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<document id="D719F646B70C8F88BE49CF891745D665" ID-CLB-Dataset="31455" ID-DOI="10.11646/zootaxa.4084.3.9" ID-GBIF-Dataset="3da62e39-7304-4933-a4eb-aecbc1c5d77e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052247" ID-ZooBank="5FEBA8FA-FA3B-421A-8F8C-2F5DD3243D30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510872430653" checkinUser="plazi" docAuthor="Sankaran, Pradeep M. &amp; Sebastian, Pothalil A." docDate="2016" docId="7973E75CE231FFCB72F6FBD307B8FB12" docLanguage="en" docName="zootaxa.4084.3.9.pdf" docOrigin="Zootaxa 4084 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Shearella alii Sankaran &amp; Sebastian 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="446" masterDocId="854A9F24E230FFC87261FFEF0576FFAF" masterDocTitle="A checklist of Indian armored spiders (Araneae, Tetrablemmidae) with the description of a new species from the Western Ghats" masterLastPageNumber="450" masterPageNumber="443" pageNumber="444" updateTime="1698498153148" updateUser="ExternalLinkService">
<mods:mods id="0BABAB6A4AA68B2A15ADF9F4BE1D31B8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7901E6829ABA79D67B8211528D218396">
<mods:title id="C3A46B339C116445C5667281857C74DD">A checklist of Indian armored spiders (Araneae, Tetrablemmidae) with the description of a new species from the Western Ghats</mods:title>
</mods:titleInfo>
<mods:name id="ED2A1A68FB197DBC92083463663AC7ED" type="personal">
<mods:role id="3FBC31CFD3A40E3C933DCED71CD2D0B1">
<mods:roleTerm id="0799A2F3F989873FF26E84251419FDA2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B071B259197E5E41C5E56185EC691D9">Sankaran, Pradeep M.</mods:namePart>
</mods:name>
<mods:name id="946197896FA031D8188B764783E1E077" type="personal">
<mods:role id="ED1BAAEFCB8CDC490FC8CF5A64AD5200">
<mods:roleTerm id="5BF4ED5F64B5FEBACCC27B404CEF3C7D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13BA676C1804FD90D090B89DBFAF1F61">Sebastian, Pothalil A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="03502DC04283DE950C8537F9FBC124A8">text</mods:typeOfResource>
<mods:relatedItem id="22DBCB7A076BB7E071DB676384BFE09A" type="host">
<mods:titleInfo id="AD378C3AB5DCF7DC4F01B999363F2A21">
<mods:title id="B039311317BBEB4DE188C3EE886B8DBD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="81481DA86878D29B9B0C7472B64F4D59">
<mods:date id="EF4C952E0FAC0BC756939023677917AE">2016</mods:date>
<mods:detail id="7E19312900CE52C023381F1B57C1A1E7" type="pubDate">
<mods:number id="92B85604CE7CDE0E58266B12395D16B9">2016-02-26</mods:number>
</mods:detail>
<mods:detail id="3B400BF6150829F75D27BAA1B68CDE23" type="volume">
<mods:number id="282D4CBADD4E98A9F265021B0983997A">4084</mods:number>
</mods:detail>
<mods:detail id="804E891EFE08A9FEC90A58E91A644F8E" type="issue">
<mods:number id="73C7E36C4F1871E3EDE720C1746BACFB">3</mods:number>
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<mods:start id="8FCF089FBC25E3CB1EEA901B05997720">443</mods:start>
<mods:end id="74F105813A31259AD2B5DA9D35687F57">450</mods:end>
</mods:extent>
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<mods:classification id="D602211866302F332134F6E7335CFC52">journal article</mods:classification>
<mods:identifier id="3BD7900FFD068574301CA933DD631AA0" type="CLB-Dataset">31455</mods:identifier>
<mods:identifier id="D7405912D20FE6BC15AA2B461563E972" type="DOI">10.11646/zootaxa.4084.3.9</mods:identifier>
<mods:identifier id="89EDDE4D8A91AEF96F22A08EB0795F10" type="GBIF-Dataset">3da62e39-7304-4933-a4eb-aecbc1c5d77e</mods:identifier>
<mods:identifier id="E6B6F5DD63EF8AE336A3A71E3330FB0A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2AE602985762CF9A0A87EB12DF193C75" type="Zenodo-Dep">1052247</mods:identifier>
<mods:identifier id="FD7888DDCB2E800A57E210150750DAC8" type="ZooBank">5FEBA8FA-FA3B-421A-8F8C-2F5DD3243D30</mods:identifier>
</mods:mods>
<treatment id="7973E75CE231FFCB72F6FBD307B8FB12" ID-DOI="http://doi.org/10.5281/zenodo.5316031" ID-GBIF-Taxon="137308582" ID-Zenodo-Dep="5316031" LSID="urn:lsid:plazi:treatment:7973E75CE231FFCB72F6FBD307B8FB12" httpUri="http://treatment.plazi.org/id/7973E75CE231FFCB72F6FBD307B8FB12" lastPageId="3" lastPageNumber="446" pageId="1" pageNumber="444">
<subSubSection id="B9C005C1E231FFC972F6FBD304B7FBD7" pageId="1" pageNumber="444" type="nomenclature">
<paragraph id="F165564AE231FFC972F6FBD304E8FBF8" blockId="1.[151,449,1084,1145]" box="[151,414,1084,1111]" pageId="1" pageNumber="444">
<heading id="AA2DE126E231FFC972F6FBD304E8FBF8" bold="true" box="[151,414,1084,1111]" fontSize="11" level="1" pageId="1" pageNumber="444" reason="1">
<emphasis id="C3AE8A58E231FFC972F6FBD304E8FBF8" bold="true" box="[151,414,1084,1111]" pageId="1" pageNumber="444">
<taxonomicName id="36DA2DC9E231FFC972F6FBD3044FFBF9" ID-CoL="4X48V" authority="Sankaran &amp; Sebastian, 2016" authorityName="Sankaran &amp; Sebastian" authorityYear="2016" box="[151,313,1084,1110]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="alii" status="sp. nov.">
<emphasis id="C3AE8A58E231FFC972F6FBD3044FFBF9" bold="true" box="[151,313,1084,1110]" italics="true" pageId="1" pageNumber="444">Shearella alii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D89D3723E231FFC97321FBD204E8FBF8" box="[320,414,1085,1111]" pageId="1" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F165564AE231FFC972F6FB8F04B7FBD7" blockId="1.[151,449,1084,1145]" box="[151,449,1120,1145]" pageId="1" pageNumber="444">
(
<figureCitation id="69E14ACFE231FFC972FEFB8F046AFBD7" box="[159,284,1120,1145]" captionStart="FIGURE 1" captionStartId="2.[151,250,1385,1407]" captionTargetBox="[160,1427,527,1354]" captionTargetId="figure@2.[151,1436,517,1363]" captionTargetPageId="2" captionText="FIGURE 1. Shearella alii sp. nov. A Male holotype, dorsal view; B Female paratype, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1052249/files/figure.png" pageId="1" pageNumber="444">Figs 1A–B</figureCitation>
,
<figureCitation id="69E14ACFE231FFC97349FB8F041FFBD7" box="[296,361,1120,1144]" captionStart="FIGURE 2" captionStartId="3.[151,250,1029,1051]" captionTargetBox="[158,1428,198,1008]" captionTargetId="figure@3.[150,1436,193,1008]" captionTargetPageId="3" captionText="FIGURE 2. Shearella alii sp. nov. A Male holotype habitus, retrolateral view; B Female paratype, habitus, retrolateral view; C Male, frontal view; D Female, frontal view; E Male, left chelicera, retrolateral view; F Male, left leg I, retrolateral view. Scale bars: A – B 0.15 mm; C, 0.08 mm; D, 0.06 mm; E, 0.02 mm; F, 0.40 mm." httpUri="https://zenodo.org/record/1052251/files/figure.png" pageId="1" pageNumber="444">2A–F</figureCitation>
,
<figureCitation id="69E14ACFE231FFC97315FB8F04CCFBD7" box="[372,442,1120,1144]" captionStart="FIGURE 3" captionStartId="4.[151,250,1694,1716]" captionTargetBox="[334,1252,201,1670]" captionTargetId="figure@4.[327,1259,195,1672]" captionTargetPageId="4" captionText="FIGURE 3. Shearella alii sp. nov. A Male, left pedipalp, prolateral view; B Same, retrolateral view; C Female, opisthosoma, ventral view; D Female, genital area, dorsal view. Abbreviations: B bulb, CP central process, E embolus, EF epigastric fold, GO genital orifice, IVP inner vulval plate, PA preanal plate, PEG perigenital plate, POG postgenital plate, S spermophor, SR seminal receptacle, VS vulval stem. Scale bars: A – B 0.16 mm; C, 0.10 mm; D, 0.01 mm." httpUri="https://zenodo.org/record/1052253/files/figure.png" pageId="1" pageNumber="444">3A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B9C005C1E231FFC972F6FB470623FAA3" pageId="1" pageNumber="444" type="materials_examined">
<paragraph id="F165564AE231FFC972F6FB470623FAA3" blockId="1.[151,1437,1192,2012]" pageId="1" pageNumber="444">
<materialsCitation id="41B25C17E231FFC972F6FB470714FB48" ID-GBIF-Occurrence="1701200752" collectionCode="ADSH" country="India" county="Male" elevation="67" latitude="10.433416" location="Chimmony Wildlife Sanctuary" longLatPrecision="1" longitude="76.48234" municipality="Thrissur" pageId="1" pageNumber="444" specimenCode="ADSH100261" specimenCount="67" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="C3AE8A58E231FFC972F6FB4704A4FB6E" bold="true" box="[151,466,1192,1217]" pageId="1" pageNumber="444">
<typeStatus id="2E61E8E8E231FFC972F6FB4705A4FB6E" box="[151,210,1192,1217]" pageId="1" pageNumber="444">Type</typeStatus>
material:
<typeStatus id="2E61E8E8E231FFC9733DFB4704BBFB6E" box="[348,461,1192,1217]" pageId="1" pageNumber="444" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCounty id="18042EC6E231FFC97381FB47076CFB6E" box="[480,538,1192,1217]" pageId="1" pageNumber="444">Male</collectingCounty>
(
<specimenCode id="A17CFE31E231FFC97051FB4707A0FB6E" box="[560,726,1192,1217]" collectionCode="ADSH" country="India" httpUri="http://grbio.org/cool/7csr-pk9c" name="Arachnology Division, Sacred Heart College" pageId="1" pageNumber="444">
ADSH
<date id="8564708AE231FFC970E0FB4707A0FB6E" box="[641,726,1192,1217]" pageId="1" pageNumber="444" value="61-10-02">100261</date>
</specimenCode>
),
<emphasis id="C3AE8A58E231FFC97091FB4706C6FB6E" bold="true" box="[752,944,1192,1217]" pageId="1" pageNumber="444">
<collectingCountry id="89CD16DAE231FFC97091FB470630FB6E" box="[752,838,1192,1217]" name="India" pageId="1" pageNumber="444">INDIA</collectingCountry>
:
<collectingRegion id="331E98A8E231FFC9713BFB4706DEFB6E" box="[858,936,1192,1217]" country="India" name="Kerala" pageId="1" pageNumber="444">
<emphasis id="C3AE8A58E231FFC9713BFB4706DEFB6E" bold="true" box="[858,936,1192,1217]" italics="true" pageId="1" pageNumber="444">Kerala</emphasis>
</collectingRegion>
:
</emphasis>
<collectingMunicipality id="1101CC30E231FFC971DEFB470168FB6E" box="[959,1054,1192,1217]" pageId="1" pageNumber="444">Thrissur</collectingMunicipality>
,
<location id="F4050091E231FFC97651FB4700EEFB6F" LSID="urn:lsid:plazi:treatment:7973E75CE231FFCB72F6FBD307B8FB12:F4050091E231FFC97651FB4700EEFB6F" box="[1072,1432,1192,1217]" country="India" county="Male" latitude="10.433416" longLatPrecision="1" longitude="76.48234" municipality="Thrissur" name="Chimmony Wildlife Sanctuary" pageId="1" pageNumber="444" stateProvince="Kerala">Chimmony Wildlife Sanctuary</location>
,
<geoCoordinate id="94EE308DE231FFC972F6FB20044DFB48" box="[151,315,1227,1255]" direction="north" orientation="latitude" pageId="1" pageNumber="444" precision="1" value="10.433416">10o26'00.30''N</geoCoordinate>
,
<geoCoordinate id="94EE308DE231FFC97326FB21049EFB48" box="[327,488,1227,1255]" direction="east" orientation="longitude" pageId="1" pageNumber="444" precision="1" value="76.48234">76o28'56.41''E</geoCoordinate>
,
<specimenCount id="E7DC9DC3E231FFC97394FB210743FB48" box="[501,565,1230,1255]" pageId="1" pageNumber="444" type="generic">
<elevation id="7AF7B179E231FFC97394FB210747FB48" box="[501,561,1230,1255]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="444" unit="m" value="67.0">
<quantity id="3622FBAFE231FFC97394FB210747FB48" box="[501,561,1230,1255]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="444" unit="m" value="67.0">67 m</quantity>
</elevation>
.
</specimenCount>
alt.
</materialsCitation>
,
<date id="8564708AE231FFC9700FFB200671FB48" box="[622,775,1231,1256]" pageId="1" pageNumber="444" value="2013-05-31">31 May 2013</date>
, M.S. Pradeep leg., from forest litter, by hand;
<materialsCitation id="41B25C17E231FFC9777AFB210627FAA3" ID-GBIF-Occurrence="1701200751" collectionCode="ADSH" country="India" county="Male" elevation="67" latitude="10.433416" location="Chimmony Wildlife Sanctuary" longLatPrecision="1" longitude="76.48234" municipality="Thrissur" pageId="1" pageNumber="444" specimenCode="ADSH100262" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Kerala" typeStatus="paratype">
<emphasis id="C3AE8A58E231FFC9777AFB2100EAFB48" bold="true" box="[1307,1436,1230,1255]" pageId="1" pageNumber="444">
<typeStatus id="2E61E8E8E231FFC9777AFB2100E1FB48" box="[1307,1431,1230,1255]" pageId="1" pageNumber="444" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="E7DC9DC3E231FFC972F6FB1C0586FAA3" box="[151,240,1267,1292]" pageId="1" pageNumber="444" type="male">2 males</specimenCount>
,
<specimenCount id="E7DC9DC3E231FFC9729AFB1C0411FAA3" box="[251,359,1267,1292]" pageId="1" pageNumber="444" type="female">3 females</specimenCount>
(
<specimenCode id="A17CFE31E231FFC97317FB1C076AFAA3" box="[374,540,1267,1292]" collectionCode="ADSH" country="India" httpUri="http://grbio.org/cool/7csr-pk9c" name="Arachnology Division, Sacred Heart College" pageId="1" pageNumber="444">
ADSH
<date id="8564708AE231FFC973A7FB1C076AFAA3" box="[454,540,1267,1292]" pageId="1" pageNumber="444" value="62-10-02">100262</date>
</specimenCode>
), same data as for
<typeStatus id="2E61E8E8E231FFC9708AFB1C0627FAA3" box="[747,849,1267,1292]" pageId="1" pageNumber="444" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B9C005C1E231FFC972A6FAF90603F984" pageId="1" pageNumber="444" type="diagnosis">
<paragraph id="F165564AE231FFC972A6FAF90603F984" blockId="1.[151,1437,1192,2012]" pageId="1" pageNumber="444">
<emphasis id="C3AE8A58E231FFC972A6FAF90437FA80" bold="true" box="[199,321,1302,1327]" pageId="1" pageNumber="444">Diagnosis.</emphasis>
<taxonomicName id="36DA2DC9E231FFC97329FAF804FCFA80" authority="Sankaran &amp; Sebastian, 2016" authorityName="Sankaran &amp; Sebastian" authorityYear="2016" box="[328,394,1302,1327]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="alii" status="sp. nov.">
<emphasis id="C3AE8A58E231FFC97329FAF804FCFA80" box="[328,394,1302,1327]" italics="true" pageId="1" pageNumber="444">S. alii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D89D3723E231FFC973F0FAF9049DFA80" box="[401,491,1302,1327]" pageId="1" pageNumber="444" rank="species">
<emphasis id="C3AE8A58E231FFC973F0FAF9049DFA80" bold="true" box="[401,491,1302,1327]" pageId="1" pageNumber="444">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="36DA2DC9E231FFC970DCFAF80659FA80" authorityName="Lehtinen" authorityYear="1981" box="[701,815,1302,1327]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC970DCFAF80659FA80" box="[701,815,1302,1327]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
but can be distinguished by the following combination of characters:
<emphasis id="C3AE8A58E231FFC97356FAD4040DFAFB" box="[311,379,1339,1364]" italics="true" pageId="1" pageNumber="444">Males</emphasis>
: Eye tubercle with pointed apex (eye tubercle in
<taxonomicName id="36DA2DC9E231FFC971C8FAD3016DFAFB" authorityName="Lehtinen" authorityYear="1981" box="[937,1051,1339,1364]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC971C8FAD3016DFAFB" box="[937,1051,1339,1364]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
with blunt apex), triangular PLEs (
<taxonomicName id="36DA2DC9E231FFC972C1FAB00467FAD8" authorityName="Lehtinen" authorityYear="1981" box="[160,273,1374,1399]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC972C1FAB00467FAD8" box="[160,273,1374,1399]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
with circular PLEs), bulb with apple-like shape (in
<taxonomicName id="36DA2DC9E231FFC97136FAB006BEFAD8" authorityName="Lehtinen" authorityYear="1981" box="[855,968,1374,1399]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC97136FAB006BEFAD8" box="[855,968,1374,1399]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
, bulb piriform), very long embolus (short embolus in
<taxonomicName id="36DA2DC9E231FFC97342FA6B04EEFA33" authorityName="Lehtinen" authorityYear="1981" box="[291,408,1411,1436]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC97342FA6B04EEFA33" box="[291,408,1411,1436]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
) and embolus with disto-prolateral origin (in
<taxonomicName id="36DA2DC9E231FFC971D8FA6B0159FA33" authorityName="Lehtinen" authorityYear="1981" box="[953,1071,1411,1436]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC971D8FA6B0159FA33" box="[953,1071,1411,1436]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
, embolus originating apically).
<emphasis id="C3AE8A58E231FFC972F6FA490580FA10" box="[151,246,1446,1471]" italics="true" pageId="1" pageNumber="444">Females</emphasis>
: 2 pairs of perigenital plates (1 pair in
<taxonomicName id="36DA2DC9E231FFC970DBFA48065BFA10" authorityName="Lehtinen" authorityYear="1981" box="[698,813,1446,1471]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC970DBFA48065BFA10" box="[698,813,1446,1471]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
), preanal plate with single row of pits and setae (in
<taxonomicName id="36DA2DC9E231FFC977E6FA48059AFA4B" authorityName="Lehtinen" authorityYear="1981" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC977E6FA48059AFA4B" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
, preanal plate bearing numerous pits and setae in several irregular rows) and genital orifice situated in middle of epigastric fold (genital orifice in
<taxonomicName id="36DA2DC9E231FFC970E0FA000785F9A8" authorityName="Lehtinen" authorityYear="1981" box="[641,755,1518,1543]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="lilawati">
<emphasis id="C3AE8A58E231FFC970E0FA000785F9A8" box="[641,755,1518,1543]" italics="true" pageId="1" pageNumber="444">S. lilawati</emphasis>
</taxonomicName>
at posterior margin of epigastric fold) (compare figures 2A, C, 3A–D with
<bibRefCitation id="954B2BBBE231FFC9735DF9FC0494F984" author="Lehtinen" box="[316,482,1555,1580]" pageId="1" pageNumber="444" refString="Lehtinen, P. T. (1981) Spiders of the Oriental-Australian region. III. Tetrablemmidae, with a world revision. Acta Zoologica Fennica, 162, 1 - 151." type="journal article" year="1981">Lehtinen 1981</bibRefCitation>
: figs 198A, 202, 203, 208 and 214).
</paragraph>
</subSubSection>
<subSubSection id="B9C005C1E231FFCA72A6F9D901E1F8C0" lastPageId="2" lastPageNumber="445" pageId="1" pageNumber="444" type="description">
<paragraph id="F165564AE231FFCA72A6F9D9018CFE60" blockId="1.[151,1437,1192,2012]" lastBlockId="2.[151,1437,151,464]" lastPageId="2" lastPageNumber="445" pageId="1" pageNumber="444">
<emphasis id="C3AE8A58E231FFC972A6F9D9042FF9E0" bold="true" box="[199,345,1590,1615]" pageId="1" pageNumber="444">Description.</emphasis>
Male (
<typeStatus id="2E61E8E8E231FFC973D0F9D90761F9E0" box="[433,535,1590,1615]" pageId="1" pageNumber="444" type="holotype">holotype</typeStatus>
, figs 1A, 2A, C, 2E–F). Colouration: body brownish yellow; legs and palps yellowish brown. Prosoma finely retiform, with fine brownish streaks; cephalic region excluding the eye tubercle flat, thoracic margin rugose. Fovea indistinct. Ocular area black; six eyes in a compact group, ALE largest, PME smallest, raised on short, conical tubercle. Clypeus high, finely reticulated, slightly sloping forward. Chelicerae frontally with basally wide, pointed, upward-curving apophysis/horn (
<figureCitation id="69E14ACFE231FFC971DFF929016FF970" box="[958,1049,1734,1759]" captionStart="FIGURE 2" captionStartId="3.[151,250,1029,1051]" captionTargetBox="[158,1428,198,1008]" captionTargetId="figure@3.[150,1436,193,1008]" captionTargetPageId="3" captionText="FIGURE 2. Shearella alii sp. nov. A Male holotype habitus, retrolateral view; B Female paratype, habitus, retrolateral view; C Male, frontal view; D Female, frontal view; E Male, left chelicera, retrolateral view; F Male, left leg I, retrolateral view. Scale bars: A – B 0.15 mm; C, 0.08 mm; D, 0.06 mm; E, 0.02 mm; F, 0.40 mm." httpUri="https://zenodo.org/record/1052251/files/figure.png" pageId="1" pageNumber="444">Fig. 2E</figureCitation>
). Sternum finely reticulated, with sparse setae, fused at margin with prosoma. Opisthosoma oval; dorsal scutum large, oval, glossy, densely pitted, covered with fine setae; lateral scuta glossy, with paired rows of pits accommodating backward-directed setae; lateral plate/sclerite IV long, narrower than plates/sclerites II and III; pulmonary plate glossy, with pits for setae; postgenital plate slightly overlapping anterior margin of preanal plate, narrower than preanal plate, laterally tapering, with median row of pits; preanal plate slightly longer and much wider than postgenital plate, with numerous pits; 2 pairs of perigenital plates, 1 pair on each side. Leg segments smooth, hirsute; leg I not modified; femora I–IV ventrally with a longitudinal row of granules; tibiae I–IV with 3 dorsal trichobothria each; metatarsi I– IV with 1 dorsal trichobothrium each. Body length 1.23. Prosoma length 0.45, width (in middle) 0.42, height (in middle) 0.25. Opisthosoma length 0.78, width (in middle) 0.66, height (in middle) 0.61. Eye diameters: ALE 0.06. PME 0.01. PLE 0.04. Distance between PLEs 0.04. Clypeus height at ALE 0.21. Length of chelicerae 0.19. Measurements of palp and legs: palp 1.07 [0.22, 0.12, 0.16, 0.57], I 1.36 [0.44, 0.13, 0.32, 0.24, 0.23], II 1.21 [0.38, 0.12, 0.28, 0.21, 0.22], III 1.10 [0.32, 0.12, 0.24, 0.21, 0.21], IV 1.42 [0.43, 0.13, 0.36, 0.27, 0.23]. Leg formula (from longest to shortest leg): 4123.
<emphasis id="C3AE8A58E232FFCA7057FEA307EDFECA" box="[566,667,332,357]" italics="true" pageId="2" pageNumber="445">Pedipalp</emphasis>
(
<figureCitation id="69E14ACFE232FFCA70CAFEA3065BFECB" box="[683,813,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1694,1716]" captionTargetBox="[334,1252,201,1670]" captionTargetId="figure@4.[327,1259,195,1672]" captionTargetPageId="4" captionText="FIGURE 3. Shearella alii sp. nov. A Male, left pedipalp, prolateral view; B Same, retrolateral view; C Female, opisthosoma, ventral view; D Female, genital area, dorsal view. Abbreviations: B bulb, CP central process, E embolus, EF epigastric fold, GO genital orifice, IVP inner vulval plate, PA preanal plate, PEG perigenital plate, POG postgenital plate, S spermophor, SR seminal receptacle, VS vulval stem. Scale bars: A – B 0.16 mm; C, 0.10 mm; D, 0.01 mm." httpUri="https://zenodo.org/record/1052253/files/figure.png" pageId="2" pageNumber="445">Figs 3A–B</figureCitation>
): Femur ventrally with a row of granules, without long setae (present in
<taxonomicName id="36DA2DC9E232FFCA7332FE9F07F1FE27" authority="Lehtinen, 1981" authorityName="Lehtinen" authorityYear="1981" box="[339,647,367,392]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="445" phylum="Arthropoda" rank="species" species="selvarani">
<emphasis id="C3AE8A58E232FFCA7332FE9F04A0FE27" box="[339,470,367,392]" italics="true" pageId="2" pageNumber="445">S. selvarani</emphasis>
<bibRefCitation id="954B2BBBE232FFCA73BCFE8007F1FE27" author="Lehtinen" box="[477,647,367,392]" pageId="2" pageNumber="445" refString="Lehtinen, P. T. (1981) Spiders of the Oriental-Australian region. III. Tetrablemmidae, with a world revision. Acta Zoologica Fennica, 162, 1 - 151." type="journal article" year="1981">Lehtinen, 1981</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="36DA2DC9E232FFCA70DDFE9F066FFE27" baseAuthorityName="Lin &amp; Li" baseAuthorityYear="2010" box="[700,793,368,392]" class="Arachnida" family="Tetrablemmidae" genus="Sinamma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="445" phylum="Arthropoda" rank="species" species="sanya">
<emphasis id="C3AE8A58E232FFCA70DDFE9F066FFE27" box="[700,793,368,392]" italics="true" pageId="2" pageNumber="445">S. sanya</emphasis>
</taxonomicName>
); tibia with 1 dorsal trichobothrium. Bulb broad, smooth, apple-like in appearance. Embolus very long (0.31), thin, originating disto-prolaterally to bulb, with broad base, curving inward, with wedge-shaped tip (
<figureCitation id="69E14ACFE232FFCA703AFE5807D8FE7F" box="[603,686,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1694,1716]" captionTargetBox="[334,1252,201,1670]" captionTargetId="figure@4.[327,1259,195,1672]" captionTargetPageId="4" captionText="FIGURE 3. Shearella alii sp. nov. A Male, left pedipalp, prolateral view; B Same, retrolateral view; C Female, opisthosoma, ventral view; D Female, genital area, dorsal view. Abbreviations: B bulb, CP central process, E embolus, EF epigastric fold, GO genital orifice, IVP inner vulval plate, PA preanal plate, PEG perigenital plate, POG postgenital plate, S spermophor, SR seminal receptacle, VS vulval stem. Scale bars: A – B 0.16 mm; C, 0.10 mm; D, 0.01 mm." httpUri="https://zenodo.org/record/1052253/files/figure.png" pageId="2" pageNumber="445">Fig 3A</figureCitation>
, see arrow). Spermophor long with broad basal part.
</paragraph>
<caption id="A5A506C2E232FFCA72F6FA860038FAD0" box="[151,1358,1385,1407]" httpUri="https://zenodo.org/record/1052249/files/figure.png" pageId="2" pageNumber="445" targetBox="[160,1427,527,1354]" targetPageId="2">
<paragraph id="F165564AE232FFCA72F6FA860038FAD0" blockId="2.[151,1358,1385,1407]" box="[151,1358,1385,1407]" pageId="2" pageNumber="445">
<emphasis id="C3AE8A58E232FFCA72F6FA860465FAD1" bold="true" box="[151,275,1385,1407]" pageId="2" pageNumber="445">FIGURE 1.</emphasis>
<taxonomicName id="36DA2DC9E232FFCA7378FA8604D7FAD0" authority="Sankaran &amp; Sebastian, 2016" authorityName="Sankaran &amp; Sebastian" authorityYear="2016" box="[281,417,1385,1407]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="445" phylum="Arthropoda" rank="species" species="alii" status="sp. nov.">
<emphasis id="C3AE8A58E232FFCA7378FA8604D7FAD0" box="[281,417,1385,1407]" italics="true" pageId="2" pageNumber="445">Shearella alii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D89D3723E232FFCA73C6FA86048EFAD1" box="[423,504,1385,1406]" pageId="2" pageNumber="445" rank="species">
<emphasis id="C3AE8A58E232FFCA73C6FA86048EFAD1" bold="true" box="[423,504,1385,1406]" pageId="2" pageNumber="445">sp. nov.</emphasis>
</taxonomicNameLabel>
A Male holotype, dorsal view; B Female paratype, dorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="F165564AE232FFCA72A6FA490124F8E4" blockId="2.[151,1437,1446,2011]" pageId="2" pageNumber="445">
Female (
<typeStatus id="2E61E8E8E232FFCA734BFA4804F9FA10" box="[298,399,1447,1471]" pageId="2" pageNumber="445" type="paratype">paratype</typeStatus>
, figs 1B, 2B, D). In all details like male except for the following: Colouration: body dark brown. Eye tubercle low and domed. Chelicera without frontal apophysis/horn but with weak frontal protrusions (
<figureCitation id="69E14ACFE232FFCA72FEFA01058BF9A8" box="[159,253,1518,1543]" captionStart="FIGURE 2" captionStartId="3.[151,250,1029,1051]" captionTargetBox="[158,1428,198,1008]" captionTargetId="figure@3.[150,1436,193,1008]" captionTargetPageId="3" captionText="FIGURE 2. Shearella alii sp. nov. A Male holotype habitus, retrolateral view; B Female paratype, habitus, retrolateral view; C Male, frontal view; D Female, frontal view; E Male, left chelicera, retrolateral view; F Male, left leg I, retrolateral view. Scale bars: A – B 0.15 mm; C, 0.08 mm; D, 0.06 mm; E, 0.02 mm; F, 0.40 mm." httpUri="https://zenodo.org/record/1052251/files/figure.png" pageId="2" pageNumber="445">Fig. 2D</figureCitation>
, see arrow). Lateral plate/sclerite IV nearly as wide as plates/sclerites II and III. Preanal plate with proximal row of pits. Body length 1.28. Prosoma length 0.49, width (in middle) 0.47, height (in middle) 0.29. Opisthosoma length 0.79, width (in middle) 0.69, height (in middle) 0.62. Eye diameters: ALE 0.04. PME 0.01. PLE 0.03. Distance between PLEs 0.05. Clypeus height at ALE 0.18. Length of chelicerae 0.19. Measurements of palp and legs: palp 0.24 [0.10, 0.03, 0.04, 0.07], I 1.24 [0.41, 0.12, 0.28, 0.20, 0.23], II 1.12 [0.34, 0.12, 0.25, 0.20, 0.21], III 1.02 [0.31, 0.12, 0.21, 0.19, 0.19], IV 1.34 [0.41, 0.12, 0.33, 0.26, 0.22].
<emphasis id="C3AE8A58E232FFCA76EAF94D0183F914" box="[1163,1269,1698,1723]" italics="true" pageId="2" pageNumber="445">Genitalia</emphasis>
(
<figureCitation id="69E14ACFE232FFCA7768F94D00E6F914" box="[1289,1424,1698,1723]" captionStart="FIGURE 3" captionStartId="4.[151,250,1694,1716]" captionTargetBox="[334,1252,201,1670]" captionTargetId="figure@4.[327,1259,195,1672]" captionTargetPageId="4" captionText="FIGURE 3. Shearella alii sp. nov. A Male, left pedipalp, prolateral view; B Same, retrolateral view; C Female, opisthosoma, ventral view; D Female, genital area, dorsal view. Abbreviations: B bulb, CP central process, E embolus, EF epigastric fold, GO genital orifice, IVP inner vulval plate, PA preanal plate, PEG perigenital plate, POG postgenital plate, S spermophor, SR seminal receptacle, VS vulval stem. Scale bars: A – B 0.16 mm; C, 0.10 mm; D, 0.01 mm." httpUri="https://zenodo.org/record/1052253/files/figure.png" pageId="2" pageNumber="445">Figs 3C–D</figureCitation>
): Epigastric fold distinct, strongly sclerotized, laterally with upward curving ends touching seminal receptacula. Genital orifice transversely oval, less obvious than surrounding dark region. Central genital process short, conical. Vulval stem broad, nearly triangular. Inner vulval plate finger-like, with blunt apex. Vulval ducts membranous originating antero-laterally from vulval stem, leading to sac-like seminal receptacula.
</paragraph>
<paragraph id="F165564AE232FFCA72A6F8B901E1F8C0" blockId="2.[151,1437,1446,2011]" box="[199,1175,1878,1903]" pageId="2" pageNumber="445">
<emphasis id="C3AE8A58E232FFCA72A6F8B9044AF8C0" bold="true" box="[199,316,1878,1903]" pageId="2" pageNumber="445">Variation.</emphasis>
Male (n = 3): Body length 1.15–1.23. Female (n=3): Body length 1.22–1.28
</paragraph>
</subSubSection>
<subSubSection id="B9C005C1E232FFCA72A6F89501ADF874" pageId="2" pageNumber="445" type="etymology">
<paragraph id="F165564AE232FFCA72A6F89507B9F818" blockId="2.[151,1437,1446,2011]" pageId="2" pageNumber="445">
<emphasis id="C3AE8A58E232FFCA72A6F8950438F83C" bold="true" box="[199,334,1914,1939]" pageId="2" pageNumber="445">Etymology.</emphasis>
The specific epithet is a patronym honouring Dr. Salim Ali, the Birdman of
<collectingCountry id="89CD16DAE232FFCA76B8F8950063F83C" box="[1241,1301,1914,1939]" name="India" pageId="2" pageNumber="445">India</collectingCountry>
, one of the greatest ornithologists and naturalists in the world.
</paragraph>
<paragraph id="F165564AE232FFCA72A6F82D01ADF874" blockId="2.[151,1437,1446,2011]" box="[199,1243,1986,2011]" pageId="2" pageNumber="445">
<emphasis id="C3AE8A58E232FFCA72A6F82D04F3F874" bold="true" box="[199,389,1986,2011]" pageId="2" pageNumber="445">Natural history.</emphasis>
<taxonomicName id="36DA2DC9E232FFCA73EDF82D0754F874" authority="Sankaran &amp; Sebastian, 2016" authorityName="Sankaran &amp; Sebastian" authorityYear="2016" box="[396,546,1986,2011]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="445" phylum="Arthropoda" rank="species" species="alii" status="sp. nov.">
<emphasis id="C3AE8A58E232FFCA73EDF82D0754F874" box="[396,546,1986,2011]" italics="true" pageId="2" pageNumber="445">Shearella alii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D89D3723E232FFCA7048F82D07F4F874" box="[553,642,1986,2011]" pageId="2" pageNumber="445" rank="species">
<emphasis id="C3AE8A58E232FFCA7048F82D07F4F874" bold="true" box="[553,642,1986,2011]" pageId="2" pageNumber="445">sp. nov.</emphasis>
</taxonomicNameLabel>
inhabits the leaf litter layer in a dry deciduous forest.
</paragraph>
</subSubSection>
<caption id="A5A506C2E233FFCB72F6FBEA0602FBF6" httpUri="https://zenodo.org/record/1052251/files/figure.png" pageId="3" pageNumber="446" targetBox="[158,1428,198,1008]" targetPageId="3">
<paragraph id="F165564AE233FFCB72F6FBEA0602FBF6" blockId="3.[151,1436,1029,1113]" pageId="3" pageNumber="446">
<emphasis id="C3AE8A58E233FFCB72F6FBEA0464FBB4" bold="true" box="[151,274,1029,1051]" pageId="3" pageNumber="446">FIGURE 2.</emphasis>
<taxonomicName id="36DA2DC9E233FFCB7379FBEA04D6FBB4" authority="Sankaran &amp; Sebastian, 2016" authorityName="Sankaran &amp; Sebastian" authorityYear="2016" box="[280,416,1029,1051]" class="Arachnida" family="Tetrablemmidae" genus="Shearella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="446" phylum="Arthropoda" rank="species" species="alii" status="sp. nov.">
<emphasis id="C3AE8A58E233FFCB7379FBEA04D6FBB4" box="[280,416,1029,1051]" italics="true" pageId="3" pageNumber="446">Shearella alii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D89D3723E233FFCB73C7FBE90481FBB4" box="[422,503,1030,1051]" pageId="3" pageNumber="446" rank="species">
<emphasis id="C3AE8A58E233FFCB73C7FBE90481FBB4" bold="true" box="[422,503,1030,1051]" pageId="3" pageNumber="446">sp. nov.</emphasis>
</taxonomicNameLabel>
A Male holotype habitus, retrolateral view; B Female paratype, habitus, retrolateral view; C Male, frontal view; D Female, frontal view; E Male, left chelicera, retrolateral view; F Male, left leg I, retrolateral view. Scale bars: A–B 0.15 mm; C, 0.08 mm; D, 0.06 mm; E, 0.02 mm; F, 0.40 mm.
</paragraph>
</caption>
<subSubSection id="B9C005C1E233FFCB72A6FB6F07B8FB12" pageId="3" pageNumber="446" type="distribution">
<paragraph id="F165564AE233FFCB72A6FB6F07B8FB12" blockId="3.[199,962,1152,1213]" pageId="3" pageNumber="446">
<emphasis id="C3AE8A58E233FFCB72A6FB6F0416FB36" bold="true" box="[199,352,1152,1177]" pageId="3" pageNumber="446">Distribution.</emphasis>
At present known only from the
<typeStatus id="2E61E8E8E233FFCB70B4FB6E0673FB36" box="[725,773,1153,1177]" pageId="3" pageNumber="446">type</typeStatus>
locality (
<figureCitation id="69E14ACFE233FFCB710EFB6F06C2FB36" box="[879,948,1152,1177]" captionStart="FIGURE 4" captionStartId="5.[151,250,1373,1395]" captionTargetBox="[183,1404,577,1330]" captionTargetId="figure@5.[159,1423,558,1349]" captionTargetPageId="5" captionText="FIGURE 4. Map showing the currently known localities of Shearella spp. ● Shearella alii sp. nov. Shearella lilawati Lehtinen, 1981 ▲ Shearella sanya Lin &amp; Li, 2010 ♦ Shearella selvarani Lehtinen, 1981." httpUri="https://zenodo.org/record/1052255/files/figure.png" pageId="3" pageNumber="446">Fig. 4</figureCitation>
).
<emphasis id="C3AE8A58E233FFCB72A6FB4B0446FB12" bold="true" box="[199,304,1188,1213]" pageId="3" pageNumber="446">Remark.</emphasis>
First record of the genus from
<collectingCountry id="89CD16DAE233FFCB70ECFB4B07BCFB12" box="[653,714,1188,1213]" name="India" pageId="3" pageNumber="446">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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