treatments-xml/data/03/CE/87/03CE87E0CC30FFE1FF3DCF86FDAF8F15.xml

<document id="141EB3E07C5CE7F29F377012FE31FA5B" ID-DOI="10.5281/zenodo.188300" ID-GBIF-Dataset="94399717-4170-49a8-81f4-5d94b701a1db" ID-ISSN="1175-5326" ID-Zenodo-Dep="188300" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461087766959" checkinUser="plazi" docAuthor="Kraus, Fred &amp; Allison, Allen" docDate="2009" docId="03CE87E0CC30FFE1FF3DCF86FDAF8F15" docLanguage="en" docName="zt02128p038.pdf" docOrigin="Zootaxa 2128" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cophixalus melanops Kraus &amp; Allison, 2009, sp. nov." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FFF7FF98CC3EFFF3FFAACE1BFF858E33" masterDocTitle="New species of Cophixalus (Anura: Microhylidae) from Papua New Guinea" masterLastPageNumber="38" masterPageNumber="1" pageNumber="15" updateTime="1698588117876" updateUser="plazi">
<mods:mods id="8DC694D3BDDE9850615A78AB235CF8F1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CAF92907C640A5022A0A0AFA9A7D416B">
<mods:title id="B14BAAA7E793A6E390D5AB37AA41036B">New species of Cophixalus (Anura: Microhylidae) from Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="DC05C640F3D7F1E52349650C158682CA" type="personal">
<mods:role id="87E79A2C5F778B0C019204B67BBD239E">
<mods:roleTerm id="6EE57AC3D1D5C97D1945AC1A59D1BB2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8612533B1B210268A925F7DE9C668E9C">Kraus, Fred</mods:namePart>
</mods:name>
<mods:name id="EB20D55973E823B087E3A6A83591F081" type="personal">
<mods:role id="D2140C504025ED801F2CFB269158BE84">
<mods:roleTerm id="EF3351784CC48F542B90DF8A15A318C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC2D13B5A3CFC90D18FE278A136F8CD4">Allison, Allen</mods:namePart>
</mods:name>
<mods:typeOfResource id="73175CBCA0C9551742F32C1FB0589620">text</mods:typeOfResource>
<mods:relatedItem id="7CBB818DE3E74CC87694628258E4C682" type="host">
<mods:titleInfo id="DB41F5CA202D6C6FD44A7E373D162126">
<mods:title id="ED4ECBCD859598BE95A22413323332D3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="010A0598B5CFCB8E4B7E329615CC3A8E">
<mods:date id="B1B07D0FC74D9A20DFE42A07776045FC">2009</mods:date>
<mods:detail id="7598D40FD78ADB5A05ED4D20DF407D02" type="volume">
<mods:number id="E5E48F4E12D890A4C9A4A9498282F883">2128</mods:number>
</mods:detail>
<mods:extent id="8434391FE5261869A0E453A49523684A" unit="page">
<mods:start id="26172B853C70DD5BB56ECE68B4B07C69">1</mods:start>
<mods:end id="B9146B2C62E329FD1FD4B853170C7EB3">38</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C585CF33FBE7841771F99E35423AC90E">journal article</mods:classification>
<mods:identifier id="F19C8BE51C86F80D3F3104D01334CDBC" type="DOI">10.5281/zenodo.188300</mods:identifier>
<mods:identifier id="C95218C14CEF084F47498792C33736E7" type="GBIF-Dataset">94399717-4170-49a8-81f4-5d94b701a1db</mods:identifier>
<mods:identifier id="3B10547B7E0E8948890B36F06C133B06" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8C58BBCDA36F8E8AC7FC37594567A3EB" type="Zenodo-Dep">188300</mods:identifier>
</mods:mods>
<treatment id="03CE87E0CC30FFE1FF3DCF86FDAF8F15" ID-DOI="http://doi.org/10.5281/zenodo.6223203" ID-GBIF-Taxon="119619444" ID-Zenodo-Dep="6223203" LSID="urn:lsid:plazi:treatment:03CE87E0CC30FFE1FF3DCF86FDAF8F15" httpUri="http://treatment.plazi.org/id/03CE87E0CC30FFE1FF3DCF86FDAF8F15" lastPageId="18" lastPageNumber="19" pageId="14" pageNumber="15">
<subSubSection id="C37D657DCC30FFFDFF3DCF86FE9B8FE9" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BD836F6CC30FFFDFF3DCF86FD8D8F84" blockId="14.[151,520,413,474]" box="[151,520,413,439]" pageId="14" pageNumber="15">
<heading id="D090819ACC30FFFDFF3DCF86FD8D8F84" bold="true" box="[151,520,413,439]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B913EAE4CC30FFFDFF3DCF86FD8D8F84" bold="true" box="[151,520,413,439]" pageId="14" pageNumber="15">
<taxonomicName id="4C674D75CC30FFFDFF3DCF86FE198F84" ID-CoL="Y44H" box="[151,412,413,439]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="melanops" status="sp. nov.">
<emphasis id="B913EAE4CC30FFFDFF3DCF86FE198F84" bold="true" box="[151,412,413,439]" italics="true" pageId="14" pageNumber="15">Cophixalus melanops</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A220579FCC30FFFDFE01CF86FD8D8F84" box="[427,520,413,439]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD836F6CC30FFFDFF3DCFDBFE9B8FE9" blockId="14.[151,520,413,474]" box="[151,286,448,474]" pageId="14" pageNumber="15">
<figureCitation id="135C2A73CC30FFFDFF3DCFDBFF628FE9" box="[151,231,448,474]" captionStart="FIGURE 2" captionStartId="15.[151,259,1744,1768]" captionTargetBox="[151,1436,197,1717]" captionTargetId="figure@15.[151,1436,197,1719]" captionTargetPageId="15" captionText="FIGURE 2. Photos in life of (A) holotype of Cophixalus melanops (BPBM 20198), (B) paratype of Cophixalus melanops (BPBM 20197), (C) holotype of Cophixalus linnaeus (BPBM 31836), (D) paratype of Cophixalus phaeobalius (BPBM 26194), (E) paratype of Cophixalus tomaiodactylus (BPBM 26182), and (F) holotype of Cophixalus tomaiodactylus (BPBM 26185)." httpUri="https://zenodo.org/record/188302/files/figure.png" pageId="14" pageNumber="15">Figs. 2</figureCitation>
A, B
</paragraph>
</subSubSection>
<subSubSection id="C37D657DCC30FFFDFF3DCC16FDA18D03" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BD836F6CC30FFFDFF3DCC16FF5C8C40" blockId="14.[151,1437,525,1879]" pageId="14" pageNumber="15">
<emphasis id="B913EAE4CC30FFFDFF3DCC16FE978C14" bold="true" box="[151,274,525,551]" pageId="14" pageNumber="15">
<typeStatus id="54DC8854CC30FFFDFF3DCC16FE978C14" box="[151,274,525,551]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<collectionCode id="ED76AE33CC30FFFDFE89CC16FEFC8C14" LSID="urn:lsid:biocol.org:col:34697" box="[291,377,525,551]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="15">BPBM</collectionCode>
20198 (field tag FK 9855), adult male, collected by F. Kraus on W slope Mt. Rio, 
<geoCoordinate id="EE535031CC30FFFDFF3DCC28FEBB8C7E" box="[151,318,563,589]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-11.508152">11.5081521ºS</geoCoordinate>
, 
<geoCoordinate id="EE535031CC30FFFDFEE3CC28FD848C7E" box="[329,513,563,589]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="153.43086">153.4308608ºE</geoCoordinate>
, 
<quantity id="4C9F9B13CC30FFFDFDA7CC28FD108C7E" box="[525,661,563,589]" metricMagnitude="2" metricUnit="m" metricValue="7.6" metricValueMax="8.0" metricValueMin="7.2" pageId="14" pageNumber="15" unit="m" value="760.0" valueMax="800.0" valueMin="720.0">720–800 m</quantity>
, Sudest Island, Milne Bay Province, 
<collectingCountry id="F3707666CC30FFFDFBEDCC28FAAB8C7E" box="[1095,1326,563,589]" name="Papua New Guinea" pageId="14" pageNumber="15">Papua New Guinea</collectingCountry>
, 
<date id="FFD91036CC30FFFDFA93CC28FF508C40" pageId="14" pageNumber="15" value="2004-04-25">25 April 2004</date>
.
</paragraph>
<paragraph id="8BD836F6CC30FFFDFF6CCC65FDA18D03" blockId="14.[151,1437,525,1879]" pageId="14" pageNumber="15">
<emphasis id="B913EAE4CC30FFFDFF6CCC65FE208CAB" bold="true" box="[198,421,638,664]" pageId="14" pageNumber="15">
<typeStatus id="54DC8854CC30FFFDFF6CCC65FEC68CAB" box="[198,323,638,664]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
(n = 9)
</emphasis>
. 
<collectionCode id="ED76AE33CC30FFFDFE1CCC64FD828CAA" LSID="urn:lsid:biocol.org:col:34697" box="[438,519,639,665]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="15">BPBM</collectionCode>
20192, W slope Mt. Rio, 
<geoCoordinate id="EE535031CC30FFFDFCE3CC64FC6A8CAA" box="[841,1007,639,665]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-11.505435">11.5054347ºS</geoCoordinate>
, 
<geoCoordinate id="EE535031CC30FFFDFC57CC64FB228CAA" box="[1021,1191,639,665]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="153.42995">153.429945ºE</geoCoordinate>
, 
<quantity id="4C9F9B13CC30FFFDFB1FCC64FABA8CAA" box="[1205,1343,639,665]" metricMagnitude="2" metricUnit="m" metricValue="6.8" metricValueMax="7.3" metricValueMin="6.3" pageId="14" pageNumber="15" unit="m" value="680.0" valueMax="730.0" valueMin="630.0">630–730 m</quantity>
, Sudest Island, 
<date id="FFD91036CC30FFFDFF45CCBFFE1F8C8D" box="[239,410,676,702]" pageId="14" pageNumber="15" value="2004-04-19">19 April 2004</date>
; 
<collectionCode id="ED76AE33CC30FFFDFE0DCCBFFE7C8C8D" LSID="urn:lsid:biocol.org:col:34697" box="[423,505,676,702]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="15">BPBM</collectionCode>
20193, W slope Mt. Rio, 
<geoCoordinate id="EE535031CC30FFFDFC99CCBFFC3B8C8D" box="[819,958,676,702]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-11.49436">11.49436ºS</geoCoordinate>
, 
<geoCoordinate id="EE535031CC30FFFDFC60CCBFFBE38C8D" box="[970,1126,676,702]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="153.42542">153.42542ºE</geoCoordinate>
, 
<quantity id="4C9F9B13CC30FFFDFBD9CCBFFB478C8D" box="[1139,1218,676,702]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="15" unit="m" value="440.0">440 m</quantity>
, Sudest Island, 
<date id="FFD91036CC30FFFDFAD4CCBFFE9F8CD7" pageId="14" pageNumber="15" value="2004-04-21">21 April 2004</date>
; 
<collectionCode id="ED76AE33CC30FFFDFE8DCCD1FEFD8CD7" LSID="urn:lsid:biocol.org:col:34697" box="[295,376,714,740]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="15">BPBM</collectionCode>
20194, W slope Mt. Rio, 
<geoCoordinate id="EE535031CC30FFFDFD06CCD1FCB08CD7" box="[684,821,714,740]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-11.50402">11.50402ºS</geoCoordinate>
, 
<geoCoordinate id="EE535031CC30FFFDFCE8CCD1FC598CD7" box="[834,988,714,740]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="153.42995">153.42995ºE</geoCoordinate>
, 
<quantity id="4C9F9B13CC30FFFDFC43CCD1FBB38CD7" box="[1001,1078,714,740]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="m" value="600.0">600 m</quantity>
, Sudest Island, 
<date id="FFD91036CC30FFFDFB45CCD1FA128CD7" box="[1263,1431,714,740]" pageId="14" pageNumber="15" value="2004-04-23">23 April 2004</date>
; 
<collectionCode id="ED76AE33CC30FFFDFF3DCCEBFF6F8D39" LSID="urn:lsid:biocol.org:col:34697" box="[151,234,752,778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="15">BPBM</collectionCode>
20195–96, same data as 
<typeStatus id="54DC8854CC30FFFDFDB5CCEBFD0C8D39" box="[543,649,752,778]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
except collected 
<date id="FFD91036CC30FFFDFCCACCEBFB8A8D39" box="[864,1039,752,778]" pageId="14" pageNumber="15" value="2004-04-23">23 April 2004</date>
; 
<collectionCode id="ED76AE33CC30FFFDFBB6CCEBFBEA8D39" LSID="urn:lsid:biocol.org:col:34697" box="[1052,1135,752,778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="15">BPBM</collectionCode>
20197, 20199, 
<collectionCode id="ED76AE33CC30FFFDFA9ACCEBFA198D39" box="[1328,1436,752,778]" pageId="14" pageNumber="15">PNGNM</collectionCode>
24066–67, same data as 
<typeStatus id="54DC8854CC30FFFDFE1CCD0DFDA58D03" box="[438,544,790,816]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D657DCC30FFFDFF6CCD20FE8C8B2F" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BD836F6CC30FFFDFF6CCD20FC9D8DDE" blockId="14.[151,1437,525,1879]" pageId="14" pageNumber="15">
<emphasis id="B913EAE4CC30FFFDFF6CCD20FEC38D66" bold="true" box="[198,326,827,853]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
A small species (male SV = 
<quantity id="4C9F9B13CC30FFFDFD08CD27FCD48D65" box="[674,849,828,854]" metricMagnitude="-2" metricUnit="m" metricValue="1.765" metricValueMax="1.89" metricValueMin="1.64" pageId="14" pageNumber="15" unit="mm" value="17.65" valueMax="18.9" valueMin="16.4">16.4–18.9 mm</quantity>
, female SV = 
<quantity id="4C9F9B13CC30FFFDFC57CD27FBED8D65" box="[1021,1128,828,854]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="14" pageNumber="15" unit="mm" value="21.8">21.8 mm</quantity>
) with finger discs smaller than toe discs (3rdF/4thT = 0.63–0.78); first finger of normal size, with disc bearing circum-marginal groove; wide head (HW/SV = 0.41–0.44); large eye (EY/SV = 0.14–0.17); small tympanum (TY/SV = 0.043–0.069), side of face uniformly black; lateral surfaces same color as dorsum or darker than dorsum but not bearing black lateral band; and short, black supratympanic bar.
</paragraph>
<paragraph id="8BD836F6CC30FFFDFF6CCDE2FE8C8B2F" blockId="14.[151,1437,525,1879]" pageId="14" pageNumber="15">
<emphasis id="B913EAE4CC30FFFDFF6CCDE2FDDE8A20" bold="true" box="[198,603,1017,1043]" pageId="14" pageNumber="15">Comparisons with other species.</emphasis>
The new species differs from all Papuan congeners except 
<taxonomicName id="4C674D75CC30FFFDFAB2CDE2FE098A0A" authority="Richards and Oliver" authorityName="Richards and Oliver" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFAB2CDE2FA1E8A21" box="[1304,1435,1017,1042]" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
Richards and Oliver
</taxonomicName>
in the combination of having finger discs distinctly smaller than toe discs and a first finger of normal size, bearing a disc with a circum-marginal groove. 
<taxonomicName id="4C674D75CC30FFFDFC76CA5EFB5A8A6D" box="[988,1247,1093,1118]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="melanops">
<emphasis id="B913EAE4CC30FFFDFC76CA5EFB5A8A6D" box="[988,1247,1093,1118]" italics="true" pageId="14" pageNumber="15">Cophixalus melanops</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C674D75CC30FFFDFA2BCA5EFF7F8AB7" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFA2BCA5EFF7F8AB7" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
</taxonomicName>
in having a wider head (HW/SV = 
<quantity id="4C9F9B13CC30FFFDFD0CCA70FCB98AB6" box="[678,828,1131,1157]" metricMagnitude="-3" metricUnit="m" metricValue="8.508999999999999" metricValueMax="9.144" metricValueMin="7.874" pageId="14" pageNumber="15" unit="in" value="0.33499999999999996" valueMax="0.36" valueMin="0.31">0.31–0.36 in</quantity>
<taxonomicName id="4C674D75CC30FFFDFCEECA70FC4F8AB7" box="[836,970,1131,1156]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFCEECA70FC4F8AB7" box="[836,970,1131,1156]" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
</taxonomicName>
), larger eye (EY/SV = 
<quantity id="4C9F9B13CC30FFFDFB48CA70FAFD8AB6" box="[1250,1400,1131,1157]" metricMagnitude="-3" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="14" pageNumber="15" unit="in" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 in</quantity>
<taxonomicName id="4C674D75CC30FFFDFA2BCA70FF788A99" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFA2BCA70FF788A99" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
</taxonomicName>
), smaller tympanum (TY/SV = 
<quantity id="4C9F9B13CC30FFFDFD27CA8AFCCD8A98" box="[653,840,1169,1195]" metricMagnitude="-3" metricUnit="m" metricValue="1.778" metricValueMax="1.905" metricValueMin="1.651" pageId="14" pageNumber="15" unit="in" value="0.07" valueMax="0.075" valueMin="0.065">0.065–0.075 in</quantity>
<taxonomicName id="4C674D75CC30FFFDFCFACA8AFC5E8A99" box="[848,987,1169,1194]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFCFACA8AFC5E8A99" box="[848,987,1169,1194]" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
</taxonomicName>
), lacking the black lateral band and inguinal black spots of the former, having the snout truncate in lateral view (projecting in 
<taxonomicName id="4C674D75CC30FFFDFB1ACAACFAB18AE3" box="[1200,1332,1207,1232]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFB1ACAACFAB18AE3" box="[1200,1332,1207,1232]" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
</taxonomicName>
), having oblique lores (vertical in 
<taxonomicName id="4C674D75CC30FFFDFE17CAC6FDC48AC5" box="[445,577,1245,1270]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFE17CAC6FDC48AC5" box="[445,577,1245,1270]" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
</taxonomicName>
), and having a black bar on the upper part of the tympanum (absent in 
<taxonomicName id="4C674D75CC30FFFDFA2BCAC6FF7C8B2F" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="misimae">
<emphasis id="B913EAE4CC30FFFDFA2BCAC6FF7C8B2F" italics="true" pageId="14" pageNumber="15">C. misimae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D657DCC30FFE3FF6CCB33FE7988C1" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="8BD836F6CC30FFFDFF6CCB33FAE18964" blockId="14.[151,1437,525,1879]" pageId="14" pageNumber="15">
<emphasis id="B913EAE4CC30FFFDFF6CCB33FE748B71" bold="true" box="[198,497,1320,1346]" pageId="14" pageNumber="15">
Description of 
<typeStatus id="54DC8854CC30FFFDFE2FCB33FE748B71" box="[389,497,1320,1346]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
</emphasis>
. An adult male with right-lateral incision, vocal slits. Head moderately wide (HW/SV = 0.42), with steep but oblique loreal region; canthus rostralis rounded, straight when viewed from above; nostrils directed laterally, much closer to tip of snout than to eyes; internarial distance slightly larger than distance from naris to eye (EN/IN = 0.90, IN/SV = 0.111, EN/SV = 0.101); snout rounded and slightly projecting when viewed from the side, shallowly angulate when viewed from above; eyes large (EY/SV = 0.15); eyelid 2/3 width of interorbital distance; tympanum small (TY/SV = 0.058), with a clear, raised annulus. Dorsal and lateral surfaces smooth; supratympanic fold absent. Ventral surfaces generally smooth but weakly granular on abdomen. Fingers unwebbed, bearing discs with terminal grooves; relative lengths 3&gt;4&gt;2&gt;1; first finger and disc well-developed. Finger discs approximately 1.5 times widths of penultimate phalanges, except for the first finger disc, which is only slightly wider than penultimate phalanx. Subarticular tubercles not obvious; metacarpal tubercles poorly developed, inner narrow and elongate, outer round. Toes unwebbed, bearing discs with terminal grooves; relative lengths 4&gt;3&gt;5&gt;2&gt;1. Toe discs larger than those of fingers (3rd F/4thT = 0.71); disc of fourth toe approximately twice width of penultimate phalanx; disc of first toe only slightly wider than penultimate phalanx. Subarticular tubercles reduced to mere skin thickenings; inner metatarsal tubercle narrow and elongate, outer lacking. 
<collectingCountry id="F3707666CC30FFFDFCCCC926FC248964" box="[870,929,1853,1879]" name="India" pageId="14" pageNumber="15">Hind</collectingCountry>
legs moderately long (TL/SV = 0.52).
</paragraph>
<caption id="DF18667ECC31FFFCFF3DC8CBFE68897C" httpUri="https://zenodo.org/record/188302/files/figure.png" pageId="15" pageNumber="16" targetBox="[151,1436,197,1717]" targetPageId="15">
<paragraph id="8BD836F6CC31FFFCFF3DC8CBFE68897C" blockId="15.[151,1437,1744,1872]" pageId="15" pageNumber="16">
<emphasis id="B913EAE4CC31FFFCFF3DC8CBFEA488DB" bold="true" box="[151,289,1744,1768]" pageId="15" pageNumber="16">FIGURE 2.</emphasis>
Photos in life of (A) holotype of 
<taxonomicName id="4C674D75CC31FFFCFD0CC8CAFC1988DB" box="[678,924,1745,1768]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="melanops">
<emphasis id="B913EAE4CC31FFFCFD0CC8CAFC1988DB" box="[678,924,1745,1768]" italics="true" pageId="15" pageNumber="16">Cophixalus melanops</emphasis>
</taxonomicName>
(BPBM 20198), (B) paratype of 
<taxonomicName id="4C674D75CC31FFFCFAB1C8CAFF788939" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="melanops">
<emphasis id="B913EAE4CC31FFFCFAB1C8CAFF788939" italics="true" pageId="15" pageNumber="16">Cophixalus melanops</emphasis>
</taxonomicName>
(BPBM 20197), (C) holotype of 
<taxonomicName id="4C674D75CC31FFFCFDF4C8E8FCBC8939" box="[606,825,1779,1802]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="linnaeus">
<emphasis id="B913EAE4CC31FFFCFDF4C8E8FCBC8939" box="[606,825,1779,1802]" italics="true" pageId="15" pageNumber="16">Cophixalus linnaeus</emphasis>
</taxonomicName>
(BPBM 31836), (D) paratype of 
<taxonomicName id="4C674D75CC31FFFCFB30C8E8FA188939" box="[1178,1437,1779,1802]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="phaeobalius">
<emphasis id="B913EAE4CC31FFFCFB30C8E8FA188939" box="[1178,1437,1779,1802]" italics="true" pageId="15" pageNumber="16">Cophixalus phaeobalius</emphasis>
</taxonomicName>
(BPBM 26194), (E) paratype of 
<taxonomicName id="4C674D75CC31FFFCFDB3C90DFCDE891E" box="[537,859,1814,1837]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="tomaiodactylus">
<emphasis id="B913EAE4CC31FFFCFDB3C90DFCDE891E" box="[537,859,1814,1837]" italics="true" pageId="15" pageNumber="16">Cophixalus tomaiodactylus</emphasis>
</taxonomicName>
(BPBM 26182), and (F) holotype of 
<taxonomicName id="4C674D75CC31FFFCFAB2C90DFEBF8963" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="tomaiodactylus">
<emphasis id="B913EAE4CC31FFFCFAB2C90DFEBF8963" italics="true" pageId="15" pageNumber="16">Cophixalus tomaiodactylus</emphasis>
</taxonomicName>
(BPBM 26185).
</paragraph>
</caption>
<paragraph id="8BD836F6CC31FFE3FF6FC965FB3C8F15" blockId="15.[151,1437,1918,2021]" lastBlockId="16.[151,1437,152,647]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">Dorsum medium brown with few, scattered, tiny dark-brown spots and dark-brown postocular blotch; sides lighter. Face dark brown to angle of jaw, with few obscure ligher brown specks along mandible. Darkbrown stripe on lower half of anterior face of upper arm. Rear of thighs dorsally and proximally as for dorsum, ventrally and distally dark brown. Venter pale straw, heavily flecked with dark brown on chin and throat, sparsely flecked on chest and abdomen, producing high-contrasting appearance of light belly with dark chin and throat. Undersides of thighs light straw anteriorly, heavily flecked with dark brown posteriorly. Front of thighs pale straw with dark-brown flecks. Iris black, heavily flecked with silver-bronze.</paragraph>
<paragraph id="8BD836F6CC2EFFE3FF6CCF29FDB18F4A" blockId="16.[151,1437,152,647]" pageId="16" pageNumber="17">
<emphasis id="B913EAE4CC2EFFE3FF6CCF29FE5F8F78" box="[198,474,306,331]" italics="true" pageId="16" pageNumber="17">Measurements (in mm).</emphasis>
SV = 18.9, TL= 9.8, HW = 8.0, HL = 7.1, IN = 2.1, EN = 1.9, SN = 2.9, EY = 2.8, TY = 1.1, 3rd F = 0.64, 4th T = 0.90.
</paragraph>
<paragraph id="8BD836F6CC2EFFE3FF6CCF9EFE178CB4" blockId="16.[151,1437,152,647]" pageId="16" pageNumber="17">
<emphasis id="B913EAE4CC2EFFE3FF6CCF9EFEC48FAC" bold="true" box="[198,321,389,415]" pageId="16" pageNumber="17">Variation.</emphasis>
Mensural variation for the 
<typeStatus id="54DC8854CC2EFFE3FD27CF9EFD458FAC" box="[653,704,389,415]" pageId="16" pageNumber="17">type</typeStatus>
series is shown in Table 8. The two adult 
<typeStatus id="54DC8854CC2EFFE3FB14CF9EFAAB8FAC" box="[1214,1326,389,415]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
have the snout more truncate than the 
<typeStatus id="54DC8854CC2EFFE3FE58CFB7FDDF8FF5" box="[498,602,428,454]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
when seen in lateral view. The adult male 
<typeStatus id="54DC8854CC2EFFE3FBF0CFB7FB458FF5" box="[1114,1216,428,454]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
(BPBM 20192) is identical to the 
<typeStatus id="54DC8854CC2EFFE3FECFCFC8FE538FDE" box="[357,470,467,493]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
in appearance. The adult female 
<typeStatus id="54DC8854CC2EFFE3FC21CFC8FC7F8FDE" box="[907,1018,467,493]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
(BPBM 20197) has dark-brown dorsolateral stripes extending on each side from scapular region to groin; sides below this stripe darker than dorsum; chest more heavily flecked with dark brown than in 
<typeStatus id="54DC8854CC2EFFE3FC39CC3BFB818C09" box="[915,1028,544,570]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
. The remaining 
<typeStatus id="54DC8854CC2EFFE3FB65CC3BFAC38C09" box="[1231,1350,544,570]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
are all juveniles and are similar in appearance to the 
<typeStatus id="54DC8854CC2EFFE3FD12CC5CFC9A8C52" box="[696,799,583,609]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
except that there are variable amounts of dark-brown flecking on the chest.
</paragraph>
<paragraph id="8BD836F6CC2EFFE3FF3DCCA0FAD28B04" pageId="16" pageNumber="17">
<table id="F967C456CC2E000CFF3DCCA0FAE08B04" box="[151,1381,699,1335]" gridcols="5" gridrows="16" pageId="16" pageNumber="17">
<tr id="355734B4CC2E000CFF3DCCA0FAE08CE0" box="[151,1381,699,723]" gridrow="0" pageId="16" pageNumber="17" rowspan-3="1" rowspan-4="1">
<th id="76865DC8CC2E000CFF3DCCA0FEF08CE0" box="[151,373,699,723]" gridcol="0" gridrow="0" pageId="16" pageNumber="17">
<emphasis id="B913EAE4CC2EFFE3FF3DCCA0FE8D8CE0" bold="true" box="[151,264,699,723]" pageId="16" pageNumber="17">TABLE 8.</emphasis>
Mensural
</th>
<th id="76865DC8CC2E000CFED6CCA0FDC08CE0" box="[380,581,699,723]" gridcol="1" gridrow="0" pageId="16" pageNumber="17">data for type series</th>
<th id="76865DC8CC2E000CFDE6CCA0FC548CE0" box="[588,977,699,723]" gridcol="2" gridrow="0" pageId="16" pageNumber="17">
of 
<taxonomicName id="4C674D75CC2EFFE3FDC3CCA0FCD58CE1" box="[617,848,699,722]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="melanops" status="sp. nov.">
<emphasis id="B913EAE4CC2EFFE3FDC3CCA0FCD58CE1" box="[617,848,699,722]" italics="true" pageId="16" pageNumber="17">Cophixalus melanops</emphasis>
</taxonomicName>
<emphasis id="B913EAE4CC2EFFE3FCFCCCA0FC298CE0" bold="true" box="[854,940,699,723]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A220579FCC2EFFE3FCFCCCA0FC298CE0" box="[854,940,699,723]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="355734B4CC2E000CFF3DCCF6FAE08D19" box="[151,1381,749,810]" gridrow="1" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCCF6FEF08D19" box="[151,373,749,810]" gridcol="0" gridrow="1" pageId="16" pageNumber="17">Character</th>
<td id="76865DC8CC2E000CFED6CCF6FDC08D19" box="[380,581,749,810]" gridcol="1" gridrow="1" pageId="16" pageNumber="17">BPBM 20192 male</td>
<td id="76865DC8CC2E000CFDE6CCF6FC548D19" box="[588,977,749,810]" gridcol="2" gridrow="1" pageId="16" pageNumber="17">BPBM 20198 BPBM 20197 male female</td>
<td id="76865DC8CC2E000CFB98CCF6FB518D19" box="[1074,1236,749,810]" gridcol="3" gridrow="1" pageId="16" pageNumber="17">Juveniles mean</td>
<td id="76865DC8CC2E000CFB71CCF6FAE08D19" box="[1243,1381,749,810]" gridcol="4" gridrow="1" pageId="16" pageNumber="17">(n = 5) range</td>
</tr>
<tr id="355734B4CC2E000CFF3DCD23FAE08D7C" box="[151,1381,824,847]" gridrow="2" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCD23FEF08D7C" box="[151,373,824,847]" gridcol="0" gridrow="2" pageId="16" pageNumber="17">SV (mm)</th>
<td id="76865DC8CC2E000CFED6CD23FDC08D7C" box="[380,581,824,847]" gridcol="1" gridrow="2" pageId="16" pageNumber="17">16.4</td>
<td id="76865DC8CC2E000CFDE6CD23FC548D7C" box="[588,977,824,847]" gridcol="2" gridrow="2" pageId="16" pageNumber="17">18.9 21.8</td>
<td id="76865DC8CC2E000CFB98CD23FB518D7C" box="[1074,1236,824,847]" gridcol="3" gridrow="2" pageId="16" pageNumber="17">11.2</td>
<td id="76865DC8CC2E000CFB71CD23FAE08D7C" box="[1243,1381,824,847]" gridcol="4" gridrow="2" pageId="16" pageNumber="17">8.7–13.3</td>
</tr>
<tr id="355734B4CC2E000CFF3DCD46FAE08D47" box="[151,1381,861,884]" gridrow="3" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCD46FEF08D47" box="[151,373,861,884]" gridcol="0" gridrow="3" pageId="16" pageNumber="17">TL/SV</th>
<td id="76865DC8CC2E000CFED6CD46FDC08D47" box="[380,581,861,884]" gridcol="1" gridrow="3" pageId="16" pageNumber="17">0.56</td>
<td id="76865DC8CC2E000CFDE6CD46FC548D47" box="[588,977,861,884]" gridcol="2" gridrow="3" pageId="16" pageNumber="17">0.52 0.52</td>
<td id="76865DC8CC2E000CFB98CD46FB518D47" box="[1074,1236,861,884]" gridcol="3" gridrow="3" pageId="16" pageNumber="17">0.53</td>
<td id="76865DC8CC2E000CFB71CD46FAE08D47" box="[1243,1381,861,884]" gridcol="4" gridrow="3" pageId="16" pageNumber="17">0.48–0.58</td>
</tr>
<tr id="355734B4CC2E000CFF3DCD98FAE08DA9" box="[151,1381,899,922]" gridrow="4" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCD98FEF08DA9" box="[151,373,899,922]" gridcol="0" gridrow="4" pageId="16" pageNumber="17">EN/SV</th>
<td id="76865DC8CC2E000CFED6CD98FDC08DA9" box="[380,581,899,922]" gridcol="1" gridrow="4" pageId="16" pageNumber="17">0.098</td>
<td id="76865DC8CC2E000CFDE6CD98FC548DA9" box="[588,977,899,922]" gridcol="2" gridrow="4" pageId="16" pageNumber="17">0.101 0.096</td>
<td id="76865DC8CC2E000CFB98CD98FB518DA9" box="[1074,1236,899,922]" gridcol="3" gridrow="4" pageId="16" pageNumber="17">0.097</td>
<td id="76865DC8CC2E000CFB71CD98FAE08DA9" box="[1243,1381,899,922]" gridcol="4" gridrow="4" pageId="16" pageNumber="17">0.081–0.105</td>
</tr>
<tr id="355734B4CC2E000CFF3DCDB3FAE08D8C" box="[151,1381,936,959]" gridrow="5" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCDB3FEF08D8C" box="[151,373,936,959]" gridcol="0" gridrow="5" pageId="16" pageNumber="17">IN/SV</th>
<td id="76865DC8CC2E000CFED6CDB3FDC08D8C" box="[380,581,936,959]" gridcol="1" gridrow="5" pageId="16" pageNumber="17">0.110</td>
<td id="76865DC8CC2E000CFDE6CDB3FC548D8C" box="[588,977,936,959]" gridcol="2" gridrow="5" pageId="16" pageNumber="17">0.111 0.119</td>
<td id="76865DC8CC2E000CFB98CDB3FB518D8C" box="[1074,1236,936,959]" gridcol="3" gridrow="5" pageId="16" pageNumber="17">0.123</td>
<td id="76865DC8CC2E000CFB71CDB3FAE08D8C" box="[1243,1381,936,959]" gridcol="4" gridrow="5" pageId="16" pageNumber="17">0.117–0.128</td>
</tr>
<tr id="355734B4CC2E000CFF3DCDD6FAE08DD7" box="[151,1381,973,996]" gridrow="6" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCDD6FEF08DD7" box="[151,373,973,996]" gridcol="0" gridrow="6" pageId="16" pageNumber="17">SN/SV</th>
<td id="76865DC8CC2E000CFED6CDD6FDC08DD7" box="[380,581,973,996]" gridcol="1" gridrow="6" pageId="16" pageNumber="17">0.15</td>
<td id="76865DC8CC2E000CFDE6CDD6FC548DD7" box="[588,977,973,996]" gridcol="2" gridrow="6" pageId="16" pageNumber="17">0.15 0.15</td>
<td id="76865DC8CC2E000CFB98CDD6FB518DD7" box="[1074,1236,973,996]" gridcol="3" gridrow="6" pageId="16" pageNumber="17">0.15</td>
<td id="76865DC8CC2E000CFB71CDD6FAE08DD7" box="[1243,1381,973,996]" gridcol="4" gridrow="6" pageId="16" pageNumber="17">0.14–0.17</td>
</tr>
<tr id="355734B4CC2E000CFF3DCDE8FAE08A39" box="[151,1381,1011,1034]" gridrow="7" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCDE8FEF08A39" box="[151,373,1011,1034]" gridcol="0" gridrow="7" pageId="16" pageNumber="17">TY/SV</th>
<td id="76865DC8CC2E000CFED6CDE8FDC08A39" box="[380,581,1011,1034]" gridcol="1" gridrow="7" pageId="16" pageNumber="17">0.043</td>
<td id="76865DC8CC2E000CFDE6CDE8FC548A39" box="[588,977,1011,1034]" gridcol="2" gridrow="7" pageId="16" pageNumber="17">0.058 0.069</td>
<td id="76865DC8CC2E000CFB98CDE8FB518A39" box="[1074,1236,1011,1034]" gridcol="3" gridrow="7" pageId="16" pageNumber="17">0.056</td>
<td id="76865DC8CC2E000CFB71CDE8FAE08A39" box="[1243,1381,1011,1034]" gridcol="4" gridrow="7" pageId="16" pageNumber="17">0.052–0.062</td>
</tr>
<tr id="355734B4CC2E000CFF3DCA03FAE08A1C" box="[151,1381,1048,1071]" gridrow="8" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCA03FEF08A1C" box="[151,373,1048,1071]" gridcol="0" gridrow="8" pageId="16" pageNumber="17">EY/SV</th>
<td id="76865DC8CC2E000CFED6CA03FDC08A1C" box="[380,581,1048,1071]" gridcol="1" gridrow="8" pageId="16" pageNumber="17">0.14</td>
<td id="76865DC8CC2E000CFDE6CA03FC548A1C" box="[588,977,1048,1071]" gridcol="2" gridrow="8" pageId="16" pageNumber="17">0.15 0.15</td>
<td id="76865DC8CC2E000CFB98CA03FB518A1C" box="[1074,1236,1048,1071]" gridcol="3" gridrow="8" pageId="16" pageNumber="17">0.16</td>
<td id="76865DC8CC2E000CFB71CA03FAE08A1C" box="[1243,1381,1048,1071]" gridcol="4" gridrow="8" pageId="16" pageNumber="17">0.14–0.17</td>
</tr>
<tr id="355734B4CC2E000CFF3DCA26FAE08A67" box="[151,1381,1085,1108]" gridrow="9" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCA26FEF08A67" box="[151,373,1085,1108]" gridcol="0" gridrow="9" pageId="16" pageNumber="17">HW/SV</th>
<td id="76865DC8CC2E000CFED6CA26FDC08A67" box="[380,581,1085,1108]" gridcol="1" gridrow="9" pageId="16" pageNumber="17">0.43</td>
<td id="76865DC8CC2E000CFDE6CA26FC548A67" box="[588,977,1085,1108]" gridcol="2" gridrow="9" pageId="16" pageNumber="17">0.42 0.41</td>
<td id="76865DC8CC2E000CFB98CA26FB518A67" box="[1074,1236,1085,1108]" gridcol="3" gridrow="9" pageId="16" pageNumber="17">0.42</td>
<td id="76865DC8CC2E000CFB71CA26FAE08A67" box="[1243,1381,1085,1108]" gridcol="4" gridrow="9" pageId="16" pageNumber="17">0.41–0.44</td>
</tr>
<tr id="355734B4CC2E000CFF3DCA78FAE08A49" box="[151,1381,1123,1146]" gridrow="10" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCA78FEF08A49" box="[151,373,1123,1146]" gridcol="0" gridrow="10" pageId="16" pageNumber="17">HL/SV</th>
<td id="76865DC8CC2E000CFED6CA78FDC08A49" box="[380,581,1123,1146]" gridcol="1" gridrow="10" pageId="16" pageNumber="17">0.40</td>
<td id="76865DC8CC2E000CFDE6CA78FC548A49" box="[588,977,1123,1146]" gridcol="2" gridrow="10" pageId="16" pageNumber="17">0.38 0.38</td>
<td id="76865DC8CC2E000CFB98CA78FB518A49" box="[1074,1236,1123,1146]" gridcol="3" gridrow="10" pageId="16" pageNumber="17">0.40</td>
<td id="76865DC8CC2E000CFB71CA78FAE08A49" box="[1243,1381,1123,1146]" gridcol="4" gridrow="10" pageId="16" pageNumber="17">0.39–0.41</td>
</tr>
<tr id="355734B4CC2E000CFF3DCA93FAE08AAC" box="[151,1381,1160,1183]" gridrow="11" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCA93FEF08AAC" box="[151,373,1160,1183]" gridcol="0" gridrow="11" pageId="16" pageNumber="17">3rdF/SV</th>
<td id="76865DC8CC2E000CFED6CA93FDC08AAC" box="[380,581,1160,1183]" gridcol="1" gridrow="11" pageId="16" pageNumber="17">0.035</td>
<td id="76865DC8CC2E000CFDE6CA93FC548AAC" box="[588,977,1160,1183]" gridcol="2" gridrow="11" pageId="16" pageNumber="17">0.034 0.034</td>
<td id="76865DC8CC2E000CFB98CA93FB518AAC" box="[1074,1236,1160,1183]" gridcol="3" gridrow="11" pageId="16" pageNumber="17">0.030</td>
<td id="76865DC8CC2E000CFB71CA93FAE08AAC" box="[1243,1381,1160,1183]" gridcol="4" gridrow="11" pageId="16" pageNumber="17">0.025–0.034</td>
</tr>
<tr id="355734B4CC2E000CFF3DCAB6FAE08AF7" box="[151,1381,1197,1220]" gridrow="12" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCAB6FEF08AF7" box="[151,373,1197,1220]" gridcol="0" gridrow="12" pageId="16" pageNumber="17">4thT/SV</th>
<td id="76865DC8CC2E000CFED6CAB6FDC08AF7" box="[380,581,1197,1220]" gridcol="1" gridrow="12" pageId="16" pageNumber="17">0.048</td>
<td id="76865DC8CC2E000CFDE6CAB6FC548AF7" box="[588,977,1197,1220]" gridcol="2" gridrow="12" pageId="16" pageNumber="17">0.048 0.050</td>
<td id="76865DC8CC2E000CFB98CAB6FB518AF7" box="[1074,1236,1197,1220]" gridcol="3" gridrow="12" pageId="16" pageNumber="17">0.044</td>
<td id="76865DC8CC2E000CFB71CAB6FAE08AF7" box="[1243,1381,1197,1220]" gridcol="4" gridrow="12" pageId="16" pageNumber="17">0.037–0.051</td>
</tr>
<tr id="355734B4CC2E000CFF3DCAC8FAE08AD9" box="[151,1381,1235,1258]" gridrow="13" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCAC8FEF08AD9" box="[151,373,1235,1258]" gridcol="0" gridrow="13" pageId="16" pageNumber="17">EN/IN</th>
<td id="76865DC8CC2E000CFED6CAC8FDC08AD9" box="[380,581,1235,1258]" gridcol="1" gridrow="13" pageId="16" pageNumber="17">0.89</td>
<td id="76865DC8CC2E000CFDE6CAC8FC548AD9" box="[588,977,1235,1258]" gridcol="2" gridrow="13" pageId="16" pageNumber="17">0.90 0.81</td>
<td id="76865DC8CC2E000CFB98CAC8FB518AD9" box="[1074,1236,1235,1258]" gridcol="3" gridrow="13" pageId="16" pageNumber="17">0.78</td>
<td id="76865DC8CC2E000CFB71CAC8FAE08AD9" box="[1243,1381,1235,1258]" gridcol="4" gridrow="13" pageId="16" pageNumber="17">0.69–0.82</td>
</tr>
<tr id="355734B4CC2E000CFF3DCAE3FAE08B21" box="[151,1381,1272,1298]" gridrow="14" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCAE3FEF08B21" box="[151,373,1272,1298]" gridcol="0" gridrow="14" pageId="16" pageNumber="17">3rd F/4th T</th>
<td id="76865DC8CC2E000CFED6CAE3FDC08B21" box="[380,581,1272,1298]" gridcol="1" gridrow="14" pageId="16" pageNumber="17">0.74</td>
<td id="76865DC8CC2E000CFDE6CAE3FC548B21" box="[588,977,1272,1298]" gridcol="2" gridrow="14" pageId="16" pageNumber="17">0.71 0.67</td>
<td id="76865DC8CC2E000CFB98CAE3FB518B21" box="[1074,1236,1272,1298]" gridcol="3" gridrow="14" pageId="16" pageNumber="17">0.68</td>
<td id="76865DC8CC2E000CFB71CAE3FAE08B21" box="[1243,1381,1272,1298]" gridcol="4" gridrow="14" pageId="16" pageNumber="17">0.63–0.72</td>
</tr>
<tr id="355734B4CC2E000CFF3DCB3BFAE08B04" box="[151,1381,1312,1335]" gridrow="15" pageId="16" pageNumber="17">
<th id="76865DC8CC2E000CFF3DCB3BFEF08B04" box="[151,373,1312,1335]" gridcol="0" gridrow="15" pageId="16" pageNumber="17">HL/HW</th>
<td id="76865DC8CC2E000CFED6CB3BFDC08B04" box="[380,581,1312,1335]" gridcol="1" gridrow="15" pageId="16" pageNumber="17">0.92</td>
<td id="76865DC8CC2E000CFDE6CB3BFC548B04" box="[588,977,1312,1335]" gridcol="2" gridrow="15" pageId="16" pageNumber="17">0.89 0.92</td>
<td id="76865DC8CC2E000CFB98CB3BFB518B04" box="[1074,1236,1312,1335]" gridcol="3" gridrow="15" pageId="16" pageNumber="17">0.95</td>
<td id="76865DC8CC2E000CFB71CB3BFAE08B04" box="[1243,1381,1312,1335]" gridcol="4" gridrow="15" pageId="16" pageNumber="17">0.92–0.98</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD836F6CC2EFFE3FF6CCB67FE1C88FF" blockId="16.[151,1436,1404,1894]" pageId="16" pageNumber="17">
<emphasis id="B913EAE4CC2EFFE3FF6CCB67FEEC8BA5" bold="true" box="[198,361,1404,1430]" pageId="16" pageNumber="17">Color in life.</emphasis>
BPBM 20192: “Dorsum uniform red-brown. Short black postocular stripe followed by a black dot. Face black. Narrow light-brown line on canthus and rim of upper eyelid. Iris dark copper. 
<collectingCountry id="F3707666CC2EFFE3FA9BCBB8FAEF8B8E" box="[1329,1386,1443,1469]" name="China" pageId="16" pageNumber="17">Chin</collectingCountry>
and throat pale straw mottled with dark gray. Chest and belly uniform straw with minute white punctations. Under thighs straw, mottled distally with dark gray. Rear of thighs same.” BPBM 20197 was described as: “Dorsum rich orange-brown with very narrow tan vertebral stripe and narrow black dosolateral stripes. Sides straw, heavily dusted with gray and with few black spots. Side of face black. Iris orange-brown. 
<collectingCountry id="F3707666CC2EFFE3FB78C825FA8A886B" box="[1234,1295,1598,1624]" name="China" pageId="16" pageNumber="17">Chin</collectingCountry>
, throat, and chest light gray mostly covered by dense charcoal-gray flecking. Belly and under thighs dull yellow with some dark gray flecks, especially toward rear of thighs. Rear of thighs orange-brown heavily covered with charcoal-gray flecks.”
</paragraph>
<paragraph id="8BD836F6CC2EFFE3FF6CC8C3FE7988C1" blockId="16.[151,1436,1404,1894]" box="[198,508,1752,1778]" pageId="16" pageNumber="17">
<emphasis id="B913EAE4CC2EFFE3FF6CC8C3FE8788C1" bold="true" box="[198,258,1752,1778]" pageId="16" pageNumber="17">Call.</emphasis>
Call not heard by us.
</paragraph>
</subSubSection>
<subSubSection id="C37D657DCC2EFFE3FF6CC8E4FB698955" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BD836F6CC2EFFE3FF6CC8E4FD288973" blockId="16.[151,1436,1404,1894]" pageId="16" pageNumber="17">
<emphasis id="B913EAE4CC2EFFE3FF6CC8E4FED6892A" bold="true" box="[198,339,1791,1817]" pageId="16" pageNumber="17">Etymology.</emphasis>
The name is a single-ending latinized Greek compound adjective combining “melanus”, meaning “black”, and “ops”, meaning “face”.
</paragraph>
<paragraph id="8BD836F6CC2EFFE3FF6CC957FB698955" blockId="16.[151,1436,1404,1894]" box="[198,1260,1868,1894]" pageId="16" pageNumber="17">
<emphasis id="B913EAE4CC2EFFE3FF6CC957FE9B8955" bold="true" box="[198,286,1868,1894]" pageId="16" pageNumber="17">Range.</emphasis>
Known only from Sudest Island, Milne Bay Province, 
<collectingCountry id="F3707666CC2EFFE3FC0EC957FB038955" box="[932,1158,1868,1894]" name="Papua New Guinea" pageId="16" pageNumber="17">Papua New Guinea</collectingCountry>
(
<figureCitation id="135C2A73CC2EFFE3FB3CC957FB5A8955" box="[1174,1247,1868,1894]" captionStart="FIGURE 4" captionStartId="17.[151,255,1944,1968]" captionTargetBox="[227,1372,1073,1928]" captionTargetId="figure@17.[227,1372,1073,1928]" captionTargetPageId="17" captionText="FIGURE 4. Map showing known distributions for Cophixalus cupricarenus (inverted triangles), C. desticans (circles), C. iovaorum (triangle), C. melanops (star), and C. tomaiodactylus (squares). Filled symbols are type localities; open symbols are additional localities, as relevant." httpUri="https://zenodo.org/record/188304/files/figure.png" pageId="16" pageNumber="17">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF18667ECC2FFFE2FF3DCD85FCB48DC8" httpUri="https://zenodo.org/record/188303/files/figure.png" pageId="17" pageNumber="18" targetBox="[239,1320,197,899]" targetPageId="17">
<paragraph id="8BD836F6CC2FFFE2FF3DCD85FCB48DC8" blockId="17.[151,1436,926,1019]" pageId="17" pageNumber="18">
<emphasis id="B913EAE4CC2FFFE2FF3DCD85FE9D8D85" bold="true" box="[151,280,926,950]" pageId="17" pageNumber="18">FIGURE 3.</emphasis>
(A) Waveform, (B) power spectrum, and (C) spectrogram of the first nine notes of a 40-note call of paratype of 
<taxonomicName id="4C674D75CC2FFFE2FF1ECDD9FE468DEA" box="[180,451,962,985]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="cupricarenus">
<emphasis id="B913EAE4CC2FFFE2FF1ECDD9FE468DEA" box="[180,451,962,985]" italics="true" pageId="17" pageNumber="18">Cophixalus cupricarenus</emphasis>
</taxonomicName>
(BPBM 20219) recorded by F. Kraus at Cheme, Rossel Island, Milne Bay Province, Papua New Guinea, at 2000 h, 4 May 2004. Air temperature 27.8 °C.
</paragraph>
</caption>
<caption id="DF18667ECC2FFFE2FF3DC983FDF289C6" httpUri="https://zenodo.org/record/188304/files/figure.png" pageId="17" pageNumber="18" targetBox="[227,1372,1073,1928]" targetPageId="17">
<paragraph id="8BD836F6CC2FFFE2FF3DC983FDF289C6" blockId="17.[151,1436,1944,2037]" pageId="17" pageNumber="18">
<emphasis id="B913EAE4CC2FFFE2FF3DC983FE9F8983" bold="true" box="[151,282,1944,1968]" pageId="17" pageNumber="18">FIGURE 4.</emphasis>
Map showing known distributions for 
<taxonomicName id="4C674D75CC2FFFE2FD6BC982FC4A8983" box="[705,975,1945,1968]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="cupricarenus">
<emphasis id="B913EAE4CC2FFFE2FD6BC982FC4A8983" box="[705,975,1945,1968]" italics="true" pageId="17" pageNumber="18">Cophixalus cupricarenus</emphasis>
</taxonomicName>
(inverted triangles), 
<taxonomicName id="4C674D75CC2FFFE2FB18C982FAB38983" box="[1202,1334,1945,1968]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="desticans">
<emphasis id="B913EAE4CC2FFFE2FB18C982FAB38983" box="[1202,1334,1945,1968]" italics="true" pageId="17" pageNumber="18">C. desticans</emphasis>
</taxonomicName>
(circles), 
<taxonomicName id="4C674D75CC2FFFE2FF3DC9A7FEA489E0" box="[151,289,1980,2003]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="iovaorum">
<emphasis id="B913EAE4CC2FFFE2FF3DC9A7FEA489E0" box="[151,289,1980,2003]" italics="true" pageId="17" pageNumber="18">C. iovaorum</emphasis>
</taxonomicName>
(triangle), 
<taxonomicName id="4C674D75CC2FFFE2FE37C9A7FDA289E0" box="[413,551,1980,2003]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="melanops">
<emphasis id="B913EAE4CC2FFFE2FE37C9A7FDA289E0" box="[413,551,1980,2003]" italics="true" pageId="17" pageNumber="18">C. melanops</emphasis>
</taxonomicName>
(star), and 
<taxonomicName id="4C674D75CC2FFFE2FD0CC9A7FCEB89E0" box="[678,878,1980,2003]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tomaiodactylus">
<emphasis id="B913EAE4CC2FFFE2FD0CC9A7FCEB89E0" box="[678,878,1980,2003]" italics="true" pageId="17" pageNumber="18">C. tomaiodactylus</emphasis>
</taxonomicName>
(squares). Filled symbols are type localities; open symbols are additional localities, as relevant.
</paragraph>
</caption>
<subSubSection id="C37D657DCC2CFFE1FF6CCE8CFDAF8F15" pageId="18" pageNumber="19" type="description">
<paragraph id="8BD836F6CC2CFFE1FF6CCE8CFDAF8F15" blockId="18.[151,1436,151,294]" pageId="18" pageNumber="19">
<emphasis id="B913EAE4CC2CFFE1FF6CCE8CFE198E82" bold="true" box="[198,412,151,177]" pageId="18" pageNumber="19">Ecological notes.</emphasis>
Animals were found active on the forest floor during the day in primary lowland rainforest along ridgelines from 
<quantity id="4C9F9B13CC2CFFE1FDB3CEA5FD258EEB" box="[537,672,190,216]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="8.0" metricValueMin="4.4" pageId="18" pageNumber="19" unit="m" value="620.0" valueMax="800.0" valueMin="440.0">440–800 m</quantity>
elevation. The upper 
<quantity id="4C9F9B13CC2CFFE1FC0CCEA5FC778EEB" box="[934,1010,190,216]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="m" value="200.0">200 m</quantity>
of this distribution consists of lowstature, dense cloud forest, and represents the highest reaches of the island. The general area of collection was described by 
<bibRefCitation id="EFF64B07CC2CFFE1FE99CF17FDA38F15" author="Brass" box="[307,550,268,294]" pageId="18" pageNumber="38" refString="Brass, L. J. (1959) Results of the Archbold Expeditions. No. 79. Summary of the Fifth Archbold Expedition to New Guinea (1956 - 1957). Bulletin of the American Museum of Natural History, 118, 5 - 69." type="journal article" year="1959">Brass (1959: 52–53)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>