maximal pronotal width =
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<quantity id="4CA6C482AF7B5D52FDA5ADB474D0DE39" box="[573,700,1015,1038]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="5" pageNumber="112" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9-1.1 mm</quantity>
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. Body dark brown wih appendages slightly paler; antennomeres 10-11 paler than antennomeres 1-9. Dorsal pubescence denser on head and pronotum than on elytral disc, becoming denser along medial groove of pronotum and on anterior portion of elytral disc; frontal setae directed forward; elytral and pronotal setae slightly arcuate, recumbent; metasternal pubescence becoming denser anteriorly and medially, as long as or longer than that of prosternum; pubescence on abdominal tergites parallel, uniform; that on sternites IV-VIII uniform. Frons, pronotum, lateral portion of elytral disc and anterior portion of prohypomera granulate; frontal granulation conspicuous, with granula about as high as their diameter, or higher; central, adsutural and posterior area of elytral disc punctate, coarsely; metasternum coarsely granulofossulate.
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</paragraph>
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<paragraph id="8BE16967AF7B5D52FF7BAB0D725EDFF5" blockId="5.[151,1074,983,1662]" pageId="5" pageNumber="112">Frons forming above clypeus a sharp ridge, the latter finely evenly carinate, nearly straight in middle and laterally oblique in dorsal view; mesal portion of disc strongly evenly convex in lateral view; U-shaped frontal impression shallow. Temples strongly convex in dorsal view. Antennae (Fig. 61) 2.1-2.3 times longer than pronotum.</paragraph>
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<paragraph id="8BE16967AF7B5D52FF7BAB887565DC49" blockId="5.[151,1074,983,1662]" pageId="5" pageNumber="112">Pronotum (Fig. 68) with center strongly convex in frontal view; disc deeply depressed near middle of lateral edges, shallowly depressed along anterior and posterior margins; medial groove slightly arcuate in lateral view, deep, narrowed in middle; hypomera ridged from anterior margin to laterobasal angle, disc without pit. Prosternal medial ridge absent, or absent. Scutellum with anterior margin rounded and posterior margin slighly arcuate toward right-angled apex.</paragraph>
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<paragraph id="8BE16967AF785D51FF4CAEA07514DB0D" blockId="6.[136,1059,227,666]" pageId="6" pageNumber="113">Elytra gradually widened (Fig. 5); humeral callus low, moderately convex; disc with low swellings, moderately depressed posteriorly along lateral edges; the latter very finely carinate, finely denticulate, slightly sinuate in dorsal view.</paragraph>
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<paragraph id="8BE16967AF785D51FF4CAF007452DB4D" blockId="6.[136,1059,227,666]" pageId="6" pageNumber="113">Abdominal sternites II and III with medial processes as in Fig. 66, posterior portion of process of sternite III straight.</paragraph>
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<paragraph id="8BE16967AF785D51FF4CAFC07429D84D" blockId="6.[136,1059,227,666]" pageId="6" pageNumber="113">Male: Frontoclypeal area not modified. Protarsomeres 1 lacking tenent setae. Mesofemorar (Fig. 62) as long as metafemora. Mesotibiae (Fig. 60) shorter than metatibia (Fig. 59). Metatarsomere 1 about as long as combined length of metatarsomeres 2-4. Peg-like setae arranged in a single row on mesotrochanters (Fig. 62) and mesotibiae, arranged in double row on metatibiae, and absent from protrochanters, profemora, protibiae, mesofemora, metatrochanters and metafemora. Apex of abdominal tergite VIII as in Figs 63, 67. Sternite 8 as in Figs 64-65. Sternite IX lacking subbasal protuberance. Aedeagus as in Figs 57-58.</paragraph>
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<paragraph id="8BE16967AF785D51FF4CACC077CED8AD" blockId="6.[136,1059,227,666]" box="[212,418,643,666]" pageId="6" pageNumber="113">Female: Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C3443AECAF785D51FF4CACED7716D912" pageId="6" pageNumber="113" type="distribution">
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<paragraph id="8BE16967AF785D51FF4CACED7716D912" blockId="6.[136,1059,686,805]" pageId="6" pageNumber="113">
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<materialsCitation id="3B36633AAF785D51FF4CACED7719D912" collectorName="A. Bierig" country="Costa Rica" elevation="1800" location="Costa Rica" pageId="6" pageNumber="113" specimenCount="1" typeStatus="holotype">
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DISTRIBUTION AND NATURAL HISTORY: The species is known only from the type locality in
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<collectingCountry id="F34929F7AF785D51FE98AC8D7716D8D2" box="[256,378,718,741]" name="Costa Rica" pageId="6" pageNumber="113">Costa Rica</collectingCountry>
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. According to the original description,
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<collectorName id="26AB0CB1AF785D51FCB1AC8D75E3D8D2" box="[809,911,718,741]" pageId="6" pageNumber="113">A. Bierig</collectorName>
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collected the
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<typeStatus id="54E5D7C5AF785D51FF10ACAD7689D932" box="[136,229,750,773]" pageId="6" pageNumber="113" type="holotype">holotype</typeStatus>
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at
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<quantity id="4CA6C482AF785D51FE92ACAD7732D932" box="[266,350,750,773]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="113" unit="m" value="1800.0">
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<elevation id="00738E54AF785D51FE92ACAD7732D932" box="[266,350,750,773]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="113" unit="m" value="1800.0">1800 m</elevation>
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</quantity>
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from sifted leaf litter of a fallen tree in forest (together with the types of
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<taxonomicName id="4C5E12E4AF785D51FF7DAD4D7705D912" authorityName="Bierig" authorityYear="2011" box="[229,361,782,805]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="adelphus">
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<emphasis id="B92AB575AF785D51FF7DAD4D7705D912" box="[229,361,782,805]" italics="true" pageId="6" pageNumber="113">M. adelphus</emphasis>
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</taxonomicName>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3443AECAF785D51FF4CAD7A75B6D9A7" pageId="6" pageNumber="113" type="discussion">
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<paragraph id="8BE16967AF785D51FF4CAD7A75B6D9A7" blockId="6.[136,1059,825,912]" pageId="6" pageNumber="113">
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COMMENTS: The male abdominal sternite VIII with a pair of projecting tips of
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<taxonomicName id="4C5E12E4AF785D51FF10AD1A7756D947" authorityName="Bierig" authorityYear="1940" box="[136,314,857,880]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="flavosignatus">
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<emphasis id="B92AB575AF785D51FF10AD1A7756D947" box="[136,314,857,880]" italics="true" pageId="6" pageNumber="113">M. flavosignatus</emphasis>
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</taxonomicName>
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is particularly notable. This feature is shared only with
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<taxonomicName id="4C5E12E4AF785D51FC17AD1A7277D947" authorityName="Sharp. These" authorityYear="1887" box="[911,1051,857,880]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="zunilensis">
|
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<emphasis id="B92AB575AF785D51FC17AD1A7277D947" box="[911,1051,857,880]" italics="true" pageId="6" pageNumber="113">M. zunilensis</emphasis>
|
||
|
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</taxonomicName>
|
||
|
|
, which has different male sexual characters. See comments under
|
||
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<taxonomicName id="4C5E12E4AF785D51FCDEAD3A75BFD9A7" authorityName="Sharp. These" authorityYear="1887" box="[838,979,889,912]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="zunilensis">
|
||
|
|
<emphasis id="B92AB575AF785D51FCDEAD3A75BFD9A7" box="[838,979,889,912]" italics="true" pageId="6" pageNumber="113">M. zunilensis</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
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</document>
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