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<document id="A7F43EE4F35709BEE75E4651E4A7B7F8" ID-CLB-Dataset="2886" ID-DOI="10.1206/0003-0090.454.1.1" ID-GBIF-Dataset="3a7cdaa4-c2f2-4370-9301-c2ab339b2e0d" ID-ISSN="0003-0090" ID-Zenodo-Dep="5788835" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639747398861" checkinUser="felipe" docAuthor="Esselstyn, Jacob A., Achmadi, Anang S., Handika, Heru, Swanson, Mark T., Giarla, Thomas C. &amp; Rowe, Kevin C." docDate="2021" docId="038AB3180155E9414FFEFAFCFC73B19B" docLanguage="en" docName="BulAmeMusNatHist.454.1-108.pdf" docOrigin="Bulletin of the American Museum of Natural History 2021 (454)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-454/issue-1/0003-0090.454.1.1/Fourteen-New-Endemic-Species-of-Shrew-Genus-Crocidura-from-Sulawesi/10.1206/0003-0090.454.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Crocidura musseri Ruedi 1995" docType="treatment" docVersion="7" lastPageNumber="78" masterDocId="FFB3CB600119E90F4F33FF82FF9DB404" masterDocTitle="Fourteen New, Endemic Species Of Shrew (Genus Crocidura) From Sulawesi Reveal A Spectacular Island Radiation" masterLastPageNumber="109" masterPageNumber="1" pageNumber="76" updateTime="1699472789874" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8B03E336718B8311F69A1FEBF5F53C87">Fourteen New, Endemic Species Of Shrew (Genus Crocidura) From Sulawesi Reveal A Spectacular Island Radiation</mods:title>
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<mods:namePart id="ED4B0BD896EA4E598928D7D8AA55424C">Esselstyn, Jacob A.</mods:namePart>
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<mods:namePart id="87FAE15698CF35F6374F33AF14C8B3CA">Achmadi, Anang S.</mods:namePart>
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<mods:namePart id="2D4275D331E99B30B1F5E37681221587">Handika, Heru</mods:namePart>
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<mods:namePart id="AAAC06B522DCC70628722A260D721E4C">Swanson, Mark T.</mods:namePart>
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<mods:namePart id="66DD74417AD1FC53C072F4E4A9B181F0">Giarla, Thomas C.</mods:namePart>
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<mods:namePart id="08538E35910BE6FAA6D81739D873E921">Rowe, Kevin C.</mods:namePart>
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<treatment id="038AB3180155E9414FFEFAFCFC73B19B" ID-DOI="http://doi.org/10.5281/zenodo.5795546" ID-GBIF-Taxon="191690915" ID-Zenodo-Dep="5795546" LSID="urn:lsid:plazi:treatment:038AB3180155E9414FFEFAFCFC73B19B" httpUri="http://treatment.plazi.org/id/038AB3180155E9414FFEFAFCFC73B19B" lastPageId="78" lastPageNumber="78" pageId="76" pageNumber="76">
<subSubSection id="C33951850155E9434FFEFAFCFD86B192" box="[205,539,1406,1430]" pageId="76" pageNumber="76" type="nomenclature">
<paragraph id="8B9C020E0155E9434FFEFAFCFD86B192" blockId="76.[205,539,1406,1430]" box="[205,539,1406,1430]" pageId="76" pageNumber="76">
<heading id="D0D4B5620155E9434FFEFAFCFD86B192" box="[205,539,1406,1430]" centered="true" fontSize="9" level="2" pageId="76" pageNumber="76" reason="2">
<taxonomicName id="4C23798D0155E9434FFEFAFCFD86B192" ID-CoL="6BGCY" authority="Ruedi, 1995" authorityName="Ruedi" authorityYear="1995" box="[205,539,1406,1430]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="76" pageNumber="76" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0155E9434FFEFAFCFE0FB192" box="[205,402,1406,1430]" italics="true" pageId="76" pageNumber="76">Crocidura musseri</emphasis>
<bibRefCitation id="EFB27FFF0155E9434EAAFAFCFD86B192" author="Ruedi, M." box="[409,539,1406,1430]" pageId="76" pageNumber="76" pagination="211 - 265" refId="ref60758" refString="Ruedi, M. 1995. Taxonomic revision of shrews of the genus Crocidura from the Sunda Shelf and Sulawesi with description of two new species (Mammalia: Soricidae). Zoological Journal of the Linnean Society 115: 211 - 265." type="journal article" year="1995">Ruedi, 1995</bibRefCitation>
</taxonomicName>
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</paragraph>
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<paragraph id="8B9C020E0155E9434F5FFA36FE80B1EA" blockId="76.[108,590,1460,1518]" pageId="76" pageNumber="76">
<treatmentCitationGroup id="AB3325200155E9434F5FFA36FE80B1EA" pageId="76" pageNumber="76">
<treatmentCitation id="0A82241F0155E9434F5FFA36FE77B1C8" author="Ruedi, M." box="[108,490,1460,1484]" page="254" pageId="76" pageNumber="76" year="1995">
<taxonomicName id="4C23798D0155E9434F5FFA36FE77B1C8" ID-CoL="6BGCY" authority="Ruedi, 1995: 254" authorityName="Ruedi" authorityPageNumber="254" authorityYear="1995" box="[108,490,1460,1484]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="76" pageNumber="76" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0155E9434F5FFA36FEB0B1C8" box="[108,301,1460,1484]" italics="true" pageId="76" pageNumber="76">Crocidura musseri</emphasis>
<bibRefCitation id="EFB27FFF0155E9434E07FA36FE77B1C8" author="Ruedi, M." box="[308,490,1460,1484]" pageId="76" pageNumber="76" pagination="211 - 265" refId="ref60758" refString="Ruedi, M. 1995. Taxonomic revision of shrews of the genus Crocidura from the Sunda Shelf and Sulawesi with description of two new species (Mammalia: Soricidae). Zoological Journal of the Linnean Society 115: 211 - 265." type="journal article" year="1995">Ruedi, 1995: 254</bibRefCitation>
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. Original description.
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</paragraph>
</subSubSection>
<subSubSection id="C33951850155E9434FBFF989FB28B036" pageId="76" pageNumber="76" type="materials_examined">
<paragraph id="8B9C020E0155E9434FBFF989FB28B036" blockId="76.[108,636,1547,1737]" lastBlockId="76.[684,1214,1017,1735]" pageId="76" pageNumber="76">
<materialsCitation id="3B4B08530155E9434FBFF989FB28B036" ID-GBIF-Occurrence="3421079419" accessionNumber="MZB 16791" collectingDate="1991-08-18" collectionCode="MZB" elevation="2230" latitude="-1.2666667" location="Central Sulawesi" longLatPrecision="1308" longitude="120.25" pageId="76" pageNumber="76" specimenCode="MZB 16791, IZEA 4397" specimenCount="1" specimenCount-female="1" stateProvince="Central Sulawesi" typeStatus="holotype">
<typeStatus id="5498BCAC0155E9434FBFF989FE98B227" box="[140,261,1547,1571]" pageId="76" pageNumber="76" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCode id="DB85AA750155E9434E26F989FE06B227" box="[277,411,1547,1571]" pageId="76" pageNumber="76">MZB 16791</specimenCode>
(=
<specimenCode id="DB85AA750155E9434EF9F989FDD6B227" box="[458,587,1547,1571]" collectionCode="IZEA" pageId="76" pageNumber="76">IZEA 4397</specimenCode>
), an adult
<specimenCount id="9D25C9870155E9434F9DF9AEFF6AB240" box="[174,247,1580,1604]" pageId="76" pageNumber="76" type="female">female</specimenCount>
caught
<date id="FF9D24CE0155E9434E67F9AEFD95B240" box="[340,520,1580,1604]" pageId="76" pageNumber="76" value="1991-08-18">
<collectingDate id="EFD9DD260155E9434E67F9AEFD95B240" box="[340,520,1580,1604]" pageId="76" pageNumber="76" value="1991-08-18">18 August 1991</collectingDate>
</date>
. Measurements from the holotype are TC (head-andbody length):
<quantity id="4CDBAFEB0155E9434E39F9ECFEC7B281" box="[266,346,1646,1670]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="76" pageNumber="76" unit="mm" value="66.0">66 mm</quantity>
× Q (tail length):
<quantity id="4CDBAFEB0155E9434D1FF9ECFDE1B281" box="[556,636,1646,1670]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="76" pageNumber="76" unit="mm" value="57.0">57 mm</quantity>
× PP (hind-foot length):
<quantity id="4CDBAFEB0155E9434E80F90DFDBCB2A2" box="[435,545,1679,1703]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="76" pageNumber="76" unit="mm" value="13.5">13.5 mm</quantity>
= Pds (weight):
<quantity id="4CDBAFEB0155E9434FE5F932FE93B2CD" box="[214,270,1712,1737]" metricMagnitude="-3" metricUnit="kg" metricValue="6.5" pageId="76" pageNumber="76" unit="g" value="6.5">6.5 g</quantity>
. TYPE LOCALITY: Mt. Rorekatimbo,
<collectingRegion id="49E7CCEC0155E9434B59FC7BFC9AB036" country="Indonesia" name="Sulawesi Tengah" pageId="76" pageNumber="76">Central Sulawesi</collectingRegion>
, at
<geoCoordinate id="EE1764C90155E9434C19FB98FCE7B036" box="[810,890,1050,1074]" degrees="1" direction="south" minutes="16" orientation="latitude" pageId="76" pageNumber="76" precision="925" value="-1.2666667">1° 16’ S</geoCoordinate>
,
<geoCoordinate id="EE1764C90155E9434CB6FB98FC6DB036" box="[901,1008,1050,1074]" degrees="120" direction="east" minutes="15" orientation="longitude" pageId="76" pageNumber="76" precision="925" value="120.25">120° 15’ E</geoCoordinate>
,
<quantity id="4CDBAFEB0155E9434CC8FB98FBD7B036" box="[1019,1098,1050,1074]" metricMagnitude="3" metricUnit="m" metricValue="2.23" pageId="76" pageNumber="76" unit="m" value="2230.0">
<elevation id="000EE53D0155E9434CC8FB98FBD7B036" box="[1019,1098,1050,1074]" metricMagnitude="3" metricUnit="m" metricValue="2.23" pageId="76" pageNumber="76" unit="m" value="2230.0">2230 m</elevation>
</quantity>
elevation.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33951850155E9434DFFFBB9FB02B247" pageId="76" pageNumber="76" type="distribution">
<paragraph id="8B9C020E0155E9434DFFFBB9FB02B247" blockId="76.[684,1214,1017,1735]" pageId="76" pageNumber="76">
GEOGRAPHIC DISTRIBUTION: Apparently restricted to high elevation areas (&gt;
<quantity id="4CDBAFEB0155E9434B2CFBDEFBECB070" box="[1055,1137,1116,1140]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="76" pageNumber="76" unit="m" value="1800.0">1800 m</quantity>
) in the northern portions of the west-central area of endemism east of the Palu-Koro Fault in
<collectingRegion id="49E7CCEC0155E9434B5FFB1CFCE8B0D3" country="Indonesia" name="Sulawesi Tengah" pageId="76" pageNumber="76">Central Sulawesi Province</collectingRegion>
(
<figureCitation id="13181E8B0155E9434CBBFB3DFC41B0DC" box="[904,988,1215,1240]" captionStart="FIG" captionStartId="31.[104,146,1395,1419]" captionTargetBox="[104,1603,469,1356]" captionTargetId="graphics-9@31.[188,1530,466,1251]" captionTargetPageId="31" captionText="FIG. 13. Elevational records of all species of Crocidura known from Sulawesi. Each point represents a specimen. For specimens associated with a minimum and maximum elevation, we used the center of the given elevational range. Sample sizes are given above the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5788859" httpUri="https://zenodo.org/record/5788859/files/figure.png" pageId="76" pageNumber="76">figs. 13</figureCitation>
,
<figureCitation id="13181E8B0155E9434CDFFB42FB95B0DC" box="[1004,1032,1216,1240]" captionStart="FIG" captionStartId="83.[108,150,1052,1073]" captionTargetId="graphics-267@83.[275,1045,225,1023]" captionTargetPageId="83" captionText="FIG. 39. Map of Sulawesi showing localities sampled for shrews. Colored areas enclose localities with known records of members of the Ordinary Group. To maintain clarity of presentation, the widespread Crocidura nigripes is not included." figureDoi="http://doi.org/10.5281/zenodo.5789164" httpUri="https://zenodo.org/record/5789164/files/figure.png" pageId="76" pageNumber="76">39</figureCitation>
;
<tableCitation id="C6A137B50155E9434B24FB3DFBF8B0DC" box="[1047,1125,1215,1240]" captionStart="TABLE 3" captionStartId="27.[611,691,226,250]" captionTargetBox="[117,1200,329,1675]" captionTargetPageId="27" captionText="TABLE 3 Elevational Ranges (m) and Species Richness of Crocidura on Mountains of Sulawesia" httpUri="http://table.plazi.org/id/DF5C52860102E9144D50FF60FBDEB51A" pageId="76" pageNumber="76" tableUuid="DF5C52860102E9144D50FF60FBDEB51A">table 3</tableCitation>
).
<bibRefCitation id="EFB27FFF0155E9434B4DFB3DFD6EB0FC" author="Ruedi, M." pageId="76" pageNumber="76" pagination="211 - 265" refId="ref60758" refString="Ruedi, M. 1995. Taxonomic revision of shrews of the genus Crocidura from the Sunda Shelf and Sulawesi with description of two new species (Mammalia: Soricidae). Zoological Journal of the Linnean Society 115: 211 - 265." type="journal article" year="1995">Ruedi (1995)</bibRefCitation>
reported specimens of this species from only the
<typeStatus id="5498BCAC0155E9434C21FA80FCDCB11E" box="[786,833,1282,1306]" pageId="76" pageNumber="76">type</typeStatus>
locality on Mt. Rorekatimbo. We collected specimens that match Ruedi’s description and are mitochondrially identical (based on cytochrome
<emphasis id="B957DE1C0155E9434C00FAE7FCDDB179" box="[819,832,1381,1405]" italics="true" pageId="76" pageNumber="76">b</emphasis>
sequences from
<bibRefCitation id="EFB27FFF0155E9434CC4FAE7FB29B179" author="Ruedi, M. &amp; M. Auberson &amp; V. Savolainen" box="[1015,1204,1381,1405]" pageId="76" pageNumber="76" pagination="567 - 571" refId="ref60802" refString="Ruedi, M., M. Auberson, and V. Savolainen. 1998. Biogeography of Sulawesian shrews: testing for their origin with a parametric bootstrap on molecular data. Molecular Phylogenetics and Evolution 9: 567 - 571." type="journal article" year="1998">Ruedi et al., 1998</bibRefCitation>
) from the same mountain, at three nearby sites (
<geoCoordinate id="EE1764C90155E9434D87FA25FCB8B1BB" box="[692,805,1447,1471]" degrees="1.3093" direction="south" orientation="latitude" pageId="76" pageNumber="76" precision="5" value="-1.3093">1.3093° S</geoCoordinate>
,
<geoCoordinate id="EE1764C90155E9434C0AFA25FC58B1BB" box="[825,965,1447,1471]" degrees="120.3092" direction="east" orientation="longitude" pageId="76" pageNumber="76" precision="5" value="120.3092">120.3092 °E</geoCoordinate>
,
<quantity id="4CDBAFEB0155E9434CEAFA25FBA9B1BA" box="[985,1076,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="76" pageNumber="76" unit="m" value="2020.0">2020 m</quantity>
;
<geoCoordinate id="EE1764C90155E9434B7BFA25FB25B1BB" box="[1096,1208,1447,1471]" degrees="1.2926" direction="south" orientation="latitude" pageId="76" pageNumber="76" precision="5" value="-1.2926">1.2926° S</geoCoordinate>
,
<geoCoordinate id="EE1764C90155E9434D9FFA4AFCB1B1E4" box="[684,812,1480,1504]" degrees="120.3064" direction="east" orientation="longitude" pageId="76" pageNumber="76" precision="5" value="120.3064">120.3064° E</geoCoordinate>
,
<quantity id="4CDBAFEB0155E9434C05FA4AFC1AB1E4" box="[822,903,1480,1504]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="76" pageNumber="76" unit="m" value="2180.0">2180 m</quantity>
;
<geoCoordinate id="EE1764C90155E9434CA2FA4AFC69B1E4" box="[913,1012,1480,1504]" degrees="1.2884" direction="south" orientation="latitude" pageId="76" pageNumber="76" precision="5" value="-1.2884">1.2884° S</geoCoordinate>
,
<geoCoordinate id="EE1764C90155E9434CCCFA4AFBE3B1E4" box="[1023,1150,1480,1504]" degrees="120.3104" direction="east" orientation="longitude" pageId="76" pageNumber="76" precision="5" value="120.3104">120.3104° E</geoCoordinate>
,
<quantity id="4CDBAFEB0155E9434BBAFA4AFD58B205" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="76" pageNumber="76" unit="m" value="2250.0">2250 m</quantity>
). Musser collected additional specimens from
<quantity id="4CDBAFEB0155E9434D9FF988FCDCB226" box="[684,833,1546,1570]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="76" pageNumber="76" unit="m" value="2050.0" valueMax="2300.0" valueMin="1800.0">1800–2300 m</quantity>
elevation on the neighboring Mt. Nokilalaki (ca.
<quantity id="4CDBAFEB0155E9434C7CF9A9FC0EB247" box="[847,915,1579,1603]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="76" pageNumber="76" unit="km" value="15.0">15 km</quantity>
W of Mt. Rorekatimbo).
</paragraph>
</subSubSection>
<subSubSection id="C33951850155E9424DFFF9CFFEEBB241" lastPageId="77" lastPageNumber="77" pageId="76" pageNumber="76" type="diagnosis">
<paragraph id="8B9C020E0155E9434DFFF9CFFB23B2C3" blockId="76.[684,1214,1017,1735]" pageId="76" pageNumber="76">
EMENDED DIAGNOSIS: A medium-sized shrew (
<tableCitation id="C6A137B50155E9434D86F9EFFC95B281" box="[693,776,1645,1669]" captionStart="TABLE 2" captionStartId="25.[610,690,226,250]" captionTargetBox="[119,1184,329,1682]" captionTargetPageId="25" captionText="TABLE 2 Descriptive Statisticsa for External Measurements (mm) and Mass (g) for Species of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860100E9164D51FF60FBE5B523" pageId="76" pageNumber="76" tableUuid="DF5C52860100E9164D51FF60FBE5B523">tables 2</tableCitation>
,
<tableCitation id="C6A137B50155E9434C20F9EFFCB3B281" box="[787,814,1645,1669]" captionStart="TABLE 14" captionStartId="77.[603,683,226,250]" captionTargetBox="[119,1190,358,1139]" captionTargetPageId="77" captionText="TABLE 14 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Ordinary Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860154E9424D68FF60FCC5B540" pageId="76" pageNumber="76" tableUuid="DF5C52860154E9424D68FF60FCC5B540">14</tableCitation>
) with a thick, dark brown pelage that is only slightly paler on the venter. Foot, ear, tail, and rhinarium color closely match the pelage. Tail
</paragraph>
<caption id="DF5C52860154E9424D68FF60FCC5B540" ID-Table-UUID="DF5C52860154E9424D68FF60FCC5B540" httpUri="http://table.plazi.org/id/DF5C52860154E9424D68FF60FCC5B540" pageId="77" pageNumber="77" startId="77.[603,683,226,250]" targetBox="[119,1190,358,1139]" targetIsTable="true" targetPageId="77">
<paragraph id="8B9C020E0154E9424D68FF60FD51B4FE" blockId="77.[265,1055,226,324]" box="[603,716,226,250]" pageId="77" pageNumber="77">
<heading id="D0D4B5620154E9424D68FF60FD51B4FE" box="[603,716,226,250]" centered="true" fontSize="10" level="2" pageId="77" pageNumber="77" reason="4">TABLE 14</heading>
</paragraph>
<paragraph id="8B9C020E0154E9424E3AFE93FCC5B540" blockId="77.[265,1055,226,324]" pageId="77" pageNumber="77">
<emphasis id="B957DE1C0154E9424E3AFE93FE41B523" bold="true" box="[265,476,273,295]" pageId="77" pageNumber="77">Descriptive Statistics</emphasis>
<superScript id="7C56AF460154E9424EEFFE92FE7EB51A" attach="left" box="[476,483,272,286]" fontSize="6" pageId="77" pageNumber="77">a</superScript>
<emphasis id="B957DE1C0154E9424EDAFE93FCC5B540" bold="true" pageId="77" pageNumber="77">
for Craniodental Measurements (mm) for Species of the Ordinary Group of
<collectingRegion id="49E7CCEC0154E9424DA4FEACFD70B540" box="[663,749,302,324]" country="Indonesia" name="Sulawesi Utara" pageId="77" pageNumber="77">Sulawesi</collectingRegion>
<taxonomicName id="4C23798D0154E9424DC7FEACFCC5B540" box="[756,856,302,324]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="B957DE1C0154E9424DC7FEACFCC5B540" bold="true" box="[756,856,302,324]" italics="true" pageId="77" pageNumber="77">Crocidura</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</caption>
<paragraph id="8B9C020E0154E9424E76FEE4FB01B077" pageId="77" pageNumber="77">
<table id="F923F0AE015416F04F44FEE4FB3BB077" box="[119,1190,358,1139]" gridcols="6" gridrows="13" pageId="77" pageNumber="77">
<tr id="3513004C015416F04F44FEE4FB3BB57D" box="[119,1190,358,377]" gridrow="0" pageId="77" pageNumber="77" rowspan-0="1">
<th id="76C26930015416F04E1AFEE4FE24B57D" box="[297,441,358,377]" gridcol="1" gridrow="0" pageId="77" pageNumber="77">
<taxonomicName id="4C23798D0154E9424E76FEE4FE00B57D" authorityName="Ruedi" authorityYear="1995" box="[325,413,358,377]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="musseri">C. musseri</taxonomicName>
</th>
<th id="76C26930015416F04ED2FEE4FDEDB57D" box="[481,624,358,377]" gridcol="2" gridrow="0" pageId="77" pageNumber="77">
<taxonomicName id="4C23798D0154E9424ECFFEE4FDCBB57D" authorityName="Miller and Hollister" authorityYear="1921" box="[508,598,358,377]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="nigripes">C. nigripes</taxonomicName>
</th>
<th id="76C26930015416F04DAEFEE4FCB0B57D" box="[669,813,358,377]" gridcol="3" gridrow="0" pageId="77" pageNumber="77">
<taxonomicName id="4C23798D0154E9424D87FEE4FC8BB57D" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[692,790,358,377]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="normalis">C. normalis</taxonomicName>
</th>
<th id="76C26930015416F04C68FEE4FC77B57D" box="[859,1002,358,377]" gridcol="4" gridrow="0" pageId="77" pageNumber="77">
<taxonomicName id="4C23798D0154E9424C5CFEE4FC4BB57D" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[879,982,358,377]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="ordinaria">C. ordinaria</taxonomicName>
</th>
<th id="76C26930015416F04B24FEE4FB3BB57D" box="[1047,1190,358,377]" gridcol="5" gridrow="0" pageId="77" pageNumber="77">
<taxonomicName id="4C23798D0154E9424B0FFEE4FB1CB57D" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[1084,1153,358,377]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C0154E9424B0FFEE4FB1CB57D" box="[1084,1153,358,377]" italics="true" pageId="77" pageNumber="77">C. solita</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3513004C015416F04F44FE09FB3BB5BD" box="[119,1190,395,441]" gridrow="1" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FE09FE97B5BD" box="[119,266,395,441]" gridcol="0" gridrow="1" pageId="77" pageNumber="77">Condyloincisive length</th>
<td id="76C26930015416F04E1AFE09FE24B5BD" box="[297,441,395,441]" gridcol="1" gridrow="1" pageId="77" pageNumber="77">20.13 ± 0.446 (19.17–20.78) 17</td>
<td id="76C26930015416F04ED2FE09FDEDB5BD" box="[481,624,395,441]" gridcol="2" gridrow="1" pageId="77" pageNumber="77">22.47 ± 0.733 (20.91–24.32) 80</td>
<td id="76C26930015416F04DAEFE09FCB0B5BD" box="[669,813,395,441]" gridcol="3" gridrow="1" pageId="77" pageNumber="77">18.73 ± 0.276 (18.29–19.66) 27</td>
<td id="76C26930015416F04C68FE09FC77B5BD" box="[859,1002,395,441]" gridcol="4" gridrow="1" pageId="77" pageNumber="77">20.38 ± 0.491 (19.47–21.59) 15</td>
<td id="76C26930015416F04B24FE09FB3BB5BD" box="[1047,1190,395,441]" gridcol="5" gridrow="1" pageId="77" pageNumber="77">20.13 ± 0.584 (19.25–21.55) 43</td>
</tr>
<tr id="3513004C015416F04F44FE48FB3BB5FC" box="[119,1190,458,504]" gridrow="2" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FE48FE97B5FC" box="[119,266,458,504]" gridcol="0" gridrow="2" pageId="77" pageNumber="77">Braincase breadth</th>
<td id="76C26930015416F04E1AFE48FE24B5FC" box="[297,441,458,504]" gridcol="1" gridrow="2" pageId="77" pageNumber="77">9.74 ± 0.233 (9.42–10.26) 18</td>
<td id="76C26930015416F04ED2FE48FDEDB5FC" box="[481,624,458,504]" gridcol="2" gridrow="2" pageId="77" pageNumber="77">9.94 ± 0.315 (9.32–10.78) 77</td>
<td id="76C26930015416F04DAEFE48FCB0B5FC" box="[669,813,458,504]" gridcol="3" gridrow="2" pageId="77" pageNumber="77">8.61 ± 0.168 (8.45–9.06) 27</td>
<td id="76C26930015416F04C68FE48FC77B5FC" box="[859,1002,458,504]" gridcol="4" gridrow="2" pageId="77" pageNumber="77">9.79 ± 0.178 (9.48–10.06) 15</td>
<td id="76C26930015416F04B24FE48FB3BB5FC" box="[1047,1190,458,504]" gridcol="5" gridrow="2" pageId="77" pageNumber="77">9.51 ± 0.263 (8.8–10.12) 43</td>
</tr>
<tr id="3513004C015416F04F44FD88FB3BB63C" box="[119,1190,522,568]" gridrow="3" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FD88FE97B63C" box="[119,266,522,568]" gridcol="0" gridrow="3" pageId="77" pageNumber="77">Interorbital width</th>
<td id="76C26930015416F04E1AFD88FE24B63C" box="[297,441,522,568]" gridcol="1" gridrow="3" pageId="77" pageNumber="77">4.91 ± 0.164 (4.61–5.2) 18</td>
<td id="76C26930015416F04ED2FD88FDEDB63C" box="[481,624,522,568]" gridcol="2" gridrow="3" pageId="77" pageNumber="77">4.71 ± 0.196 (4.3–5.08) 85</td>
<td id="76C26930015416F04DAEFD88FCB0B63C" box="[669,813,522,568]" gridcol="3" gridrow="3" pageId="77" pageNumber="77">4.3 ± 0.114 (4.11–4.58) 28</td>
<td id="76C26930015416F04C68FD88FC77B63C" box="[859,1002,522,568]" gridcol="4" gridrow="3" pageId="77" pageNumber="77">4.86 ± 0.137 (4.65–5.12) 16</td>
<td id="76C26930015416F04B24FD88FB3BB63C" box="[1047,1190,522,568]" gridcol="5" gridrow="3" pageId="77" pageNumber="77">4.68 ± 0.167 (4.36–5.06) 43</td>
</tr>
<tr id="3513004C015416F04F44FDCBFB3BB673" box="[119,1190,585,631]" gridrow="4" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FDCBFE97B673" box="[119,266,585,631]" gridcol="0" gridrow="4" pageId="77" pageNumber="77">Rostral length</th>
<td id="76C26930015416F04E1AFDCBFE24B673" box="[297,441,585,631]" gridcol="1" gridrow="4" pageId="77" pageNumber="77">8.16 ± 0.205 (7.76–8.39) 17</td>
<td id="76C26930015416F04ED2FDCBFDEDB673" box="[481,624,585,631]" gridcol="2" gridrow="4" pageId="77" pageNumber="77">9.31 ± 0.337 (8.45–10.09) 86</td>
<td id="76C26930015416F04DAEFDCBFCB0B673" box="[669,813,585,631]" gridcol="3" gridrow="4" pageId="77" pageNumber="77">7.36 ± 0.163 (7.09–7.75) 28</td>
<td id="76C26930015416F04C68FDCBFC77B673" box="[859,1002,585,631]" gridcol="4" gridrow="4" pageId="77" pageNumber="77">8.25 ± 0.285 (7.77–8.95) 16</td>
<td id="76C26930015416F04B24FDCBFB3BB673" box="[1047,1190,585,631]" gridcol="5" gridrow="4" pageId="77" pageNumber="77">8.15 ± 0.363 (7.2–9.17) 43</td>
</tr>
<tr id="3513004C015416F04F44FD0BFB3BB6B3" box="[119,1190,649,695]" gridrow="5" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FD0BFE97B6B3" box="[119,266,649,695]" gridcol="0" gridrow="5" pageId="77" pageNumber="77">Postpalatal width</th>
<td id="76C26930015416F04E1AFD0BFE24B6B3" box="[297,441,649,695]" gridcol="1" gridrow="5" pageId="77" pageNumber="77">3.84 ± 0.12 (3.59–4.03) 18</td>
<td id="76C26930015416F04ED2FD0BFDEDB6B3" box="[481,624,649,695]" gridcol="2" gridrow="5" pageId="77" pageNumber="77">4.02 ± 0.156 (3.7–4.38) 85</td>
<td id="76C26930015416F04DAEFD0BFCB0B6B3" box="[669,813,649,695]" gridcol="3" gridrow="5" pageId="77" pageNumber="77">3.34 ± 0.106 (3.15–3.68) 28</td>
<td id="76C26930015416F04C68FD0BFC77B6B3" box="[859,1002,649,695]" gridcol="4" gridrow="5" pageId="77" pageNumber="77">3.96 ± 0.18 (3.7–4.36) 16</td>
<td id="76C26930015416F04B24FD0BFB3BB6B3" box="[1047,1190,649,695]" gridcol="5" gridrow="5" pageId="77" pageNumber="77">3.88 ± 0.147 (3.57–4.22) 43</td>
</tr>
<tr id="3513004C015416F04F44FD4AFB3BB6F2" box="[119,1190,712,758]" gridrow="6" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FD4AFE97B6F2" box="[119,266,712,758]" gridcol="0" gridrow="6" pageId="77" pageNumber="77">Rostral width</th>
<td id="76C26930015416F04E1AFD4AFE24B6F2" box="[297,441,712,758]" gridcol="1" gridrow="6" pageId="77" pageNumber="77">2.65 ± 0.147 (2.46–3.03) 17</td>
<td id="76C26930015416F04ED2FD4AFDEDB6F2" box="[481,624,712,758]" gridcol="2" gridrow="6" pageId="77" pageNumber="77">3.02 ± 0.17 (2.62–3.58) 86</td>
<td id="76C26930015416F04DAEFD4AFCB0B6F2" box="[669,813,712,758]" gridcol="3" gridrow="6" pageId="77" pageNumber="77">2.47 ± 0.172 (2.08–2.71) 28</td>
<td id="76C26930015416F04C68FD4AFC77B6F2" box="[859,1002,712,758]" gridcol="4" gridrow="6" pageId="77" pageNumber="77">2.96 ± 0.139 (2.73–3.18) 16</td>
<td id="76C26930015416F04B24FD4AFB3BB6F2" box="[1047,1190,712,758]" gridcol="5" gridrow="6" pageId="77" pageNumber="77">2.73 ± 0.198 (2.31–3.2) 43</td>
</tr>
<tr id="3513004C015416F04F44FC8AFB3BB732" box="[119,1190,776,822]" gridrow="7" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FC8AFE97B732" box="[119,266,776,822]" gridcol="0" gridrow="7" pageId="77" pageNumber="77">Postpalatal length</th>
<td id="76C26930015416F04E1AFC8AFE24B732" box="[297,441,776,822]" gridcol="1" gridrow="7" pageId="77" pageNumber="77">8.89 ± 0.222 (8.4–9.23) 18</td>
<td id="76C26930015416F04ED2FC8AFDEDB732" box="[481,624,776,822]" gridcol="2" gridrow="7" pageId="77" pageNumber="77">9.79 ± 0.366 (8.9–10.54) 80</td>
<td id="76C26930015416F04DAEFC8AFCB0B732" box="[669,813,776,822]" gridcol="3" gridrow="7" pageId="77" pageNumber="77">8.41 ± 0.231 (8.08–9.03) 27</td>
<td id="76C26930015416F04C68FC8AFC77B732" box="[859,1002,776,822]" gridcol="4" gridrow="7" pageId="77" pageNumber="77">9.1 ± 0.297 (8.7–9.87) 15</td>
<td id="76C26930015416F04B24FC8AFB3BB732" box="[1047,1190,776,822]" gridcol="5" gridrow="7" pageId="77" pageNumber="77">8.89 ± 0.304 (8.41–9.51) 44</td>
</tr>
<tr id="3513004C015416F04F44FCC5FB3BB771" box="[119,1190,839,885]" gridrow="8" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FCC5FE97B771" box="[119,266,839,885]" gridcol="0" gridrow="8" pageId="77" pageNumber="77">Condyle to gle- noid fossa</th>
<td id="76C26930015416F04E1AFCC5FE24B771" box="[297,441,839,885]" gridcol="1" gridrow="8" pageId="77" pageNumber="77">7.93 ± 0.186 (7.52–8.24) 18</td>
<td id="76C26930015416F04ED2FCC5FDEDB771" box="[481,624,839,885]" gridcol="2" gridrow="8" pageId="77" pageNumber="77">8.37 ± 0.281 (7.78–8.9) 80</td>
<td id="76C26930015416F04DAEFCC5FCB0B771" box="[669,813,839,885]" gridcol="3" gridrow="8" pageId="77" pageNumber="77">7.72 ± 0.134 (7.47–8.01) 27</td>
<td id="76C26930015416F04C68FCC5FC77B771" box="[859,1002,839,885]" gridcol="4" gridrow="8" pageId="77" pageNumber="77">8.09 ± 0.194 (7.77–8.43) 15</td>
<td id="76C26930015416F04B24FCC5FB3BB771" box="[1047,1190,839,885]" gridcol="5" gridrow="8" pageId="77" pageNumber="77">7.93 ± 0.224 (7.35–8.49) 43</td>
</tr>
<tr id="3513004C015416F04F44FC05FB3BB7B1" box="[119,1190,903,949]" gridrow="9" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FC05FE97B7B1" box="[119,266,903,949]" gridcol="0" gridrow="9" pageId="77" pageNumber="77">Upper toothrow length</th>
<td id="76C26930015416F04E1AFC05FE24B7B1" box="[297,441,903,949]" gridcol="1" gridrow="9" pageId="77" pageNumber="77">8.84 ± 0.229 (8.35–9.19) 17</td>
<td id="76C26930015416F04ED2FC05FDEDB7B1" box="[481,624,903,949]" gridcol="2" gridrow="9" pageId="77" pageNumber="77">10.2 ± 0.359 (9.24–11.02) 86</td>
<td id="76C26930015416F04DAEFC05FCB0B7B1" box="[669,813,903,949]" gridcol="3" gridrow="9" pageId="77" pageNumber="77">8.07 ± 0.211 (7.69–8.7) 28</td>
<td id="76C26930015416F04C68FC05FC77B7B1" box="[859,1002,903,949]" gridcol="4" gridrow="9" pageId="77" pageNumber="77">9.04 ± 0.254 (8.63–9.68) 16</td>
<td id="76C26930015416F04B24FC05FB3BB7B1" box="[1047,1190,903,949]" gridcol="5" gridrow="9" pageId="77" pageNumber="77">8.91 ± 0.327 (8.07–9.89) 43</td>
</tr>
<tr id="3513004C015416F04F44FC44FB3BB7F0" box="[119,1190,966,1012]" gridrow="10" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FC44FE97B7F0" box="[119,266,966,1012]" gridcol="0" gridrow="10" pageId="77" pageNumber="77">P4 to M3 length</th>
<td id="76C26930015416F04E1AFC44FE24B7F0" box="[297,441,966,1012]" gridcol="1" gridrow="10" pageId="77" pageNumber="77">5.08 ± 0.182 (4.7–5.34) 18</td>
<td id="76C26930015416F04ED2FC44FDEDB7F0" box="[481,624,966,1012]" gridcol="2" gridrow="10" pageId="77" pageNumber="77">5.82 ± 0.238 (5.17–6.24) 86</td>
<td id="76C26930015416F04DAEFC44FCB0B7F0" box="[669,813,966,1012]" gridcol="3" gridrow="10" pageId="77" pageNumber="77">4.49 ± 0.163 (4.23–5.02) 28</td>
<td id="76C26930015416F04C68FC44FC77B7F0" box="[859,1002,966,1012]" gridcol="4" gridrow="10" pageId="77" pageNumber="77">5.24 ± 0.167 (4.99–5.62) 16</td>
<td id="76C26930015416F04B24FC44FB3BB7F0" box="[1047,1190,966,1012]" gridcol="5" gridrow="10" pageId="77" pageNumber="77">5.11 ± 0.195 (4.72–5.67) 43</td>
</tr>
<tr id="3513004C015416F04F44FB84FB3BB030" box="[119,1190,1030,1076]" gridrow="11" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FB84FE97B030" box="[119,266,1030,1076]" gridcol="0" gridrow="11" pageId="77" pageNumber="77">M2 to M2 labial width</th>
<td id="76C26930015416F04E1AFB84FE24B030" box="[297,441,1030,1076]" gridcol="1" gridrow="11" pageId="77" pageNumber="77">5.86 ± 0.238 (5.44–6.16) 17</td>
<td id="76C26930015416F04ED2FB84FDEDB030" box="[481,624,1030,1076]" gridcol="2" gridrow="11" pageId="77" pageNumber="77">6.84 ± 0.253 (6.37–7.52) 86</td>
<td id="76C26930015416F04DAEFB84FCB0B030" box="[669,813,1030,1076]" gridcol="3" gridrow="11" pageId="77" pageNumber="77">5.12 ± 0.152 (4.85–5.63) 28</td>
<td id="76C26930015416F04C68FB84FC77B030" box="[859,1002,1030,1076]" gridcol="4" gridrow="11" pageId="77" pageNumber="77">6.08 ± 0.286 (5.69–6.77) 16</td>
<td id="76C26930015416F04B24FB84FB3BB030" box="[1047,1190,1030,1076]" gridcol="5" gridrow="11" pageId="77" pageNumber="77">5.9 ± 0.217 (5.53–6.61) 43</td>
</tr>
<tr id="3513004C015416F04F44FBC7FB3BB077" box="[119,1190,1093,1139]" gridrow="12" pageId="77" pageNumber="77">
<th id="76C26930015416F04F44FBC7FE97B077" box="[119,266,1093,1139]" gridcol="0" gridrow="12" pageId="77" pageNumber="77">Palatal width</th>
<td id="76C26930015416F04E1AFBC7FE24B077" box="[297,441,1093,1139]" gridcol="1" gridrow="12" pageId="77" pageNumber="77">2.52 ± 0.085 (2.36–2.64) 18</td>
<td id="76C26930015416F04ED2FBC7FDEDB077" box="[481,624,1093,1139]" gridcol="2" gridrow="12" pageId="77" pageNumber="77">2.74 ± 0.134 (2.45–3.19) 85</td>
<td id="76C26930015416F04DAEFBC7FCB0B077" box="[669,813,1093,1139]" gridcol="3" gridrow="12" pageId="77" pageNumber="77">2.25 ± 0.096 (2.01–2.43) 28</td>
<td id="76C26930015416F04C68FBC7FC77B077" box="[859,1002,1093,1139]" gridcol="4" gridrow="12" pageId="77" pageNumber="77">2.49 ± 0.118 (2.31–2.8) 16</td>
<td id="76C26930015416F04B24FBC7FB3BB077" box="[1047,1190,1093,1139]" gridcol="5" gridrow="12" pageId="77" pageNumber="77">2.46 ± 0.105 (2.24–2.71) 43</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9C020E0154E9424F5FFB09FC49B09A" blockId="77.[108,980,1163,1183]" box="[108,980,1163,1183]" pageId="77" pageNumber="77">
<tableNote id="76C503800154E9424F5FFB09FC49B09A" box="[108,980,1163,1183]" pageId="77" pageNumber="77" targetBox="[119,1190,358,1139]" targetPageId="77">
<superScript id="7C56AF460154E9424F5FFB09FFEFB093" attach="left" box="[108,114,1163,1175]" fontSize="5" pageId="77" pageNumber="77">a</superScript>
The sample mean ± one standard deviation, the observed range in parentheses, and the sample size.
</tableNote>
</paragraph>
<subSection id="E2AC1BE50154E9424F5FFB40FEEBB241" pageId="77" pageNumber="77" type="multiple">
<paragraph id="8B9C020E0154E9424F5FFB40FEEBB241" blockId="77.[108,638,1218,1738]" pageId="77" pageNumber="77">
length is shorter than head-and-body length. The tail has inconspicuous applied hairs that are visible with the naked eye only on the distal third of tail length and sparse bristle hairs spread from the tail base to at least its midpoint (
<figureCitation id="13181E8B0154E9424ED3FAC4FDA3B15A" box="[480,574,1350,1374]" captionStart="FIG" captionStartId="78.[107,149,902,923]" captionTargetBox="[108,1212,224,872]" captionTargetId="figure-292@78.[108,1212,224,872]" captionTargetPageId="78" captionText="FIG. 36. Images showing the ventral surface of the right hind foot and dorsal surface of the tail base (approximately 1 cm from rump) and tail tip from two darkly pigmented members of the Ordinary Group: A, Crocidura musseri, FMNH 213267; and B, C. nigripes, LSUMZ 37018. Scale bars apply to all images in panel A and to foot (upper) and tail (lower) in panel B." figureDoi="http://doi.org/10.5281/zenodo.5788899" httpUri="https://zenodo.org/record/5788899/files/figure.png" pageId="77" pageNumber="77">fig. 36A</figureCitation>
). The braincase is notably inflated, with great width, giving an overall wedge-shaped appearance when viewed from the dorsal aspect (
<figureCitation id="13181E8B0154E9424EE8FA2BFDA1B1C6" box="[475,572,1449,1474]" captionStart="FIG" captionStartId="79.[108,150,1388,1409]" captionTargetBox="[108,1209,224,1362]" captionTargetId="figure-121@79.[108,1209,224,1362]" captionTargetPageId="79" captionText="FIG. 37. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of two darkly pigmented members of the Ordinary Group: A, Crocidura musseri, FMNH 213255; and B, C. nigripes, FMNH 210611." figureDoi="http://doi.org/10.5281/zenodo.5788901" httpUri="https://zenodo.org/record/5788901/files/figure.png" pageId="77" pageNumber="77">fig. 37A</figureCitation>
). The squamosal-parietal suture is often open, leaving a tear-drop-shaped opening anteriorly, directly below the anterior opening of the sinus canal (sensu
<bibRefCitation id="EFB27FFF0154E9424F8BF9ACFEF7B241" author="McDowell, S. B., Jr." box="[184,362,1581,1606]" pageId="77" pageNumber="77" pagination="131 - 214" refId="ref59152" refString="McDowell, S. B., Jr. 1958. The greater Antillean insectivores. Bulletin of the American Museum of Natural History 115 (3): 131 - 214." type="journal article" year="1958">McDowell, 1958</bibRefCitation>
).
</paragraph>
</subSection>
</subSubSection>
<subSubSection id="C33951850154E9414FBFF9CDFE90B1E5" lastPageId="78" lastPageNumber="78" pageId="77" pageNumber="77" type="description">
<paragraph id="8B9C020E0154E9414FBFF9CDFE90B1E5" blockId="77.[108,638,1218,1738]" lastBlockId="78.[108,639,1051,1737]" lastPageId="78" lastPageNumber="78" pageId="77" pageNumber="77">
COMPARISONS:
<taxonomicName id="4C23798D0154E9424E09F9CDFD9FB262" authorityName="Ruedi" authorityYear="1995" box="[314,514,1615,1639]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0154E9424E09F9CDFD9FB262" box="[314,514,1615,1639]" italics="true" pageId="77" pageNumber="77">Crocidura musseri</emphasis>
</taxonomicName>
is smaller than members of the Elongata Subgroup and all Rhoditis Group members except
<taxonomicName id="4C23798D0154E9424EEAF913FDDCB2AD" box="[473,577,1681,1705]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="pallida">
<emphasis id="B957DE1C0154E9424EEAF913FDDCB2AD" box="[473,577,1681,1705]" italics="true" pageId="77" pageNumber="77">C. pallida</emphasis>
</taxonomicName>
. It is larger than all members of the Small-Bodied Group. Within the Ordinary Group,
<taxonomicName id="4C23798D0154E9424B06FB40FB3EB0DE" authorityName="Ruedi" authorityYear="1995" box="[1077,1187,1218,1242]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0154E9424B06FB40FB3EB0DE" box="[1077,1187,1218,1242]" italics="true" pageId="77" pageNumber="77">C. musseri</emphasis>
</taxonomicName>
is smaller than
<taxonomicName id="4C23798D0154E9424C0CFB61FC2FB0FF" authorityName="Miller and Hollister" authorityYear="1921" box="[831,946,1251,1275]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B957DE1C0154E9424C0CFB61FC2FB0FF" box="[831,946,1251,1275]" italics="true" pageId="77" pageNumber="77">C. nigripes</emphasis>
</taxonomicName>
, slightly larger than the sympatric
<taxonomicName id="4C23798D0154E9424C29FA86FC0EB118" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[794,915,1284,1308]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="normalis">
<emphasis id="B957DE1C0154E9424C29FA86FC0EB118" box="[794,915,1284,1308]" italics="true" pageId="77" pageNumber="77">C. normalis</emphasis>
</taxonomicName>
, and comparable to
<taxonomicName id="4C23798D0154E9424B5EFA86FD78B139" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C0154E9424B5EFA86FD78B139" italics="true" pageId="77" pageNumber="77">C. ordinaria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C23798D0154E9424C15FAA4FC1CB139" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[806,897,1317,1341]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C0154E9424C15FAA4FC1CB139" box="[806,897,1317,1341]" italics="true" pageId="77" pageNumber="77">C. solita</emphasis>
</taxonomicName>
. Relative to head-and-body length,
<taxonomicName id="4C23798D0154E9424DC8FAC5FCF5B15A" authorityName="Ruedi" authorityYear="1995" box="[763,872,1351,1374]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0154E9424DC8FAC5FCF5B15A" box="[763,872,1351,1374]" italics="true" pageId="77" pageNumber="77">C. musseri</emphasis>
</taxonomicName>
has a shorter tail than all members of the Long-Tailed and Rhoditis groups,
<taxonomicName id="4C23798D0154E9424B96FAEAFD66B1A4" authorityName="Miller and Hollister" authorityYear="1921" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B957DE1C0154E9424B96FAEAFD66B1A4" italics="true" pageId="77" pageNumber="77">C. nigripes</emphasis>
</taxonomicName>
, and the sympatric
<taxonomicName id="4C23798D0154E9424CFCFA0BFBBCB1A4" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[975,1057,1416,1440]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C0154E9424CFCFA0BFBBCB1A4" box="[975,1057,1416,1440]" italics="true" pageId="77" pageNumber="77">C. solita</emphasis>
</taxonomicName>
. The pelage of
<taxonomicName id="4C23798D0154E9424D9EFA28FC81B1C5" authorityName="Ruedi" authorityYear="1995" box="[685,796,1450,1473]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0154E9424D9EFA28FC81B1C5" box="[685,796,1450,1473]" italics="true" pageId="77" pageNumber="77">C. musseri</emphasis>
</taxonomicName>
is thicker than that of any other shrew species we recognize from
<collectingRegion id="49E7CCEC0154E9424CF5FA48FBBCB1E6" box="[966,1057,1482,1506]" country="Indonesia" name="Sulawesi Utara" pageId="77" pageNumber="77">Sulawesi</collectingRegion>
, except
<taxonomicName id="4C23798D0154E9424B41FA49FD60B207" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="caudicrassa">
<emphasis id="B957DE1C0154E9424B41FA49FD60B207" italics="true" pageId="77" pageNumber="77">C. caudicrassa</emphasis>
</taxonomicName>
. Overall, the color of
<taxonomicName id="4C23798D0154E9424CEFFA6EFBD5B207" authorityName="Ruedi" authorityYear="1995" box="[988,1096,1516,1539]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0154E9424CEFFA6EFBD5B207" box="[988,1096,1516,1539]" italics="true" pageId="77" pageNumber="77">C. musseri</emphasis>
</taxonomicName>
is substantially darker than specimens from the Long-Tailed and Rhoditis groups,
<taxonomicName id="4C23798D0154E9424C97F9ACFC7BB242" box="[932,998,1582,1606]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="lea">
<emphasis id="B957DE1C0154E9424C97F9ACFC7BB242" box="[932,998,1582,1606]" italics="true" pageId="77" pageNumber="77">C. lea</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C23798D0154E9424B1EF9ACFB2BB242" box="[1069,1206,1582,1606]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="mediocris">
<emphasis id="B957DE1C0154E9424B1EF9ACFB2BB242" box="[1069,1206,1582,1606]" italics="true" pageId="77" pageNumber="77">C. mediocris</emphasis>
</taxonomicName>
. Within the Ordinary Group,
<taxonomicName id="4C23798D0154E9424CD6F9CDFBCEB262" authorityName="Ruedi" authorityYear="1995" box="[997,1107,1615,1638]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0154E9424CD6F9CDFBCEB262" box="[997,1107,1615,1638]" italics="true" pageId="77" pageNumber="77">C. musseri</emphasis>
</taxonomicName>
is slightly darker than
<taxonomicName id="4C23798D0154E9424C01F9F2FC14B28C" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[818,905,1648,1672]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C0154E9424C01F9F2FC14B28C" box="[818,905,1648,1672]" italics="true" pageId="77" pageNumber="77">C. solita</emphasis>
</taxonomicName>
and
<taxonomicName id="4C23798D0154E9424CF2F9F2FBA1B28C" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[961,1084,1648,1672]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="77" pageNumber="77" phylum="Chordata" rank="species" species="normalis">
<emphasis id="B957DE1C0154E9424CF2F9F2FBA1B28C" box="[961,1084,1648,1672]" italics="true" pageId="77" pageNumber="77">C. normalis</emphasis>
</taxonomicName>
. Relative to skull length, skull width at both the braincase and interorbital region is greater than most other spe- cies, including all members of the Ordinary Group (
<figureCitation id="13181E8B0157E9414F40FBBEFF5DB050" box="[115,192,1084,1108]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="78" pageNumber="78">figs. 10</figureCitation>
,
<figureCitation id="13181E8B0157E9414FF8FBBEFF75B050" box="[203,232,1084,1108]" captionStart="FIG" captionStartId="79.[108,150,1388,1409]" captionTargetBox="[108,1209,224,1362]" captionTargetId="figure-121@79.[108,1209,224,1362]" captionTargetPageId="79" captionText="FIG. 37. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of two darkly pigmented members of the Ordinary Group: A, Crocidura musseri, FMNH 213255; and B, C. nigripes, FMNH 210611." figureDoi="http://doi.org/10.5281/zenodo.5788901" httpUri="https://zenodo.org/record/5788901/files/figure.png" pageId="78" pageNumber="78">37</figureCitation>
). The unusual width of the skull (
<tableCitation id="C6A137B50157E9414D7BFBBEFF15B071" captionStart="TABLE 14" captionStartId="77.[603,683,226,250]" captionTargetBox="[119,1190,358,1139]" captionTargetPageId="77" captionText="TABLE 14 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Ordinary Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860154E9424D68FF60FCC5B540" pageId="78" pageNumber="78" tableUuid="DF5C52860154E9424D68FF60FCC5B540">table 14</tableCitation>
) gives
<taxonomicName id="4C23798D0157E9414FE3FBDCFEA1B071" authorityName="Ruedi" authorityYear="1995" box="[208,316,1118,1141]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0157E9414FE3FBDCFEA1B071" box="[208,316,1118,1141]" italics="true" pageId="78" pageNumber="78">C. musseri</emphasis>
</taxonomicName>
a more wedge-shaped appearance than any other species when viewed from the dorsal aspect (
<figureCitation id="13181E8B0157E9414E3AFB1DFEF8B0BC" box="[265,357,1183,1208]" captionStart="FIG" captionStartId="79.[108,150,1388,1409]" captionTargetBox="[108,1209,224,1362]" captionTargetId="figure-121@79.[108,1209,224,1362]" captionTargetPageId="79" captionText="FIG. 37. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of two darkly pigmented members of the Ordinary Group: A, Crocidura musseri, FMNH 213255; and B, C. nigripes, FMNH 210611." figureDoi="http://doi.org/10.5281/zenodo.5788901" httpUri="https://zenodo.org/record/5788901/files/figure.png" pageId="78" pageNumber="78">fig. 37A</figureCitation>
).
<taxonomicName id="4C23798D0157E9414E4AFB1DFDA7B0B3" authorityName="Miller and Hollister" authorityYear="1921" box="[377,570,1183,1207]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B957DE1C0157E9414E4AFB1DFDA7B0B3" box="[377,570,1183,1207]" italics="true" pageId="78" pageNumber="78">Crocidura nigripes</emphasis>
</taxonomicName>
has a more angular braincase, a more robust dentition, and a much narrower relative interorbital region (IOW/CIL). Relative rostral length (RL/CIL) in
<taxonomicName id="4C23798D0157E9414D55FA81FF26B13F" authorityName="Ruedi" authorityYear="1995" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0157E9414D55FA81FF26B13F" italics="true" pageId="78" pageNumber="78">C. musseri</emphasis>
</taxonomicName>
is less than in
<taxonomicName id="4C23798D0157E9414E69FAA6FE58B13F" authorityName="Miller and Hollister" authorityYear="1921" box="[346,453,1316,1339]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B957DE1C0157E9414E69FAA6FE58B13F" box="[346,453,1316,1339]" italics="true" pageId="78" pageNumber="78">C. nigripes</emphasis>
</taxonomicName>
, but greater than in
<taxonomicName id="4C23798D0157E9414FB4FAC7FF62B159" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[135,255,1349,1373]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="normalis">
<emphasis id="B957DE1C0157E9414FB4FAC7FF62B159" box="[135,255,1349,1373]" italics="true" pageId="78" pageNumber="78">C. normalis</emphasis>
</taxonomicName>
(
<figureCitation id="13181E8B0157E9414E38FAC7FED2B159" box="[267,335,1349,1373]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="78" pageNumber="78">fig. 10</figureCitation>
).
<taxonomicName id="4C23798D0157E9414E52FAC7FD81B159" authorityName="Ruedi" authorityYear="1995" box="[353,540,1349,1373]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0157E9414E52FAC7FD81B159" box="[353,540,1349,1373]" italics="true" pageId="78" pageNumber="78">Crocidura musseri</emphasis>
</taxonomicName>
is distinguished from all sympatric shrews by a combination of its short tail length, thick and dark pelage, and wide, wedge-shaped skull with small teeth (
<figureCitation id="13181E8B0157E9414F40FA4BFF2EB1E5" box="[115,179,1481,1505]" captionStart="FIG" captionStartId="24.[108,150,1587,1608]" captionTargetBox="[137,1192,224,1561]" captionTargetId="graphics-9@24.[203,1191,224,1433]" captionTargetPageId="24" captionText="FIG. 9. Box plots showing variation in external measurements from all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick- Tailed Group). All measurements in mm. HBL = head-and-body length." figureDoi="http://doi.org/10.5281/zenodo.5788853" httpUri="https://zenodo.org/record/5788853/files/figure.png" pageId="78" pageNumber="78">figs. 9</figureCitation>
,
<figureCitation id="13181E8B0157E9414F8DFA4BFF44B1E5" box="[190,217,1481,1505]" captionStart="FIG" captionStartId="78.[107,149,902,923]" captionTargetBox="[108,1212,224,872]" captionTargetId="figure-292@78.[108,1212,224,872]" captionTargetPageId="78" captionText="FIG. 36. Images showing the ventral surface of the right hind foot and dorsal surface of the tail base (approximately 1 cm from rump) and tail tip from two darkly pigmented members of the Ordinary Group: A, Crocidura musseri, FMNH 213267; and B, C. nigripes, LSUMZ 37018. Scale bars apply to all images in panel A and to foot (upper) and tail (lower) in panel B." figureDoi="http://doi.org/10.5281/zenodo.5788899" httpUri="https://zenodo.org/record/5788899/files/figure.png" pageId="78" pageNumber="78">36</figureCitation>
,
<figureCitation id="13181E8B0157E9414FD7FA4BFE9CB1E5" box="[228,257,1481,1505]" captionStart="FIG" captionStartId="84.[104,146,1285,1309]" captionTargetBox="[104,1612,588,1258]" captionTargetId="figure-10@84.[104,1612,588,1258]" captionTargetPageId="84" captionText="FIG. 40. Images showing the ventral surface of the hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from three members of the Ordinary Group: A, Crocidura normalis, FMNH 213174 (left hind foot); B, C. ordinaria, FMNH 218726 (right hind foot); and C, C. solita, FMNH 213015 (left hind foot). Upper scale bar for foot, lower bar corresponds to both tail images." figureDoi="http://doi.org/10.5281/zenodo.5788905" httpUri="https://zenodo.org/record/5788905/files/figure.png" pageId="78" pageNumber="78">40</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5C52860157E9414F58FC04FDD7B7F7" ID-DOI="http://doi.org/10.5281/zenodo.5788899" ID-Zenodo-Dep="5788899" httpUri="https://zenodo.org/record/5788899/files/figure.png" pageId="78" pageNumber="78" startId="78.[107,149,902,923]" targetBox="[108,1212,224,872]" targetPageId="78">
<paragraph id="8B9C020E0157E9414F58FC04FDD7B7F7" blockId="78.[107,1211,900,1012]" pageId="78" pageNumber="78">
FIG. 36. Images showing the ventral surface of the right hind foot and dorsal surface of the tail base (approximately 1 cm from rump) and tail tip from two darkly pigmented members of the Ordinary Group:
<emphasis id="B957DE1C0157E9414B55FC23FBE3B7BD" bold="true" box="[1126,1150,929,953]" pageId="78" pageNumber="78">A,</emphasis>
<taxonomicName id="4C23798D0157E9414BB7FC20FED0B7D2" authority=", FMNH" authorityName="FMNH" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0157E9414BB7FC20FF6BB7D2" italics="true" pageId="78" pageNumber="78">Crocidura musseri</emphasis>
, FMNH
</taxonomicName>
213267; and
<emphasis id="B957DE1C0157E9414EE9FC3DFE6DB7D3" bold="true" box="[474,496,959,983]" pageId="78" pageNumber="78">B,</emphasis>
<taxonomicName id="4C23798D0157E9414ECBFC3DFD5EB7D3" authority=", LSUMZ" authorityName="LSUMZ" box="[504,707,959,983]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B957DE1C0157E9414ECBFC3DFDFEB7D2" box="[504,611,959,982]" italics="true" pageId="78" pageNumber="78">C. nigripes</emphasis>
, LSUMZ
</taxonomicName>
37018. Scale bars apply to all images in panel A and to foot (upper) and tail (lower) in panel B.
</paragraph>
</caption>
<subSubSection id="C33951850157E9414FBEFA68FB15B0FE" pageId="78" pageNumber="78" type="discussion">
<paragraph id="8B9C020E0157E9414FBEFA68FB15B0FE" blockId="78.[108,639,1051,1737]" lastBlockId="78.[684,1213,1051,1439]" pageId="78" pageNumber="78">
COMMENTS: Our description of
<taxonomicName id="4C23798D0157E9414EE8FA68FF0DB227" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0157E9414EE8FA68FF0DB227" italics="true" pageId="78" pageNumber="78">Crocidura musseri</emphasis>
</taxonomicName>
is based primarily on the series of specimens we collected in 2011 very near the type locality. This series matches the
<typeStatus id="5498BCAC0157E9414E53F9CFFE21B261" box="[352,444,1613,1637]" pageId="78" pageNumber="78" type="holotype">holotype</typeStatus>
(MZB 16791) and the description of
<bibRefCitation id="EFB27FFF0157E9414E0EF9ECFE4DB282" author="Ruedi, M." box="[317,464,1646,1670]" pageId="78" pageNumber="78" pagination="211 - 265" refId="ref60758" refString="Ruedi, M. 1995. Taxonomic revision of shrews of the genus Crocidura from the Sunda Shelf and Sulawesi with description of two new species (Mammalia: Soricidae). Zoological Journal of the Linnean Society 115: 211 - 265." type="journal article" year="1995">Ruedi (1995)</bibRefCitation>
. Cytochrome
<emphasis id="B957DE1C0157E9414D5CF9ECFDE1B282" box="[623,636,1646,1670]" italics="true" pageId="78" pageNumber="78">b</emphasis>
sequences from our series are identical to one from a
<typeStatus id="5498BCAC0157E9414F4EF933FF45B2CD" box="[125,216,1713,1737]" pageId="78" pageNumber="78" type="paratype">paratype</typeStatus>
(IZEA 4403;
<bibRefCitation id="EFB27FFF0157E9414E50F932FD8AB2CD" author="Ruedi, M. &amp; M. Auberson &amp; V. Savolainen" box="[355,535,1712,1737]" pageId="78" pageNumber="78" pagination="567 - 571" refId="ref60802" refString="Ruedi, M., M. Auberson, and V. Savolainen. 1998. Biogeography of Sulawesian shrews: testing for their origin with a parametric bootstrap on molecular data. Molecular Phylogenetics and Evolution 9: 567 - 571." type="journal article" year="1998">Ruedi et al., 1998</bibRefCitation>
). None of our phylogenetic analyses supported a close relationship of
<taxonomicName id="4C23798D0157E9414C18FBBFFC01B050" box="[811,924,1085,1108]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0157E9414C18FBBFFC01B050" box="[811,924,1085,1108]" italics="true" pageId="78" pageNumber="78">C. musseri</emphasis>
</taxonomicName>
to any other species. Our mitogenome result placed
<taxonomicName id="4C23798D0157E9414CF3FBDCFBB6B071" box="[960,1067,1118,1141]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C0157E9414CF3FBDCFBB6B071" box="[960,1067,1118,1141]" italics="true" pageId="78" pageNumber="78">C. musseri</emphasis>
</taxonomicName>
as sister to all other members of Sulawesi’s endemic radiation with some support (
<figureCitation id="13181E8B0157E9414CB3FB1DFC2AB0B3" box="[896,951,1183,1207]" captionStart="FIG" captionStartId="18.[108,150,1596,1617]" captionTargetBox="[111,1203,227,1567]" captionTargetId="graphics-32@18.[111,872,237,1559]" captionTargetPageId="18" captionText="FIG. 5. Maximum likelihood estimate of the mitochondrial gene tree derived from an analysis of 14,007 characters (representing protein-coding and rRNA genes) from 83 samples. Bootstrap support values &lt;95 are shown at nodes. Tips are labeled with the species, locality, and voucher number." figureDoi="http://doi.org/10.5281/zenodo.5788845" httpUri="https://zenodo.org/record/5788845/files/figure.png" pageId="78" pageNumber="78">fig. 5</figureCitation>
), while our UCE species tree placed it as sister to
<taxonomicName id="4C23798D0157E9414C88FB43FBF1B0DC" box="[955,1132,1216,1240]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="microelongata">
<emphasis id="B957DE1C0157E9414C88FB43FBF1B0DC" box="[955,1132,1216,1240]" italics="true" pageId="78" pageNumber="78">C. microelongata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C23798D0157E9414B96FB43FD6FB0FE" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="78" pageNumber="78" phylum="Chordata" rank="species" species="pallida">
<emphasis id="B957DE1C0157E9414B96FB43FD6FB0FE" italics="true" pageId="78" pageNumber="78">C. pallida</emphasis>
</taxonomicName>
, but without statistical support (
<figureCitation id="13181E8B0157E9414B77FB60FBE1B0FE" box="[1092,1148,1250,1274]" captionStart="FIG" captionStartId="21.[108,149,1653,1674]" captionTargetBox="[209,1136,226,1678]" captionTargetId="graphics-38@20.[180,704,233,1669]" captionTargetPageId="20" captionText="FIG. 7. Estimated species tree from analysis of 3940 ultraconserved element loci in ASTRAL. Samples from Sulawesi are labeled with the species name, locality, and catalog number. Asterisks indicate type specimens from Miller and Hollister (1921). Local posterior probabilities &lt;0.95 are shown. Tip branch lengths are arbitrary." figureDoi="http://doi.org/10.5281/zenodo.5789145" httpUri="https://zenodo.org/record/5789145/files/figure.png" pageId="78" pageNumber="78">fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33951850157E9414DFFFA81FC73B19B" pageId="78" pageNumber="78" type="materials_examined">
<paragraph id="8B9C020E0157E9414DFFFA81FC73B19B" blockId="78.[684,1213,1051,1439]" pageId="78" pageNumber="78">
<collectionCode id="ED329ACB0157E9414DFFFA81FC25B11F" box="[716,952,1283,1307]" pageId="78" pageNumber="78">SPECIMENS EXAMINED</collectionCode>
:
<materialsCitation id="3B4B08530157E9414CF0FA81FCE9B159" ID-GBIF-Occurrence="3421079334" collectionCode="AMNH" location="Mt. Nokilalaki" pageId="78" pageNumber="78" specimenCode="AMNH 223495, 225516, 225517, 225521, 225525-225527, 225530, 225531" specimenCount="1">
<location id="8EFC54D50157E9414CF0FA81FBC6B11F" LSID="urn:lsid:plazi:treatment:038AB3180155E9414FFEFAFCFC73B19B:8EFC54D50157E9414CF0FA81FBC6B11F" box="[963,1115,1283,1307]" name="Mt. Nokilalaki" pageId="78" pageNumber="78">Mt. Nokilalaki</location>
(
<specimenCode id="DB85AA750157E9414B54FA81FD66B138" pageId="78" pageNumber="78">AMNH 223495</specimenCode>
,
<specimenCode id="DB85AA750157E9414C37FAA6FCCEB138" box="[772,851,1316,1340]" pageId="78" pageNumber="78">225516</specimenCode>
,
<specimenCode id="DB85AA750157E9414C6EFAA6FC31B138" box="[861,940,1316,1340]" pageId="78" pageNumber="78">225517</specimenCode>
,
<specimenCode id="DB85AA750157E9414C86FAA6FB99B138" box="[949,1028,1316,1340]" pageId="78" pageNumber="78">225521</specimenCode>
,
<specimenCode id="DB85AA750157E9414B3DFAA6FB25B138" box="[1038,1208,1316,1340]" pageId="78" pageNumber="78">225525–225527</specimenCode>
,
<specimenCode id="DB85AA750157E9414D9FFAC7FD62B159" box="[684,767,1349,1373]" pageId="78" pageNumber="78">225530</specimenCode>
,
<specimenCode id="DB85AA750157E9414C27FAC7FCF4B159" box="[788,873,1349,1373]" pageId="78" pageNumber="78">225531</specimenCode>
),
</materialsCitation>
<materialsCitation id="3B4B08530157E9414CB5FAC7FCC4B19B" ID-GBIF-Occurrence="3421079304" collectionCode="FMNH" location="Mt. Rorekatimbo" pageId="78" pageNumber="78" specimenCode="FMNH 213249-213251, 213253-213256, 213258-213269, 213442, 213443" specimenCount="1">
<location id="8EFC54D50157E9414CB5FAC7FBCDB159" LSID="urn:lsid:plazi:treatment:038AB3180155E9414FFEFAFCFC73B19B:8EFC54D50157E9414CB5FAC7FBCDB159" box="[902,1104,1349,1373]" name="Mt. Rorekatimbo" pageId="78" pageNumber="78">Mt. Rorekatimbo</location>
(
<specimenCode id="DB85AA750157E9414B5AFAC7FCC8B17A" pageId="78" pageNumber="78">FMNH 213249–213251</specimenCode>
,
<specimenCode id="DB85AA750157E9414C6DFAE4FB9BB17A" box="[862,1030,1382,1406]" pageId="78" pageNumber="78">213253–213256</specimenCode>
,
<specimenCode id="DB85AA750157E9414B23FAE4FB24B17A" box="[1040,1209,1382,1406]" pageId="78" pageNumber="78">213258–213269</specimenCode>
,
<specimenCode id="DB85AA750157E9414D9FFA05FD66B19B" box="[684,763,1415,1439]" pageId="78" pageNumber="78">213442</specimenCode>
,
<specimenCode id="DB85AA750157E9414C36FA05FCC9B19B" box="[773,852,1415,1439]" pageId="78" pageNumber="78">213443</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B4B08530157E9414C6CFA05FC73B19B" ID-GBIF-Occurrence="3421079316" box="[863,1006,1415,1439]" collectionCode="MZB" pageId="78" pageNumber="78" specimenCode="MZB 16791" specimenCount="1">
<specimenCode id="DB85AA750157E9414C6CFA05FC7FB19B" box="[863,994,1415,1439]" pageId="78" pageNumber="78">MZB 16791</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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