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<document id="CE37B5C867FC3D0F86437A207A31C298" ID-DOI="10.5281/zenodo.5301245" ID-ISSN="0374-1036" ID-Zenodo-Dep="5301245" ID-ZooBank="D86FB04C-BD99-44FC-A724-B9DA84D079FEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630100391386" checkinUser="marcus" docAuthor="Sehnal, Richard" docDate="2014" docId="420387B18041D344FE4EFC3AFF1CFE09" docLanguage="en" docName="ActaEntMusNatPra.2014.54.2.563-569.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Engistanoxia sagala Sehnal 2014, sp. nov." docType="treatment" docVersion="4" lastPageNumber="568" masterDocId="BE3AFFC98040D341FFF5FFE4FFD7FFFA" masterDocTitle="Engistanoxia sagala sp. nov. from Kenya (Coleoptera: Scarabaeidae: Melolonthinae: Melolonthini)" masterLastPageNumber="569" masterPageNumber="563" pageNumber="564" updateTime="1699152508716" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="61D7BB1BDAD909B5FA7AA24D94ADCAB7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="43D14B2DDEA3A555A6DCE620AD075417">
<mods:title id="71FB1031EBB78C8ABEB3A5543242AA92">Engistanoxia sagala sp. nov. from Kenya (Coleoptera: Scarabaeidae: Melolonthinae: Melolonthini)</mods:title>
</mods:titleInfo>
<mods:name id="0FD2E9DA78AFE13F5CBB92E12AE426B9" type="personal">
<mods:role id="D8C01BF4BB849BA592D44345A1D0BCC9">
<mods:roleTerm id="89AB24B932C6387235ED905688ED03BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE7C18FB49B9072AE59B4EA47826F26A">Sehnal, Richard</mods:namePart>
</mods:name>
<mods:typeOfResource id="CFC762492B7F038A91F772EC446E9D06">text</mods:typeOfResource>
<mods:relatedItem id="F3BD41A7829BB56CADB2702167CA9126" type="host">
<mods:titleInfo id="4514878E736DEAB14D964904989E0B58">
<mods:title id="D98C883C998A3F06B1C38A62BBA72503">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="C13F719830F74D8C18702553917945D2">
<mods:date id="1EE3CE23DDAF18CA4504EB351D539823">2014</mods:date>
<mods:detail id="88A62EBEB50DAD5FD9B93F23C8AEFC22" type="pubDate">
<mods:number id="4858AB4D45F044667D396B28B3E2F987">2014-12-15</mods:number>
</mods:detail>
<mods:detail id="60FB286C5DAD327BC8CDE97C79A37C27" type="volume">
<mods:number id="36FA16BE3141C0D52724DC7CE85ABBD1">54</mods:number>
</mods:detail>
<mods:detail id="579BAEA06338A5EE1EBBE22FD9878C19" type="issue">
<mods:number id="23668134184A3393FE6F6A885CAB7516">2</mods:number>
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<mods:start id="C20932FB2D58838EEE06CA75E24868AA">563</mods:start>
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<mods:classification id="459EFDE127AE5EE75DE55E697D4A63F5">journal article</mods:classification>
<mods:identifier id="92F971E47C9FD2E19D21CB1809798ED0" type="DOI">10.5281/zenodo.5301245</mods:identifier>
<mods:identifier id="34DC944BB71FE46DD35FAC09C131B74A" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="0362CF292F1F2F0BCAE71B5E701FC841" type="Zenodo-Dep">5301245</mods:identifier>
<mods:identifier id="485516755D02E9E068D2C76C143053C2" type="ZooBank">D86FB04C-BD99-44FC-A724-B9DA84D079FEC</mods:identifier>
</mods:mods>
<treatment id="420387B18041D344FE4EFC3AFF1CFE09" ID-DOI="http://doi.org/10.5281/zenodo.5330556" ID-Zenodo-Dep="5330556" LSID="urn:lsid:plazi:treatment:420387B18041D344FE4EFC3AFF1CFE09" httpUri="http://treatment.plazi.org/id/420387B18041D344FE4EFC3AFF1CFE09" lastPageId="5" lastPageNumber="568" pageId="1" pageNumber="564">
<subSubSection id="82B0652C8041D340FE4EFC3AFD20FC0C" box="[443,759,990,1014]" pageId="1" pageNumber="564" type="nomenclature">
<paragraph id="CA1536A78041D340FE4EFC3AFD20FC0C" blockId="1.[443,759,990,1014]" box="[443,759,990,1014]" pageId="1" pageNumber="564">
<heading id="915D81CB8041D340FE4EFC3AFD20FC0C" bold="true" box="[443,759,990,1014]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="564" reason="3">
<emphasis id="F8DEEAB58041D340FE4EFC3AFD20FC0C" bold="true" box="[443,759,990,1014]" pageId="1" pageNumber="564">
<taxonomicName id="0DAA4D248041D340FE4EFC3AFD4CFC0C" authority="Sehnal, 2014" authorityName="Sehnal" authorityYear="2014" box="[443,667,990,1014]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="sagala" status="sp. nov.">
<emphasis id="F8DEEAB58041D340FE4EFC3AFD4CFC0C" bold="true" box="[443,667,990,1014]" italics="true" pageId="1" pageNumber="564">Engistanoxia sagala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3ED57CE8041D340FD57FC3BFD20FC0C" box="[674,759,991,1014]" pageId="1" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="82B0652C8041D340FDD8FC19FD52FBEA" box="[557,645,1021,1040]" pageId="1" pageNumber="564" type="description">
<paragraph id="CA1536A78041D340FDD8FC19FD52FBEA" blockId="1.[557,645,1021,1040]" box="[557,645,1021,1040]" pageId="1" pageNumber="564">
(
<figureCitation id="52912A228041D340FDC6FC19FDA9FBEA" box="[563,638,1021,1040]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="1" pageNumber="564">Figs 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="82B0652C8041D340FF84FBC3FEFAFB20" pageId="1" pageNumber="564" type="materials_examined">
<paragraph id="CA1536A78041D340FF84FBC3FD77FBC0" blockId="1.[113,1089,1063,1242]" box="[113,672,1063,1082]" pageId="1" pageNumber="564">
<emphasis id="F8DEEAB58041D340FF84FBC3FF3DFBC0" bold="true" box="[113,234,1063,1082]" pageId="1" pageNumber="564">
<typeStatus id="151188058041D340FF84FBC3FF49FBC0" box="[113,158,1063,1082]" pageId="1" pageNumber="564">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="B2BD76378041D340FF1AFBC3FEFEFBC0" box="[239,297,1063,1082]" name="Kenya" pageId="1" pageNumber="564">Kenya</collectingCountry>
,
<collectingRegion id="086EF8458041D340FEC4FBC3FE0EFBC0" box="[305,473,1063,1082]" country="Kenya" name="Taita/Taveta" pageId="1" pageNumber="564">Taita Taveta county</collectingRegion>
, Voi env.,
<taxonomicName id="0DAA4D248041D340FDC4FBC3FD77FBC0" authority="Hills." authorityName="Hills." box="[561,672,1063,1082]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="sagala">Sagala Hills.</taxonomicName>
</paragraph>
<paragraph id="CA1536A78041D340FF84FBA5FEFAFB20" blockId="1.[113,1089,1063,1242]" pageId="1" pageNumber="564">
<emphasis id="F8DEEAB58041D340FF84FBA5FF21FBAE" bold="true" box="[113,246,1089,1108]" pageId="1" pageNumber="564">Type material.</emphasis>
<materialsCitation id="7AC23CFA8041D340FF09FBA5FD86FB95" collectingDate="2000-11-23" collectingDateMax="2000-12-04" collectingDateMin="2000-11-23" collectionCode="NMPC" collectorName="M. BednaGik" country="Kenya" location="Coast prov." municipality="Voi" pageId="1" pageNumber="564" specimenCount="1" typeStatus="holotype">
<typeStatus id="151188058041D340FF09FBA5FE98FBA8" box="[252,335,1089,1108]" pageId="1" pageNumber="564" type="holotype">HOLOTYPE</typeStatus>
: J, ދ
<collectingCountry id="B2BD76378041D340FE88FBA5FE1FFBAE" box="[381,456,1089,1108]" name="Kenya" pageId="1" pageNumber="564">KENYA</collectingCountry>
,
<location id="CF75607C8041D340FE25FBA5FDE4FBAE" LSID="urn:lsid:plazi:treatment:420387B18041D344FE4EFC3AFF1CFE09:CF75607C8041D340FE25FBA5FDE4FBAE" box="[464,563,1089,1108]" country="Kenya" municipality="Voi" name="Coast prov." pageId="1" pageNumber="564">Coast prov.</location>
[the earlier division of the country] /
<collectingMunicipality id="2A71ACDD8041D340FC83FBA5FC44FBAE" box="[886,915,1089,1108]" pageId="1" pageNumber="564">Voi</collectingMunicipality>
env. &amp;
<taxonomicName id="0DAA4D248041D340FC2CFBA5FB96FBAE" authority="Hills" authorityName="Hills" box="[985,1089,1089,1108]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="sagala">Sagala Hills</taxonomicName>
/
<date id="BE1410678041D340FF88FBB8FEE1FB95" box="[125,310,1116,1135]" pageId="1" pageNumber="564" value="2000-11-23" valueMax="2000-12-04" valueMin="2000-11-23">
<collectingDate id="AE50E98F8041D340FF88FBB8FEE1FB95" box="[125,310,1116,1135]" pageId="1" pageNumber="564" value="2000-11-23" valueMax="2000-12-04" valueMin="2000-11-23">23. XI. - 4. XII. 2000</collectingDate>
</date>
/
<collectorName id="675F53718041D340FEB2FBB8FE63FB95" box="[327,436,1114,1136]" pageId="1" pageNumber="564">M. BednaĜík</collectorName>
leg. [p]ތ (
<collectionCode id="ACBBAE628041D340FDFFFBB8FD9CFB95" box="[522,587,1116,1135]" country="Czech Republic" name="National Museum Prague" pageId="1" pageNumber="564">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7AC23CFA8041D340FDAFFBB8FF29FB70" collectingDate="2000-11-23" collectingDateMax="2000-12-04" collectingDateMin="2000-11-23" collectionCode="NMPC" collectorName="Nos &amp; No." country="Kenya" location="Coast prov." municipality="Voi" pageId="1" pageNumber="564" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="151188058041D340FDAFFBB8FD66FB97" box="[602,689,1116,1135]" pageId="1" pageNumber="564" type="paratype">PARATYPES</typeStatus>
: No. 1,
<specimenCount id="DCACFD2E8041D340FD0CFBBCFCD0FB95" box="[761,775,1112,1135]" pageId="1" pageNumber="564" type="female">♀</specimenCount>
and
<collectorName id="675F53718041D340FCC7FBB8FC83FB95" box="[818,852,1116,1135]" pageId="1" pageNumber="564">Nos</collectorName>
2–6 and
<collectorName id="675F53718041D340FC50FBB8FC12FB95" box="[933,965,1116,1135]" pageId="1" pageNumber="564">No.</collectorName>
12, JJ, same data as holotype
</materialsCitation>
;
<materialsCitation id="7AC23CFA8041D340FEF2FB93FF0DFB5E" collectingDate="2000-11-23" collectingDateMax="2000-12-04" collectingDateMin="2000-11-23" country="Kenya" county="Voi" location="Nos" municipality="Coast prov." pageId="1" pageNumber="564" specimenCount="1" typeStatus="paratype">
<location id="CF75607C8041D340FEF2FB93FEFEFB70" LSID="urn:lsid:plazi:treatment:420387B18041D344FE4EFC3AFF1CFE09:CF75607C8041D340FEF2FB93FEFEFB70" box="[263,297,1143,1162]" country="Kenya" county="Voi" municipality="Coast prov." name="Nos" pageId="1" pageNumber="564">Nos</location>
7–11, JJ, ދ
<collectingCountry id="B2BD76378041D340FE61FB93FE08FB70" box="[404,479,1143,1162]" name="Kenya" pageId="1" pageNumber="564">KENYA</collectingCountry>
,
<collectingMunicipality id="2A71ACDD8041D340FE13FB93FD9EFB70" box="[486,585,1143,1162]" pageId="1" pageNumber="564">Coast prov.</collectingMunicipality>
/
<collectingCounty id="23744E2B8041D340FDA2FB93FDA3FB70" box="[599,628,1143,1162]" pageId="1" pageNumber="564">Voi</collectingCounty>
env. &amp;
<taxonomicName id="0DAA4D248041D340FD42FB93FCC9FB70" authority="Hills" authorityName="Hills" box="[695,798,1143,1162]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="sagala">Sagala Hills</taxonomicName>
/
<date id="BE1410678041D340FCDAFB93FC32FB70" box="[815,997,1143,1162]" pageId="1" pageNumber="564" value="2000-11-23" valueMax="2000-12-04" valueMin="2000-11-23">
<collectingDate id="AE50E98F8041D340FCDAFB93FC32FB70" box="[815,997,1143,1162]" pageId="1" pageNumber="564" value="2000-11-23" valueMax="2000-12-04" valueMin="2000-11-23">23. XI. - 4. XII. 2000</collectingDate>
</date>
/ Ivo MartinĤ leg. [p]ތ
</materialsCitation>
;
<materialsCitation id="7AC23CFA8041D340FF1CFB75FD59FB45" collectorName="Voi &amp; Sagala &amp; Werner &amp; Coll. M. Lacroix" country="Kenya" location="KENYA" pageId="1" pageNumber="564" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
Nos 13–14, JJand No. 15,
<specimenCount id="DCACFD2E8041D340FE14FB6AFE38FB5F" box="[481,495,1166,1189]" pageId="1" pageNumber="564" type="female">♀</specimenCount>
, ދ
<collectingCountry id="B2BD76378041D340FDF6FB75FD9DFB5E" box="[515,586,1169,1188]" name="Kenya" pageId="1" pageNumber="564">KENYA</collectingCountry>
XI-92 / near
<collectorName id="675F53718041D340FD4BFB75FD09FB5E" box="[702,734,1169,1188]" pageId="1" pageNumber="564">Voi</collectorName>
,
<collectorName id="675F53718041D340FD13FB75FCC8FB5E" box="[742,799,1169,1188]" pageId="1" pageNumber="564">Sagala</collectorName>
reg. /
<collectorName id="675F53718041D340FCA3FB75FC43FB5E" box="[854,916,1169,1188]" pageId="1" pageNumber="564">Werner</collectorName>
leg. //
<taxonomicName id="0DAA4D248041D340FC24FB75FB97FB5E" authorityName="Lacroix" authorityYear="2002" box="[977,1088,1169,1188]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus">Engistanoxia</taxonomicName>
[bold] / dessertineae / M.
<bibRefCitation id="AE3B4B568041D340FEA5FB48FE1EFB45" author="LACROIX M." box="[336,457,1196,1215]" pageId="1" pageNumber="564" pagination="153 - 168" refId="ref2934" refString="LACROIX M. 2002: Nouveaux Melolonthidae afrotropicaux (Coleoptera, Scarabaeoidea). Coleopteres 8 (9): 153 - 168. LACROIX M. 2010: Melolonthinae afrotropicaux. Genera et Catalogue. Collection Hannetons, Edition M. Lacroix, Paris, 277 pp." type="book chapter" year="2002">Lacroix, 2002</bibRefCitation>
/
<collectorName id="675F53718041D340FE2EFB48FDBCFB45" box="[475,619,1196,1215]" pageId="1" pageNumber="564">Coll. M. Lacroix</collectorName>
[p]ތ
</materialsCitation>
.
<typeStatus id="151188058041D340FD68FB48FD27FB45" box="[669,752,1196,1215]" pageId="1" pageNumber="564" type="paratype">Paratypes</typeStatus>
Nos
<quantity id="0D529B428041D340FCEBFB48FCB7FB45" box="[798,864,1196,1215]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="2.54" metricValueMin="0.254" pageId="1" pageNumber="564" unit="in" value="5.5" valueMax="10.0" valueMin="1.0">1–10 in</quantity>
<collectionCode id="ACBBAE628041D340FC90FB48FC4BFB45" box="[869,924,1196,1215]" pageId="1" pageNumber="564">RSCV</collectionCode>
, No.
<quantity id="0D529B428041D340FC3EFB48FC20FB45" box="[971,1015,1196,1215]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="1" pageNumber="564" unit="in" value="11.0">11 in</quantity>
<collectionCode id="ACBBAE628041D340FC09FB48FBEAFB45" box="[1020,1085,1196,1215]" pageId="1" pageNumber="564">MBCO</collectionCode>
, Nos
<quantity id="0D529B428041D340FF6CFB23FF32FB20" box="[153,229,1223,1242]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="1" pageNumber="564" unit="in" value="13.5" valueMax="15.0" valueMin="12.0">12–15 in</quantity>
<collectionCode id="ACBBAE628041D340FF1FFB23FEFEFB20" box="[234,297,1223,1242]" pageId="1" pageNumber="564">MLCR</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="82B0652C8041D345FF84FB16FEC0F9AC" lastPageId="4" lastPageNumber="567" pageId="1" pageNumber="564" type="description">
<paragraph id="CA1536A78041D340FF84FB16FD8FFAB3" blockId="1.[113,1090,1265,1609]" pageId="1" pageNumber="564">
<emphasis id="F8DEEAB58041D340FF84FB16FEEAFAF3" bold="true" box="[113,317,1265,1289]" pageId="1" pageNumber="564">
Description.
<emphasis id="F8DEEAB58041D340FEF0FB15FEEAFAF3" bold="true" box="[261,317,1265,1289]" italics="true" pageId="1" pageNumber="564">Male</emphasis>
</emphasis>
(
<typeStatus id="151188058041D340FEB8FB16FE66FAF3" box="[333,433,1266,1289]" pageId="1" pageNumber="564" type="holotype">holotype</typeStatus>
). Body length
<quantity id="0D529B428041D340FDA7FB16FD65FAF3" box="[594,690,1266,1289]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="1" pageNumber="564" unit="mm" value="23.2">23.2 mm</quantity>
(without pygidium). Body elongate, posteriorly slightly dilated. Dorsal surface and abdomen reddish brown, legs partly lighter, antennae pale brown, elytra pruinose (
<figureCitation id="52912A228041D340FDFCFAD6FD9FFAB3" box="[521,584,1330,1353]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="1" pageNumber="564">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="CA1536A78041D345FF65FAB6FD2BFE0C" blockId="1.[113,1090,1265,1609]" lastBlockId="4.[151,1128,223,1622]" lastPageId="4" lastPageNumber="567" pageId="1" pageNumber="564">
<emphasis id="F8DEEAB58041D340FF65FAB6FF06FA90" box="[144,209,1362,1386]" italics="true" pageId="1" pageNumber="564">Head.</emphasis>
Clypeus 2.19 times wider than long, with anterior angles broadly round and anterior margin strongly upturned and bisinuate; surface punctate, with smooth facets between punctures; posterolaterally oriented, pointed, semierect scale issues from centre of each puncture; some medial scales narrower than others, fronto-clypeal suture weakly indicated. Frons densely and coarsely punctate, with a scale issuing from each puncture; scales near midline point laterally toward a longitudinal bulge in each half of frons, forming a bisinuate pattern (
<figureCitation id="52912A228041D340FF3EF9F6FED9F9D3" box="[203,270,1554,1577]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="1" pageNumber="564">Fig. 2</figureCitation>
). Labrum bilobed, anterior angles obtuse but distinct, macrosetae longest at edge of clypeus and around eye canthus. Genae rugo-punctate, with group of short macrosetae. Occiput deeply but sparsely punctate, with pointed scales oriented posteriorly. Eye canthus long, with macrosetae in two regular rows; anterior conical macrosetae long, posterior conical macrosetae ¿ve times shorter than clypeal scales. Antennae decamerous. Antennal club hexamerous, straight and 1.5 times longer than antennal shaft (antennomeres 1–4 combined). Antennal club on apex with irregular, ¿ne, short macrosetae with sensilla pits. Antennomere 1 flat, ventrally with long setae, as long as antennomers II and III combined; antennomere 4 short, anteriorly pointed (
<figureCitation id="52912A228044D345FE54FE7BFE37FE4C" box="[417,480,415,438]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="4" pageNumber="567">Fig. 3</figureCitation>
). Terminal maxillary palpomere widest at midlength, apically truncate, with a terminal tubercle on inner, more angular side, and on dorsal surface with a large oval alutaceous area widening toward apex (
<figureCitation id="52912A228044D345FD58FE3BFD38FE0C" box="[685,751,479,502]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="4" pageNumber="567">Fig. 4</figureCitation>
).
</paragraph>
<caption id="9ED5662F8042D343FF62F9E5FE62F9B3" ID-DOI="http://doi.org/10.5281/zenodo.5301247" ID-Zenodo-Dep="5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="2" pageNumber="565" startId="2.[151,188,1537,1556]" targetBox="[164,1126,235,1429]" targetPageId="2">
<paragraph id="CA1536A78042D343FF62F9E5FE62F9B3" blockId="2.[151,1127,1537,1609]" pageId="2" pageNumber="565">
Figs 1–7.
<taxonomicName id="0DAA4D248042D343FF1FF9E5FE4EF9EE" authority="Sehnal, 2014" authorityName="Sehnal" authorityYear="2014" box="[234,409,1537,1556]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="sagala" status="sp. nov.">
<emphasis id="F8DEEAB58042D343FF1FF9E5FE4EF9EE" box="[234,409,1537,1556]" italics="true" pageId="2" pageNumber="565">Engistanoxia sagala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3ED57CE8042D343FE6BF9E5FE35F9EE" box="[414,482,1537,1556]" pageId="2" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm.
</paragraph>
</caption>
<caption id="9ED5662F8043D342FF84F9EAFC5DF9AC" ID-DOI="http://doi.org/10.5281/zenodo.5301249" ID-Zenodo-Dep="5301249" httpUri="https://zenodo.org/record/5301249/files/figure.png" pageId="3" pageNumber="566" startId="3.[113,150,1550,1569]" targetBox="[117,1067,222,1470]" targetPageId="3">
<paragraph id="CA1536A78043D342FF84F9EAFC5DF9AC" blockId="3.[113,1089,1549,1622]" pageId="3" pageNumber="566">
Figs 8–15.
<taxonomicName id="0DAA4D248043D342FF3AF9EAFDFBF9DB" authority="Lacroix, 2002" authorityName="Lacroix" authorityYear="2002" box="[207,556,1549,1569]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="dessertineae">
<emphasis id="F8DEEAB58043D342FF3AF9EAFE78F9DA" box="[207,431,1549,1568]" italics="true" pageId="3" pageNumber="566">Engistanoxia dessertineae</emphasis>
<bibRefCitation id="AE3B4B568043D342FE46F9EAFDFBF9DB" author="LACROIX M." box="[435,556,1550,1569]" pageId="3" pageNumber="566" pagination="153 - 168" refId="ref2934" refString="LACROIX M. 2002: Nouveaux Melolonthidae afrotropicaux (Coleoptera, Scarabaeoidea). Coleopteres 8 (9): 153 - 168. LACROIX M. 2010: Melolonthinae afrotropicaux. Genera et Catalogue. Collection Hannetons, Edition M. Lacroix, Paris, 277 pp." type="book chapter" year="2002">Lacroix, 2002</bibRefCitation>
</taxonomicName>
, holotype male. 8 – habitus, dorsal view; 9 – head, dorsal view; 10 – antenna, dorsal view; 11 – maxillary palp, dorsolateral view; 12 – pronotum, dorsal view; 13 – aedeagus, dorsal view; 14 – aedeagus, lateral view, 15 – lateral margin of pronotum, dorsal view. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="CA1536A78044D345FF40FE1BFBD8FD4C" blockId="4.[151,1128,223,1622]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF40FE1BFECAFDEC" box="[181,285,511,534]" italics="true" pageId="4" pageNumber="567">Pronotum</emphasis>
weekly convex, approximately heptagonal (
<figureCitation id="52912A228044D345FD03FE1BFCE0FDEC" box="[758,823,511,534]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="4" pageNumber="567">Fig. 5</figureCitation>
), 1.7 times wider than long, widest in posterior third, with distinct impressed medial line.Anterior and posterior angles round but discernible. Lateral margins crenulate, with a yellow hook-shaped macroseta issuing from each crenulation. Surface densely punctate; punctures unevenly distributed, spaces between them smooth and matte; each puncture bearing pointed scale only about half as long as scales on clypeus and frons. Scales forming groups oriented mostly toward midline (
<figureCitation id="52912A228044D345FC36FD7BFBD5FD4C" box="[963,1026,671,694]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="4" pageNumber="567">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="CA1536A78044D345FF40FD5AFBB0FD0C" blockId="4.[151,1128,223,1622]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF40FD5AFE8AFD2C" box="[181,349,702,726]" italics="true" pageId="4" pageNumber="567">Scutellar shield</emphasis>
wider than long, triangular, evenly punctate in anterior half, with scales similar to those on pronotum (
<figureCitation id="52912A228044D345FE21FD3BFDC4FD0C" box="[468,531,735,758]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="4" pageNumber="567">Fig. 5</figureCitation>
) and elytra, along midline with a strip of long, ¿ne hairs.
</paragraph>
<paragraph id="CA1536A78044D345FF40FD1AFCE6FC4C" blockId="4.[151,1128,223,1622]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF40FD1AFF2FFCEC" box="[181,248,766,790]" italics="true" pageId="4" pageNumber="567">Elytra</emphasis>
convex, elytron 1.4 times longer than wide. Each elytron quadricostate (including sutural costa), costae reaching from humeral to terminal bulge, costae 2 and 4 somewhat shorter. Part of elytron between terminal bulge and apex short, steeply inclined. Crests of costae glabrous, their sides and intervals unevenly covered by punctures and scales similar to those on pronotum. Spaces between punctures and scales ¿nely rugate and matte. Lateral margins with hairs twice as long as those on pronotal margins.
</paragraph>
<paragraph id="CA1536A78044D345FF40FC5BFE86FC2C" blockId="4.[151,1128,223,1622]" box="[181,337,959,982]" pageId="4" pageNumber="567">Macropterous.</paragraph>
<paragraph id="CA1536A78044D345FF40FC3BFE7BFB0C" blockId="4.[151,1128,223,1622]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF40FC3BFF38FC0C" box="[181,239,991,1014]" italics="true" pageId="4" pageNumber="567">Legs.</emphasis>
All femora shiny, irregularly punctate, macrosetae long. Protibia tridentate, claws strongly curved, microrugate, each with a minute basal tooth and a much larger tooth at midlength. Meso- and metatibiae expanded apically, densely covered by broad macrosetae, spaces between individual macrosetae larger than diameter of macrosetae, with one oblique interrupted carina externally. Apical edge with a row of short, stout macrosetae of equal length, terminal calcars stout, long, lower calcar little shorter than upper; pro-, meso- and metatarsomeres without patches of short, dense macrosetae; metatarsomeres covered by isolated, sparse macrosetae ventrally; meso- and metatibiae shallowly puctate, ¿ne long macrosetation recumbent.
</paragraph>
<paragraph id="CA1536A78044D345FF40FB1AFD96FA8C" blockId="4.[151,1128,223,1622]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF40FB1AFE59FAEC" box="[181,398,1278,1302]" italics="true" pageId="4" pageNumber="567">Abdominal ventrites</emphasis>
reddish brown, matte, ¿nely punctate, covered by oval white scales and a row of semierect yellowish-white macrosetae. Pygidium slightly concave, shallowly punctate, with yellowish-white caudally oriented scales becoming toward posterior margin ¿ne, short, yellowish-white macrosetae.
</paragraph>
<paragraph id="CA1536A78044D345FF40FA9AFDFDFA0C" blockId="4.[151,1128,223,1622]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF40FA9AFE84FA6C" box="[181,339,1406,1430]" italics="true" pageId="4" pageNumber="567">Male genitalia</emphasis>
(
<figureCitation id="52912A228044D345FE97FA9BFE11FA6C" box="[354,454,1407,1430]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="4" pageNumber="567">Figs 6–7</figureCitation>
). Aedeagus symmetrical. Phallobase in lateral view with three extensions, ventrolaterally deeply excised. Parameres forming a symmetrical lobe with medial oval summit and an oval concavity in its centre. Dorsal part of lobe steeply elevated and then reclined at right angle ventral to base.
</paragraph>
<paragraph id="CA1536A78044D345FF40FA1AFB81F9EC" blockId="4.[151,1128,223,1622]" box="[181,1110,1534,1558]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF40FA1AFED8F9EC" bold="true" box="[181,271,1534,1558]" italics="true" pageId="4" pageNumber="567">Female.</emphasis>
Similar to male but has a shorter antennal club and wider and shorter protibia.
</paragraph>
<paragraph id="CA1536A78044D345FF62F9FBFEC0F9AC" blockId="4.[151,1128,223,1622]" pageId="4" pageNumber="567">
<emphasis id="F8DEEAB58044D345FF62F9FBFEC1F9CC" bold="true" box="[151,278,1567,1590]" pageId="4" pageNumber="567">Variability.</emphasis>
Males.
<typeStatus id="151188058044D345FE9EF9FBFE04F9CC" box="[363,467,1567,1590]" pageId="4" pageNumber="567" type="paratype">Paratypes</typeStatus>
somewhat variable in body length (
<quantity id="0D529B428044D345FCBBF9FBFC38F9CC" box="[846,1007,1567,1590]" metricMagnitude="-2" metricUnit="m" metricValue="2.0799999999999996" metricValueMax="2.32" metricValueMin="1.8399999999999999" pageId="4" pageNumber="567" unit="mm" value="20.799999999999997" valueMax="23.2" valueMin="18.4">18.4–23.2 mm</quantity>
). Colour as in
<typeStatus id="151188058044D345FF46F9DBFEC3F9AC" box="[179,276,1599,1622]" pageId="4" pageNumber="567" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="82B0652C8045D344FF84FF38FCBCFE89" pageId="5" pageNumber="568" type="diagnosis">
<paragraph id="CA1536A78045D344FF84FF38FCBCFE89" blockId="5.[113,1089,219,499]" pageId="5" pageNumber="568">
<emphasis id="F8DEEAB58045D344FF84FF38FEB1FF09" bold="true" box="[113,358,220,243]" pageId="5" pageNumber="568">Differential diagnosis.</emphasis>
The habitus of
<taxonomicName id="0DAA4D248045D344FDF6FF38FD0FFF09" authority="Sehnal, 2014" authorityName="Sehnal" authorityYear="2014" box="[515,728,219,243]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="568" phylum="Arthropoda" rank="species" species="sagala" status="sp. nov.">
<emphasis id="F8DEEAB58045D344FDF6FF38FD0FFF09" box="[515,728,219,243]" italics="true" pageId="5" pageNumber="568">Engistanoxia sagala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3ED57CE8045D344FD28FF38FCFAFF09" box="[733,813,220,243]" pageId="5" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
is close to
<taxonomicName id="0DAA4D248045D344FC6EFF38FBEDFF09" authorityName="Lacroix" authorityYear="2002" box="[923,1082,219,243]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="568" phylum="Arthropoda" rank="species" species="dessertineae">
<emphasis id="F8DEEAB58045D344FC6EFF38FBEDFF09" box="[923,1082,219,243]" italics="true" pageId="5" pageNumber="568">E. dessertineae</emphasis>
</taxonomicName>
, from which it differs in the pointed but not prolonged antennomere IV, size of the accessory tooth on tarsal claws, crenulate lateral margins of the pronotum, shape and pattern of scales on the head and pronotum, shape of the terminal maxillary palpomere and its pit, frontal margin of clypeus, bisinuate phallobase and shape of the parameres (
<figureCitation id="52912A228045D344FD08FEB8FC88FE89" box="[765,863,348,371]" captionStart="Figs 1–7" captionStartId="2.[151,188,1537,1556]" captionTargetBox="[164,1126,235,1429]" captionTargetId="figure-58@2.[149,1131,218,1455]" captionTargetPageId="2" captionText="Figs 1–7. Engistanoxia sagala sp. nov., holotype male. 1 – habitus, dorsal view; 2 – head, dorsal view; 3 – antenna, dorsal view; 4 – maxillary palp, dorsolateral view; 5 – pronotum, dorsal view; 6 – aedeagus, dorsal view; 7 – aedeagus, lateral view. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301247" httpUri="https://zenodo.org/record/5301247/files/figure.png" pageId="5" pageNumber="568">Figs 6–7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="82B0652C8045D344FF84FE98FF1CFE09" pageId="5" pageNumber="568" type="etymology">
<paragraph id="CA1536A78045D344FF84FE98FD89FE49" blockId="5.[113,1089,219,499]" pageId="5" pageNumber="568">
<emphasis id="F8DEEAB58045D344FF84FE98FF25FE69" bold="true" box="[113,242,380,403]" pageId="5" pageNumber="568">Etymology.</emphasis>
The species epithet refers to the
<typeStatus id="151188058045D344FDB1FE98FDA5FE69" box="[580,626,380,403]" pageId="5" pageNumber="568">type</typeStatus>
locality,
<taxonomicName id="0DAA4D248045D344FD25FE98FC87FE69" authority="Hills" authorityName="Hills" box="[720,848,380,403]" class="Insecta" family="Melolonthidae" genus="Engistanoxia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="568" phylum="Arthropoda" rank="species" species="sagala">Sagala Hills</taxonomicName>
in
<collectingCountry id="B2BD76378045D344FC9BFE98FC64FE69" box="[878,947,380,403]" name="Kenya" pageId="5" pageNumber="568">Kenya</collectingCountry>
(
<collectingRegion id="086EF8458045D344FC4AFE98FF69FE49" country="Kenya" name="Taita/Taveta" pageId="5" pageNumber="568">Taita Taveta county</collectingRegion>
), and the name is a noun in apposition.
</paragraph>
<paragraph id="CA1536A78045D344FF84FE58FF1CFE09" blockId="5.[113,1089,219,499]" pageId="5" pageNumber="568">
<emphasis id="F8DEEAB58045D344FF84FE58FE52FE29" bold="true" box="[113,389,444,467]" pageId="5" pageNumber="568">Collection circumstaces.</emphasis>
Adults were collected at light (M. BednaĜík and I. MartinĤ, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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