114 lines
8.6 KiB
XML
114 lines
8.6 KiB
XML
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<document ID-DOI="http://doi.org/10.5281/zenodo.3679540" ID-GBIF-Dataset="75f859ce-4da6-4513-871e-e1fb3b41a55f" ID-Zenodo-Dep="3679540" checkinTime="1581972654346" checkinUser="plazi" docAuthor="Voris, Jared T., Therrien, François, Zelenitsky, Darla K. & Brown, Caleb M." docDate="2020" docId="039E87F3FFE9FFDEB9CBDB5A6C56FF75" docLanguage="en" docName="createceousResearch.110.104.388.pdf" docOrigin="Cretaceous Research 110 (104388)" docStyle="DocumentStyle{}" docTitle="Thanatotheristes Voris & Therrien & Zelenitsky & Brown 2020" docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="FFA7FF8BFFE5FFD2B95ADE6C695CFF97" masterDocTitle="A new tyrannosaurine (Theropoda: Tyrannosauridae) from the Campanian Foremost Formation of Alberta, Canada, provides insight into the evolution and biogeography of tyrannosaurids" masterLastPageNumber="15" masterPageNumber="1" pageId="12" pageNumber="13" updateTime="1643450941510" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new tyrannosaurine (Theropoda: Tyrannosauridae) from the Campanian Foremost Formation of Alberta, Canada, provides insight into the evolution and biogeography of tyrannosaurids</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Voris, Jared T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Therrien, François</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zelenitsky, Darla K.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Brown, Caleb M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Cretaceous Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-01-23</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>110</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>104388</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>15</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1016/j.cretres.2020.104388</mods:identifier>
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<mods:identifier type="GBIF-Dataset">75f859ce-4da6-4513-871e-e1fb3b41a55f</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3672003</mods:identifier>
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<mods:identifier type="ZooBank">26778F25-3135-412D-A203-752CB9C078F1</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3679540" ID-Zenodo-Dep="3679540" LSID="urn:lsid:plazi:treatment:039E87F3FFE9FFDEB9CBDB5A6C56FF75" httpUri="http://treatment.plazi.org/id/039E87F3FFE9FFDEB9CBDB5A6C56FF75" lastPageNumber="13" pageId="12" pageNumber="13">
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<subSubSection pageId="12" pageNumber="13" type="discussion">
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<paragraph blockId="12.[113,784,1334,1968]" pageId="12" pageNumber="13">
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<emphasis box="[145,443,1334,1354]" italics="true" pageId="12" pageNumber="13">
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<taxonomicName authority="Voris & Therrien & Zelenitsky & Brown, 2020" authorityName="Voris & Therrien & Zelenitsky & Brown" authorityYear="2020" box="[145,306,1334,1354]" class="Reptilia" family="Tyrannosauridae" genus="Thanatotheristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="12" pageNumber="3" phylum="Chordata" rank="genus" status="gen. nov.">Thanatotheristes</taxonomicName>
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degrootorum
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</emphasis>
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is a new genus and species of derived tyrannosaurine from the middle Campanian of
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<collectingRegion box="[704,779,1363,1382]" country="Canada" name="Alberta" pageId="12" pageNumber="13">Alberta</collectingRegion>
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,
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<collectingCountry box="[113,189,1391,1410]" name="Canada" pageId="12" pageNumber="13">Canada</collectingCountry>
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. Whereas this taxon is most closely related to
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<taxonomicName authority=", Thanatotheristes" authorityName="Thanatotheristes" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<taxonomicName genus="Daspletosaurus" pageId="12" pageNumber="13" rank="genus">
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<emphasis italics="true" pageId="12" pageNumber="13">Daspletosaurus</emphasis>
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</taxonomicName>
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,
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<emphasis box="[199,360,1417,1437]" italics="true" pageId="12" pageNumber="13">Thanatotheristes</emphasis>
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</taxonomicName>
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is morphologically distinct based on several characters and autapomorphies. Together,
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<taxonomicName authority="Voris & Therrien & Zelenitsky & Brown, 2020" authorityName="Voris & Therrien & Zelenitsky & Brown" authorityYear="2020" box="[622,783,1445,1465]" class="Reptilia" family="Tyrannosauridae" genus="Thanatotheristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="12" pageNumber="3" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis box="[622,783,1445,1465]" italics="true" pageId="12" pageNumber="13">Thanatotheristes</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[158,305,1473,1493]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis box="[158,305,1473,1493]" italics="true" pageId="12" pageNumber="13">Daspletosaurus</emphasis>
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</taxonomicName>
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form a clade of long- and deep-snouted tyrannosaurines, here referred to as Daspletosaurini. The recovery of additional clades in our phylogenetic analysis, including Albertosaurinae, Alioramini, short-snouted taxa like
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<emphasis box="[589,730,1557,1577]" italics="true" pageId="12" pageNumber="13">Teratophoneus</emphasis>
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, and gigantic taxa like
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<emphasis box="[302,451,1585,1605]" italics="true" pageId="12" pageNumber="13">
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<taxonomicName authorityName="Osborn" authorityYear="1905" box="[302,447,1585,1605]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">Tyrannosaurus</taxonomicName>
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,
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</emphasis>
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reveals that at least five major radiations occurred within
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<taxonomicName authorityName="Osborn" authorityYear="1906" box="[394,563,1614,1633]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="12" pageNumber="13" phylum="Chordata" rank="family">Tyrannosauridae</taxonomicName>
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and suggests greater disparity within the group than previously thought. Paleogeographic segregation of tyrannosaurid clades across North America during the Campanian provides renewed evidence for provinciality among Laramidian tyrannosaurids, with
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<emphasis box="[564,705,1724,1744]" italics="true" pageId="12" pageNumber="13">Teratophoneus</emphasis>
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/
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<taxonomicName class="Reptilia" family="Tyrannosauridae" genus="Lythronax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="12" pageNumber="13">Lythronax</emphasis>
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</taxonomicName>
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-
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<emphasis box="[165,301,1752,1772]" italics="true" pageId="12" pageNumber="13">Bistahieversor</emphasis>
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being endemic to southern Laramidia and Albertosaurinae-Daspletosaurini being endemic to northern Laramidia. Despite the co-occurrence of Daspletosaurini and Albertosaurinae in northern Laramidia, the former is more common in southernmost
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<collectingRegion box="[259,332,1865,1884]" country="Canada" name="Alberta" pageId="12" pageNumber="13">Alberta</collectingRegion>
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and Montana and the latter is more common farther north, in central and southcentral
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<collectingRegion box="[547,622,1893,1912]" country="Canada" name="Alberta" pageId="12" pageNumber="13">Alberta</collectingRegion>
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, suggesting the presence of a latitudinal gradient. Thus, the absence of Daspletosaurini in the lower Maastrichtian deposits from south-central
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</paragraph>
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<paragraph blockId="12.[831,1500,179,226]" pageId="12" pageNumber="13">
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<collectingRegion box="[831,904,179,198]" country="Canada" name="Alberta" pageId="12" pageNumber="13">Alberta</collectingRegion>
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(i.e. Horseshoe Canyon Formation) could reflect this latitudinal gradient rather than their extinction.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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