330 lines
39 KiB
XML
330 lines
39 KiB
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<document id="0F76C891B8246DBC0262DE2175E46080" ID-CLB-Dataset="29788" ID-DOI="10.11646/zootaxa.4441.3.12" ID-GBIF-Dataset="58c8ec85-77bc-49f4-ada7-3c923d1ec269" ID-ISSN="1175-5326" ID-Zenodo-Dep="1301560" ID-ZooBank="B2D326BE-54E6-4E6E-9DF6-00220F04EC41" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530166891776" checkinUser="plazi" docAuthor="Sazhnev, Alexey S." docDate="2018" docId="03DE878EFFCFFFCEFF17DE66FB5CE553" docLanguage="en" docName="zootaxa.4441.3.12.pdf" docOrigin="Zootaxa 4441 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterocerus fausti Reitter 1879" docType="treatment" docVersion="6" lastPageNumber="600" masterDocId="FFE7FFF6FFCFFFCDFF80D938FF94E612" masterDocTitle="Redescription of Heterocerus fausti Reitter, 1879, bona species (Coleoptera, Heteroceridae)" masterLastPageNumber="600" masterPageNumber="597" pageNumber="597" updateTime="1698551529617" updateUser="ExternalLinkService">
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<mods:titleInfo id="CE30BE7D127F9617FF9E6A7B62F9628A">
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<mods:title id="F00883F716E0054B00917C6FE71AB426">Redescription of Heterocerus fausti Reitter, 1879, bona species (Coleoptera, Heteroceridae)</mods:title>
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<mods:name id="1826986537DD65A865F20085F89FA3DA" type="personal">
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<mods:namePart id="B141196C84E8D9067B22875090A3327D">Sazhnev, Alexey S.</mods:namePart>
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<mods:typeOfResource id="E187515210CEFF6D2045B7A0F471BFB9">text</mods:typeOfResource>
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<mods:title id="A4C17D2D2B20CE2FBF2F3B7AF0987A26">Zootaxa</mods:title>
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<mods:part id="528B28514C24126400D6151CFA60C16E">
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<mods:date id="25138160AFE2CC1E4744C40F2CD53B54">2018</mods:date>
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<mods:number id="BC7CDC8F01C17FA14E7C7FB31658C9D4">2018-06-28</mods:number>
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<mods:number id="565326E393FF5EDCB8ED2DC606076117">4441</mods:number>
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<mods:detail id="493D9C53661CD17B41EE764602508AF7" type="issue">
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<mods:number id="49919E5D73082776FFD2845FC10F1DA6">3</mods:number>
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<mods:classification id="06BF0DF9C4AE23F8060EBEBAA6CED13E">journal article</mods:classification>
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<mods:identifier id="C9CFF82B1457458EBE32103146AE42ED" type="CLB-Dataset">29788</mods:identifier>
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<mods:identifier id="1ECCC8EDE8A73D2026DFFE1F7F464A05" type="DOI">10.11646/zootaxa.4441.3.12</mods:identifier>
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<mods:identifier id="8D45DC2327A134DECDE16AAC9D71F179" type="GBIF-Dataset">58c8ec85-77bc-49f4-ada7-3c923d1ec269</mods:identifier>
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<mods:identifier id="9FC63F7DC660DD954428AD2350A7B2D3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DE878EFFCFFFCEFF17DE66FB5CE553" ID-DOI="http://doi.org/10.5281/zenodo.5989271" ID-GBIF-Taxon="150871423" ID-Zenodo-Dep="5989271" LSID="urn:lsid:plazi:treatment:03DE878EFFCFFFCEFF17DE66FB5CE553" httpUri="http://treatment.plazi.org/id/03DE878EFFCFFFCEFF17DE66FB5CE553" lastPageId="3" lastPageNumber="600" pageId="0" pageNumber="597">
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<subSubSection id="C36D6513FFCFFFCDFF17DE66FD4AE16A" box="[151,734,1886,1912]" pageId="0" pageNumber="597" type="nomenclature">
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<paragraph id="8BC83698FFCFFFCDFF17DE66FD4AE16A" blockId="0.[151,1436,1886,2008]" box="[151,734,1886,1912]" pageId="0" pageNumber="597">
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<heading id="D08081F4FFCFFFCDFF17DE66FD4AE16A" bold="true" box="[151,734,1886,1912]" fontSize="11" level="1" pageId="0" pageNumber="597" reason="1">
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<emphasis id="B903EA8AFFCFFFCDFF17DE66FD4AE16A" bold="true" box="[151,734,1886,1912]" pageId="0" pageNumber="597">
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<taxonomicName id="4C774D1BFFCFFFCDFF17DE66FD88E16A" authority="Reitter, 1879" authorityName="Reitter" authorityYear="1879" box="[151,540,1886,1912]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="fausti">
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<emphasis id="B903EA8AFFCFFFCDFF17DE66FEE2E16A" bold="true" box="[151,374,1886,1912]" italics="true" pageId="0" pageNumber="597">Heterocerus fausti</emphasis>
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<bibRefCitation id="EFE64B69FFCFFFCDFEFDDE66FD88E16A" box="[381,540,1886,1912]" pageId="0" pageNumber="597" refId="ref1605" refString="Reitter, E. (1879) Neue Coleopteren aus dem sudostlichen Russland aufgefunden von Herrn J. Faust aus Helsingfors. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 29, 543 - 546. [in German]" type="journal article">Reitter, 1879</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="134C2A1DFFCFFFCDFDB3DE66FD40E16A" box="[563,724,1886,1912]" captionStart="FIGURE 1–4" captionStartId="1.[151,244,1869,1891]" captionTargetBox="[151,1436,431,1852]" captionTargetId="figure@1.[151,1436,431,1852]" captionTargetPageId="1" captionText="FIGURE 1–4. Heterocerus fausti Reitter, 1879, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers Heterocerus spp. 2A—H. parallelus Gebler, 1830, South Kazakhstan; 2B—H. fausti, Turkmenistan; 2C—H. fossor Kiesenwetter, 1843, Russia, Dagestan; 2D—H. flexuosus Stephens, 1828, Kazakhstan; 2E—H. persicus Mascagni, 1989, Iran. Male genitalia of Heterocerus fausti, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens Heterocerus spp. 4A—H. parallelus, South Kazakhstan; 4B—H. fausti, Caucasus; 4C—H. fossor, Russia, Dagestan; 4D—H. flexuosus, European part of Russia (Litovkin et al., 2013), 4E— H. persicus, Iran. Scale bar—0.5 mm." httpUri="https://zenodo.org/record/1301562/files/figure.png" pageId="0" pageNumber="597">Figs. 1, 2B, 3</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6513FFCFFFCDFF17DEB9FDCFE184" box="[151,603,1920,1942]" pageId="0" pageNumber="597" type="reference_group">
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<paragraph id="8BC83698FFCFFFCDFF17DEB9FDCFE184" blockId="0.[151,1436,1886,2008]" box="[151,603,1920,1942]" pageId="0" pageNumber="597">
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<treatmentCitationGroup id="AB6711B6FFCFFFCDFF17DEB9FDCFE184" box="[151,603,1920,1942]" pageId="0" pageNumber="597">
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<taxonomicName id="4C774D1BFFCFFFCDFF17DEB9FE6EE184" authority="Reitter, 1879: 546" authorityName="Reitter" authorityPageNumber="546" authorityYear="1879" box="[151,506,1921,1942]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="fausti" status="stat. nov.">
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<emphasis id="B903EA8AFFCFFFCDFF17DEB9FED3E184" box="[151,327,1921,1942]" italics="true" pageId="0" pageNumber="597">Heterocerus fausti</emphasis>
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<treatmentCitation id="0AD61089FFCFFFCDFECEDEB9FE6EE184" author="" box="[334,506,1921,1942]" page="546" pageId="0" pageNumber="597">
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<bibRefCitation id="EFE64B69FFCFFFCDFECEDEB9FE5AE184" box="[334,462,1921,1942]" pageId="0" pageNumber="597" refId="ref1605" refString="Reitter, E. (1879) Neue Coleopteren aus dem sudostlichen Russland aufgefunden von Herrn J. Faust aus Helsingfors. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 29, 543 - 546. [in German]" type="journal article">Reitter, 1879</bibRefCitation>
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: 546
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</treatmentCitation>
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</taxonomicName>
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<taxonomicNameLabel id="A23057F1FFCFFFCDFD80DEB8FDCFE184" box="[512,603,1920,1942]" pageId="0" pageNumber="597">
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<emphasis id="B903EA8AFFCFFFCDFD80DEB8FDCFE184" bold="true" box="[512,603,1920,1942]" pageId="0" pageNumber="597">stat. nov.</emphasis>
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</taxonomicNameLabel>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6513FFCFFFCCFF17DEA6FB8CE72B" lastPageId="1" lastPageNumber="598" pageId="0" pageNumber="597" type="materials_examined">
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<paragraph id="8BC83698FFCFFFCDFF17DEA6FE63E1A7" blockId="0.[151,1436,1886,2008]" box="[151,503,1950,1973]" pageId="0" pageNumber="597">
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<materialsCitation id="3B1F3CC5FFCFFFCDFF17DEA6FE63E1A7" ID-GBIF-Occurrence="1977784429" box="[151,503,1950,1973]" country="Azerbaijan" location="Baku" pageId="0" pageNumber="597" specimenCount="1" stateProvince="Baku" typeStatus="holotype">
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<emphasis id="B903EA8AFFCFFFCDFF17DEA6FEB8E1A7" bold="true" box="[151,300,1950,1973]" pageId="0" pageNumber="597">
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<typeStatus id="54CC883AFFCFFFCDFF17DEA6FF5BE1A7" box="[151,207,1950,1973]" pageId="0" pageNumber="597">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3607608FFCFFFCDFEB4DEA6FE39E1A7" box="[308,429,1950,1973]" name="Azerbaijan" pageId="0" pageNumber="597">Azerbaijan</collectingCountry>
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:
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<collectingRegion id="49B3F87AFFCFFFCDFE37DEA6FE67E1A7" box="[439,499,1950,1973]" country="Azerbaijan" name="Baku" pageId="0" pageNumber="597">Baku</collectingRegion>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC83698FFCFFFCDFF47DEF9FA08E1CA" blockId="0.[151,1436,1886,2008]" box="[199,1436,1985,2008]" pageId="0" pageNumber="597">
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<emphasis id="B903EA8AFFCFFFCDFF47DEF9FE70E1CA" bold="true" box="[199,484,1985,2008]" pageId="0" pageNumber="597">
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<typeStatus id="54CC883AFFCFFFCDFF47DEF9FF6BE1CA" box="[199,255,1985,2008]" pageId="0" pageNumber="597">Type</typeStatus>
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material examined.
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</emphasis>
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<typeStatus id="54CC883AFFCFFFCDFE6CDEF9FDC6E1CA" box="[492,594,1985,2008]" pageId="0" pageNumber="597" type="syntype">Syntypes</typeStatus>
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: “
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<taxonomicName id="4C774D1BFFCFFFCDFDEADEF9FCF1E1CA" authority="Fausti Rtt" authorityName="Fausti Rtt" box="[618,869,1985,2008]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EA8AFFCFFFCDFDEADEF9FCADE1CA" box="[618,825,1985,2008]" italics="true" pageId="0" pageNumber="597">Heterocerus Fausti</emphasis>
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Rtt
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</taxonomicName>
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” [Reitter's handwriting], “Caucasus” [red label], “к.
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</paragraph>
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<paragraph id="8BC83698FFCEFFCCFF17D9AFFC60E6C3" blockId="1.[151,1436,151,383]" pageId="1" pageNumber="598">
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А. Яковлева” [cоllеctiоn оf А. Jаcоvlеv—in Russiаn], 2♂♂ (ZISP); “Bаku Fаust.”, gоldеn squаrе plаtе, “
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<taxonomicName id="4C774D1BFFCEFFCCFA99D9A0FE8BE6C3" authority="Fausti Reitt." authorityName="Fausti Reitt." class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EA8AFFCEFFCCFA99D9A0FF4FE6C3" italics="true" pageId="1" pageNumber="598">Heterocerus Fausti</emphasis>
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Reitt.
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</taxonomicName>
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” [Reitter's handwriting], 2♀♀ [one specimen without head] (ZISP).
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</paragraph>
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<paragraph id="8BC83698FFCEFFCCFF47D9E5FB8CE72B" blockId="1.[151,1436,151,383]" pageId="1" pageNumber="598">
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<materialsCitation id="3B1F3CC5FFCEFFCCFF47D9E5FC1BE705" ID-GBIF-Occurrence="1977784428" collectingDate="1895-04-15" collectionCode="ZISP" collectorName="Varentsov" country="Turkmenistan" county="Velayat" location="Hazar Nature Reserve" municipality="Uzun-Ada" pageId="1" pageNumber="598" specimenCount="1" stateProvince="Balkan">
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<emphasis id="B903EA8AFFCEFFCCFF47D9E5FDB7E6E6" bold="true" box="[199,547,221,244]" pageId="1" pageNumber="598">Additional material examined.</emphasis>
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9 exs.
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<emphasis id="B903EA8AFFCEFFCCFDEED9E5FCC4E6E6" bold="true" box="[622,848,221,244]" pageId="1" pageNumber="598">
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<collectingCountry id="F3607608FFCEFFCCFDEED9E5FCDFE6E6" box="[622,843,221,244]" name="Turkmenistan" pageId="1" pageNumber="598">TURKMENISTAN</collectingCountry>
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.
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</emphasis>
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<emphasis id="B903EA8AFFCEFFCCFCD6D9E5FC60E6E6" box="[854,1012,221,244]" italics="true" pageId="1" pageNumber="598">
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<collectingRegion id="49B3F87AFFCEFFCCFCD6D9E5FC35E6E6" box="[854,929,221,244]" country="Turkmenistan" name="Balkan Welayaty" pageId="1" pageNumber="598">Balkan</collectingRegion>
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Region
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</emphasis>
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(
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<collectingCounty id="62A94E14FFCEFFCCFB84D9E5FBDBE6E6" box="[1028,1103,221,244]" pageId="1" pageNumber="598">
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<emphasis id="B903EA8AFFCEFFCCFB84D9E5FBDBE6E6" box="[1028,1103,221,244]" italics="true" pageId="1" pageNumber="598">Velayat</emphasis>
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</collectingCounty>
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):
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<collectingMunicipality id="6BACACE2FFCEFFCCFBEAD9E5FB4DE6E6" box="[1130,1241,221,244]" pageId="1" pageNumber="598">Uzun-Ada</collectingMunicipality>
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(now the territory of
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<location id="8EA86043FFCEFFCCFF37D838FE3AE705" LSID="urn:lsid:plazi:treatment:03DE878EFFCFFFCEFF17DE66FB5CE553:8EA86043FFCEFFCCFF37D838FE3AE705" box="[183,430,256,279]" country="Turkmenistan" county="Velayat" municipality="Uzun-Ada" name="Hazar Nature Reserve" pageId="1" pageNumber="598" stateProvince="Balkan">Hazar Nature Reserve</location>
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),
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<collectingDate id="EF8DE9B0FFCEFFCCFE43D838FDAAE705" box="[451,574,256,279]" pageId="1" pageNumber="598" value="1895-04-15">
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<date id="FFC91058FFCEFFCCFE43D838FDAAE705" box="[451,574,256,279]" pageId="1" pageNumber="598" value="1895-04-15">15.04.1895</date>
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</collectingDate>
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, leg.
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<collectorName id="2682534EFFCEFFCCFDFED838FD7EE705" box="[638,746,256,279]" pageId="1" pageNumber="598">Varentsov</collectorName>
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, 7 exs. (
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<collectionCode id="ED66AE5DFFCEFFCCFCCBD838FC13E705" box="[843,903,256,279]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="598">ZISP</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B1F3CC5FFCEFFCCFC1DD838FE57E72B" ID-GBIF-Occurrence="1977784427" collectionCode="ZISP" collectorName="P. Maksimovitsh" country="Turkmenistan" location="Krasnovodsk" pageId="1" pageNumber="598" specimenCount="1" stateProvince="Balkan">
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Turkmenbashi (
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<location id="8EA86043FFCEFFCCFBC8D838FB4FE705" LSID="urn:lsid:plazi:treatment:03DE878EFFCFFFCEFF17DE66FB5CE553:8EA86043FFCEFFCCFBC8D838FB4FE705" box="[1096,1243,256,279]" country="Turkmenistan" name="Krasnovodsk" pageId="1" pageNumber="598" stateProvince="Balkan">Krasnovodsk</location>
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), 0 6.1900, leg.
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<collectorName id="2682534EFFCEFFCCFA09D838FEA6E72B" pageId="1" pageNumber="598">P. Maksimovitsh</collectorName>
|
|||
|
|
, 1 ex. (
|
|||
|
|
<collectionCode id="ED66AE5DFFCEFFCCFEFFD81AFE2FE72B" box="[383,443,290,313]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="598">ZISP</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
; Esenguly, 3.07.1932, unknown collector, 1 ex. (IBIW).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C36D6513FFCEFFCFFF47D87DFC0CE193" lastPageId="2" lastPageNumber="599" pageId="1" pageNumber="598" type="diagnosis">
|
|||
|
|
<paragraph id="8BC83698FFCEFFCCFF47D87DFDF9E76D" blockId="1.[151,1436,151,383]" pageId="1" pageNumber="598">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFF47D87DFEA8E74E" bold="true" box="[199,316,325,348]" pageId="1" pageNumber="598">Diagnosis.</emphasis>
|
|||
|
|
Body size 3.8–5.0 mm. Body moderately elongated, dorsal-ventrally compressed (
|
|||
|
|
<figureCitation id="134C2A1DFFCEFFCCFB4FD87DFA84E74E" box="[1231,1296,325,348]" captionStart="FIGURE 1–4" captionStartId="1.[151,244,1869,1891]" captionTargetBox="[151,1436,431,1852]" captionTargetId="figure@1.[151,1436,431,1852]" captionTargetPageId="1" captionText="FIGURE 1–4. Heterocerus fausti Reitter, 1879, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers Heterocerus spp. 2A—H. parallelus Gebler, 1830, South Kazakhstan; 2B—H. fausti, Turkmenistan; 2C—H. fossor Kiesenwetter, 1843, Russia, Dagestan; 2D—H. flexuosus Stephens, 1828, Kazakhstan; 2E—H. persicus Mascagni, 1989, Iran. Male genitalia of Heterocerus fausti, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens Heterocerus spp. 4A—H. parallelus, South Kazakhstan; 4B—H. fausti, Caucasus; 4C—H. fossor, Russia, Dagestan; 4D—H. flexuosus, European part of Russia (Litovkin et al., 2013), 4E— H. persicus, Iran. Scale bar—0.5 mm." httpUri="https://zenodo.org/record/1301562/files/figure.png" pageId="1" pageNumber="598">Fig. 1</figureCitation>
|
|||
|
|
). Males and females do not differ. Aedeagus as in
|
|||
|
|
<figureCitation id="134C2A1DFFCEFFCCFDA9D850FDFDE76D" box="[553,617,360,383]" captionStart="FIGURE 1–4" captionStartId="1.[151,244,1869,1891]" captionTargetBox="[151,1436,431,1852]" captionTargetId="figure@1.[151,1436,431,1852]" captionTargetPageId="1" captionText="FIGURE 1–4. Heterocerus fausti Reitter, 1879, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers Heterocerus spp. 2A—H. parallelus Gebler, 1830, South Kazakhstan; 2B—H. fausti, Turkmenistan; 2C—H. fossor Kiesenwetter, 1843, Russia, Dagestan; 2D—H. flexuosus Stephens, 1828, Kazakhstan; 2E—H. persicus Mascagni, 1989, Iran. Male genitalia of Heterocerus fausti, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens Heterocerus spp. 4A—H. parallelus, South Kazakhstan; 4B—H. fausti, Caucasus; 4C—H. fossor, Russia, Dagestan; 4D—H. flexuosus, European part of Russia (Litovkin et al., 2013), 4E— H. persicus, Iran. Scale bar—0.5 mm." httpUri="https://zenodo.org/record/1301562/files/figure.png" pageId="1" pageNumber="598">Fig. 3</figureCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF086610FFCEFFCCFF17DE75FD97E1E7" httpUri="https://zenodo.org/record/1301562/files/figure.png" pageId="1" pageNumber="598" startId="1.[151,244,1869,1891]" targetBox="[151,1436,431,1852]" targetPageId="1">
|
|||
|
|
<paragraph id="8BC83698FFCEFFCCFF17DE75FD97E1E7" blockId="1.[151,1436,1869,2037]" pageId="1" pageNumber="598">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFF17DE75FEB2E171" bold="true" box="[151,294,1869,1891]" pageId="1" pageNumber="598">FIGURE 1–4.</emphasis>
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFEAFDE75FDF8E170" authority="Reitter, 1879" authorityName="Reitter" authorityYear="1879" box="[303,620,1869,1890]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFEAFDE75FE76E170" box="[303,482,1869,1890]" italics="true" pageId="1" pageNumber="598">Heterocerus fausti</emphasis>
|
|||
|
|
Reitter, 1879
|
|||
|
|
</taxonomicName>
|
|||
|
|
, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFF17DE53FE99E192" box="[151,269,1899,1920]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFF17DE53FE99E192" box="[151,269,1899,1920]" italics="true" pageId="1" pageNumber="598">Heterocerus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
spp. 2A—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFEFBDE53FD13E192" authority="Gebler, 1830" authorityName="Gebler" authorityYear="1830" box="[379,647,1899,1920]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="parallelus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFEFBDE53FE68E192" box="[379,508,1899,1920]" italics="true" pageId="1" pageNumber="598">H. parallelus</emphasis>
|
|||
|
|
Gebler, 1830
|
|||
|
|
</taxonomicName>
|
|||
|
|
, South Kazakhstan; 2B—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFC08DE53FC49E192" authorityName="Reitter" authorityYear="1879" box="[904,989,1899,1920]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFC08DE53FC49E192" box="[904,989,1899,1920]" italics="true" pageId="1" pageNumber="598">H. fausti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Turkmenistan; 2C—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFB34DE53FF5DE18F" authority="Kiesenwetter, 1843" authorityName="Kiesenwetter" authorityYear="1843" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="fossor">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFB34DE53FA84E192" box="[1204,1296,1899,1920]" italics="true" pageId="1" pageNumber="598">H. fossor</emphasis>
|
|||
|
|
Kiesenwetter, 1843
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Russia, Dagestan; 2D—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFE42DEB0FD4FE18F" authority="Stephens, 1828" authorityName="Stephens" authorityYear="1828" box="[450,731,1928,1949]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFE42DEB0FDAFE18F" box="[450,571,1928,1949]" italics="true" pageId="1" pageNumber="598">H. flexuosus</emphasis>
|
|||
|
|
Stephens, 1828
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Kazakhstan; 2E—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFC1ADEB0FB24E18F" authority="Mascagni, 1989" authorityName="Mascagni" authorityYear="1989" box="[922,1200,1928,1949]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="persicus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFC1ADEB0FB9EE18F" box="[922,1034,1928,1949]" italics="true" pageId="1" pageNumber="598">H. persicus</emphasis>
|
|||
|
|
Mascagni, 1989
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Iran. Male genitalia of
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFF17DE9DFEDCE1A8" authorityName="Reitter" authorityYear="1879" box="[151,328,1957,1978]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFF17DE9DFEDCE1A8" box="[151,328,1957,1978]" italics="true" pageId="1" pageNumber="598">Heterocerus fausti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFCF9DE9DFC7BE1A8" box="[889,1007,1957,1978]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFCF9DE9DFC7BE1A8" box="[889,1007,1957,1978]" italics="true" pageId="1" pageNumber="598">Heterocerus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
spp. 4A—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFBDADE9DFB4DE1A8" authorityName="Gebler" authorityYear="1830" box="[1114,1241,1957,1978]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="parallelus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFBDADE9DFB4DE1A8" box="[1114,1241,1957,1978]" italics="true" pageId="1" pageNumber="598">H. parallelus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, South Kazakhstan; 4B—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFF4BDEFBFE1CE1CA" authority=", Caucasus" authorityName="Caucasus" box="[203,392,1987,2008]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFF4BDEFBFE89E1CA" box="[203,285,1987,2008]" italics="true" pageId="1" pageNumber="598">H. fausti</emphasis>
|
|||
|
|
, Caucasus
|
|||
|
|
</taxonomicName>
|
|||
|
|
; 4C—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFE46DEFBFD8AE1CA" authorityName="Kiesenwetter" authorityYear="1843" box="[454,542,1987,2008]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="fossor">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFE46DEFBFD8AE1CA" box="[454,542,1987,2008]" italics="true" pageId="1" pageNumber="598">H. fossor</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Russia, Dagestan; 4D—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFC93DEFBFC1EE1CA" authorityName="Stephens" authorityYear="1828" box="[787,906,1987,2008]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFC93DEFBFC1EE1CA" box="[787,906,1987,2008]" italics="true" pageId="1" pageNumber="598">H. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, European part of Russia (Litovkin
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFB67DEFBFA86E1CA" box="[1255,1298,1987,2008]" italics="true" pageId="1" pageNumber="598">et al</emphasis>
|
|||
|
|
., 2013), 4E—
|
|||
|
|
<taxonomicName id="4C774D1BFFCEFFCCFF17DED8FE97E1E7" authorityName="Mascagni" authorityYear="1989" box="[151,259,2016,2037]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="persicus">
|
|||
|
|
<emphasis id="B903EA8AFFCEFFCCFF17DED8FE97E1E7" box="[151,259,2016,2037]" italics="true" pageId="1" pageNumber="598">H. persicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Iran. Scale bar—0.5 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF086610FFCDFFCFFF17DE3EFB9CE10E" box="[151,1032,1798,1820]" httpUri="https://zenodo.org/record/1301564/files/figure.png" pageId="2" pageNumber="599" startId="2.[151,244,1798,1820]" targetBox="[151,1436,191,1777]" targetPageId="2">
|
|||
|
|
<paragraph id="8BC83698FFCDFFCFFF17DE3EFB9CE10E" blockId="2.[151,1032,1798,1820]" box="[151,1032,1798,1820]" pageId="2" pageNumber="599">
|
|||
|
|
<emphasis id="B903EA8AFFCDFFCFFF17DE3EFE98E10E" bold="true" box="[151,268,1798,1820]" pageId="2" pageNumber="599">FIGURE 5.</emphasis>
|
|||
|
|
Distribution of
|
|||
|
|
<taxonomicName id="4C774D1BFFCDFFCFFE25DE3FFD4FE10E" authority="Reitter, 1879" authorityName="Reitter" authorityYear="1879" box="[421,731,1799,1820]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCDFFCFFE25DE3FFDC2E10E" box="[421,598,1799,1820]" italics="true" pageId="2" pageNumber="599">Heterocerus fausti</emphasis>
|
|||
|
|
Reitter, 1879
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Red circles—studied material.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC83698FFCDFFCFFF47DE7FFC0CE193" blockId="2.[151,1436,1863,2025]" pageId="2" pageNumber="599">
|
|||
|
|
<taxonomicName id="4C774D1BFFCDFFCFFF47DE7FFE04E14C" authorityName="Reitter" authorityYear="1879" box="[199,400,1863,1886]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCDFFCFFF47DE7FFE04E14C" box="[199,400,1863,1886]" italics="true" pageId="2" pageNumber="599">Heterocerus fausti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is similar in body form and drawing of elytra on
|
|||
|
|
<taxonomicName id="4C774D1BFFCDFFCFFC55DE7FFB5BE14C" authorityName="Gebler" authorityYear="1830" box="[981,1231,1863,1886]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="parallelus">
|
|||
|
|
<emphasis id="B903EA8AFFCDFFCFFC55DE7FFB5BE14C" box="[981,1231,1863,1886]" italics="true" pageId="2" pageNumber="599">Heterocerus parallelus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
but can be easily distinguished by the shape of its aedeagus (
|
|||
|
|
<figureCitation id="134C2A1DFFCDFFCFFDE4DE52FD4CE193" box="[612,728,1898,1921]" captionStart="FIGURE 1–4" captionStartId="1.[151,244,1869,1891]" captionTargetBox="[151,1436,431,1852]" captionTargetId="figure@1.[151,1436,431,1852]" captionTargetPageId="1" captionText="FIGURE 1–4. Heterocerus fausti Reitter, 1879, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers Heterocerus spp. 2A—H. parallelus Gebler, 1830, South Kazakhstan; 2B—H. fausti, Turkmenistan; 2C—H. fossor Kiesenwetter, 1843, Russia, Dagestan; 2D—H. flexuosus Stephens, 1828, Kazakhstan; 2E—H. persicus Mascagni, 1989, Iran. Male genitalia of Heterocerus fausti, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens Heterocerus spp. 4A—H. parallelus, South Kazakhstan; 4B—H. fausti, Caucasus; 4C—H. fossor, Russia, Dagestan; 4D—H. flexuosus, European part of Russia (Litovkin et al., 2013), 4E— H. persicus, Iran. Scale bar—0.5 mm." httpUri="https://zenodo.org/record/1301562/files/figure.png" pageId="2" pageNumber="599">Fig. 4A–B</figureCitation>
|
|||
|
|
) and smaller size.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C36D6513FFCDFFCEFF47DEB4FC62E486" lastPageId="3" lastPageNumber="600" pageId="2" pageNumber="599" type="description">
|
|||
|
|
<paragraph id="8BC83698FFCDFFCEFF47DEB4FA9DE72B" blockId="2.[151,1436,1863,2025]" lastBlockId="3.[151,1436,152,833]" lastPageId="3" lastPageNumber="600" pageId="2" pageNumber="599">
|
|||
|
|
<emphasis id="B903EA8AFFCDFFCFFF47DEB4FEF8E1B1" bold="true" box="[199,364,1932,1955]" pageId="2" pageNumber="599">Redescription.</emphasis>
|
|||
|
|
<emphasis id="B903EA8AFFCDFFCFFEF9DEB4FE28E1B1" box="[377,444,1932,1955]" italics="true" pageId="2" pageNumber="599">Color.</emphasis>
|
|||
|
|
Ground color is yellow or red-yellow; a combination of black-brown blotches on the pronotum, and brown blotches of elytra and abdomen form a characteristic pattern. Head and labrum are black-brown. Antennae completely yellow. Pronotum is black-brown with a more or less wide yellow edging on the sides and narrow yellow stripe along the front edge. Scutellum is black-brown. Drawing of elytra characteristic of
|
|||
|
|
<taxonomicName id="4C774D1BFFCCFFCEFB1ED9A0FA95E6BD" authorityName="Stephens" authorityYear="1828" box="[1182,1281,152,175]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="600" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFB1ED9A0FA95E6BD" box="[1182,1281,152,175]" italics="true" pageId="3" pageNumber="600">flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species-group sensu
|
|||
|
|
<bibRefCitation id="EFE64B69FFCCFFCEFF58D982FE32E6C3" box="[216,422,186,209]" pageId="3" pageNumber="600" refId="ref1338" refString="Charpentier, R. (1965) A monograph of the family Heteroceridae of the Ethiopian Region. In: South African Animal Life. II. Result of the Lund University expedition in 1950 - 1951. Vol. XI. Swedish Natural Science Research Council, Stockholm, VA, pp. 215 - 343." type="book chapter">Charpentier (1965)</bibRefCitation>
|
|||
|
|
. Yellow pattern of each elytron normally forms edging, S-shaped basal mark, M-shaped medial mark, apical spot and suture stripe. Dark pattern of the elytra brownish almost blending with the yellow background. Prosternum black-brown with yellow episternae and yellowish front margin. Mesepisternum and metepisternum blackbrown. Abdomen black-brown with yellow pattern in form of edging. Legs, except dark metacoxa, yellow.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF47D87DFE58E76D" blockId="3.[151,1436,152,833]" pageId="3" pageNumber="600">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFF47D87DFE9CE74E" box="[199,264,325,348]" italics="true" pageId="3" pageNumber="600">Head.</emphasis>
|
|||
|
|
Clypeus transversal, widely incised on the front edge. Antennae 11-segmented with 7-segment club. Hairs whitish. Puncturing uniform.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF47D8B2FCD8E7F5" blockId="3.[151,1436,152,833]" pageId="3" pageNumber="600">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFF47D8B2FE8FE7B3" box="[199,283,394,417]" italics="true" pageId="3" pageNumber="600">Thorax.</emphasis>
|
|||
|
|
Pronotum not bordered basally; slightly narrower or comparable in width with base of elytra in females, and slightly wider than elytra base in males. Hairs on the disc are double: comprise dense, short, pressed, and long and standing hairs; whitish. Puncturing very dense, fine and uniform.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF17D8CAFC90E43E" blockId="3.[151,1436,152,833]" pageId="3" pageNumber="600">Elytra covered with erect hairs; hair length is comparable with the length of few first spines of mesotibia; hairs slightly longer along sides and near the elytral apex; hairs whitish.</paragraph>
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF17DB00FEC9E463" blockId="3.[151,1436,152,833]" pageId="3" pageNumber="600">
|
|||
|
|
Legs of structure typical for family, covered with whitish hairs. Claws of tarsus noticeably longer or equal to the last tarsomere (
|
|||
|
|
<figureCitation id="134C2A1DFFCCFFCEFE8EDB62FEC4E463" box="[270,336,602,625]" captionStart="FIGURE 1–4" captionStartId="1.[151,244,1869,1891]" captionTargetBox="[151,1436,431,1852]" captionTargetId="figure@1.[151,1436,431,1852]" captionTargetPageId="1" captionText="FIGURE 1–4. Heterocerus fausti Reitter, 1879, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers Heterocerus spp. 2A—H. parallelus Gebler, 1830, South Kazakhstan; 2B—H. fausti, Turkmenistan; 2C—H. fossor Kiesenwetter, 1843, Russia, Dagestan; 2D—H. flexuosus Stephens, 1828, Kazakhstan; 2E—H. persicus Mascagni, 1989, Iran. Male genitalia of Heterocerus fausti, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens Heterocerus spp. 4A—H. parallelus, South Kazakhstan; 4B—H. fausti, Caucasus; 4C—H. fossor, Russia, Dagestan; 4D—H. flexuosus, European part of Russia (Litovkin et al., 2013), 4E— H. persicus, Iran. Scale bar—0.5 mm." httpUri="https://zenodo.org/record/1301562/files/figure.png" pageId="3" pageNumber="600">Fig. 2</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF47DB45FC62E486" blockId="3.[151,1436,152,833]" box="[199,1014,637,660]" pageId="3" pageNumber="600">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFF47DB45FEB8E486" box="[199,300,637,660]" italics="true" pageId="3" pageNumber="600">Abdomen</emphasis>
|
|||
|
|
. Abdomen covered with whitish hairs. Postmetacoxal lines absent.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C36D6513FFCCFFCEFF47DB98FF47E4EE" pageId="3" pageNumber="600" type="discussion">
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF47DB98FF47E4EE" blockId="3.[151,1436,152,833]" pageId="3" pageNumber="600">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFF47DB98FEA1E4A5" bold="true" box="[199,309,672,695]" pageId="3" pageNumber="600">Remarks.</emphasis>
|
|||
|
|
The records of
|
|||
|
|
<taxonomicName id="4C774D1BFFCCFFCEFE6CDB98FD27E4A5" authorityName="Reitter" authorityYear="1879" box="[492,691,672,695]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="600" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFE6CDB98FD27E4A5" box="[492,691,672,695]" italics="true" pageId="3" pageNumber="600">Heterocerus fausti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
for
|
|||
|
|
<collectingCountry id="F3607608FFCCFFCEFD67DB98FCCBE4A5" box="[743,863,672,695]" name="Uzbekistan" pageId="3" pageNumber="600">Uzbekistan</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFE64B69FFCCFFCEFCF0DB98FB88E4A5" box="[880,1052,672,695]" pageId="3" pageNumber="600" refId="ref1447" refString="Jacobson, G. G. (1913) Semeystvo Heteroceridae. Pilousy. In: Zhuki Rossii i zapadnoy Evropy. Rukovodstvo k opredeleniyu zhukov. Vol. 10. A. F. Davrien, Sankt-Peterburg, pp. 867 - 869. [in Russian, English translation in: Family Heteroceridae. Mud-loving beetles. Beetles of Russia, Western Europe and neighbouring countries. Guide to the determination of beetles. A. F. Devrien, St. - Petersburg, 10, pp. 867 - 869]" type="book chapter">Jacobson, 1913</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFE64B69FFCCFFCEFBABDB98FB2BE4A5" box="[1067,1215,672,695]" pageId="3" pageNumber="600" refId="ref1644" refString="Zaitsev, F. A. (1946) Rasprostranenie v Zakavkaz'e vidov sem. Pilousov (Coleoptera, Heteroceridae). Trudy Zoologicheskogo instituta AN GruzSSR, 6, 213 - 220. [in Russian, English translation in: Distribution in Transcaucasia species of the variegated mud-loving beetles (Coleoptera, Heteroceridae). Works of Zoological Institute of the Georgian Academy of Sciences, 6, 213 - 220]" type="book chapter">Zaitsev, 1946</bibRefCitation>
|
|||
|
|
) are not confirmed. Specimen from
|
|||
|
|
<collectingCountry id="F3607608FFCCFFCEFEC6DBFAFE2AE4CB" box="[326,446,706,729]" name="Uzbekistan" pageId="3" pageNumber="600">Uzbekistan</collectingCountry>
|
|||
|
|
(Qamashi) signed of F.A. Zaitsev as “
|
|||
|
|
<taxonomicName id="4C774D1BFFCCFFCEFCE7DBFAFBB9E4CB" authorityName="Reitter" authorityYear="1879" box="[871,1069,706,729]" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="600" phylum="Arthropoda" rank="species" species="fausti">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFCE7DBFAFBB9E4CB" box="[871,1069,706,729]" italics="true" pageId="3" pageNumber="600">Heterocerus fausti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
” is
|
|||
|
|
<taxonomicName id="4C774D1BFFCCFFCEFBDDDBFAFF44E4EE" authority="Kuwert, 1890" authorityName="Kuwert" authorityYear="1890" class="Insecta" family="Heteroceridae" genus="Heterocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="600" phylum="Arthropoda" rank="species" species="heydeni">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFBDDDBFAFAA8E4CB" box="[1117,1340,706,729]" italics="true" pageId="3" pageNumber="600">Heterocerus heydeni</emphasis>
|
|||
|
|
Kuwert, 1890
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C36D6513FFCCFFCEFF47DA30FBF1E50D" box="[199,1125,776,799]" pageId="3" pageNumber="600" type="biology_ecology">
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF47DA30FBF1E50D" blockId="3.[151,1436,152,833]" box="[199,1125,776,799]" pageId="3" pageNumber="600">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFF47DA30FEB2E50D" bold="true" box="[199,294,776,799]" pageId="3" pageNumber="600">Ecology.</emphasis>
|
|||
|
|
The long claws are probably an adaptation to the psammophilous way of life.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C36D6513FFCCFFCEFF47DA12FB5CE553" box="[199,1224,810,833]" pageId="3" pageNumber="600" type="distribution">
|
|||
|
|
<paragraph id="8BC83698FFCCFFCEFF47DA12FB5CE553" blockId="3.[151,1436,152,833]" box="[199,1224,810,833]" pageId="3" pageNumber="600">
|
|||
|
|
<emphasis id="B903EA8AFFCCFFCEFF47DA12FECDE553" bold="true" box="[199,345,810,833]" pageId="3" pageNumber="600">Distribution.</emphasis>
|
|||
|
|
Pre-Caspian Region including Caucasus:
|
|||
|
|
<collectingCountry id="F3607608FFCCFFCEFC9ADA12FC06E553" box="[794,914,810,833]" name="Azerbaijan" pageId="3" pageNumber="600">Azerbaijan</collectingCountry>
|
|||
|
|
, West
|
|||
|
|
<collectingCountry id="F3607608FFCCFFCEFC58DA12FBFFE553" box="[984,1131,810,833]" name="Turkmenistan" pageId="3" pageNumber="600">Turkmenistan</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="134C2A1DFFCCFFCEFBF9DA12FB2EE553" box="[1145,1210,810,833]" captionStart="FIGURE 5" captionStartId="2.[151,244,1798,1820]" captionTargetBox="[151,1436,191,1777]" captionTargetId="figure@2.[151,1436,191,1777]" captionTargetPageId="2" captionText="FIGURE 5. Distribution of Heterocerus fausti Reitter, 1879. Red circles—studied material." httpUri="https://zenodo.org/record/1301564/files/figure.png" pageId="3" pageNumber="600">Fig. 5</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|