348 lines
48 KiB
XML
348 lines
48 KiB
XML
|
<document id="8F3D23DB3D863966B013675DFCA86EFA" ID-CLB-Dataset="46838" ID-DOI="10.5281/zenodo.201061" ID-GBIF-Dataset="5e7d0d28-851a-429b-8e52-8fe8793095b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="201061" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460359522896" checkinUser="plazi" docAuthor="Markhaseva, Elena L. & Renz, Jasmin" docDate="2011" docId="DE292E1AFFE7FFA8ADE23696F167C15A" docLanguage="en" docName="zt02889p068.pdf" docOrigin="Zootaxa 2889" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Byrathis penicillatus Markhaseva & Renz, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="61" masterDocId="22105662FFE0FFA4AD753601F010C065" masterDocTitle="Two new Byrathis species (Copepoda: Calanoida) from the deep South Atlantic and Southern Ocean and first description of an adult male" masterLastPageNumber="68" masterPageNumber="49" pageNumber="56" updateTime="1698294284746" updateUser="ExternalLinkService">
|
|||
|
|
<mods:mods id="8E5E68673500BD58786604B521C91E34" xmlns:mods="http://www.loc.gov/mods/v3">
|
|||
|
|
<mods:titleInfo id="D919FD64B2130D74A6A7B979766DB971">
|
|||
|
|
<mods:title id="CBF68B2F19624682939A3974B3579E1A">Two new Byrathis species (Copepoda: Calanoida) from the deep South Atlantic and Southern Ocean and first description of an adult male</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:name id="110439ADE9E67FC358DBD241E3174A07" type="personal">
|
|||
|
|
<mods:role id="1D6A24CB38C56483DCA8FB81CF2C719B">
|
|||
|
|
<mods:roleTerm id="3437C88F76E4AF7471DDF421C4CED3A4">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="F590C6BD6DDC754C2AD09DDB8FAAEC7D">Markhaseva, Elena L.</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name id="8A96D86AAD3FB63F5B05FC5F48209831" type="personal">
|
|||
|
|
<mods:role id="6F9FF612145C402E890039E4F9BE29A6">
|
|||
|
|
<mods:roleTerm id="B798C0FCBFDF85BE3E9ED70936177B18">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="64AC01016605B0ADBBC4B8365F2E0250">Renz, Jasmin</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:typeOfResource id="1B8C9A071746E00F8C4316F6D4040176">text</mods:typeOfResource>
|
|||
|
|
<mods:relatedItem id="07EEB4C66C9F3543674DF9FB311D8367" type="host">
|
|||
|
|
<mods:titleInfo id="4225F6A967EA0CE58BCEBAB785F5F0E4">
|
|||
|
|
<mods:title id="66DC8CCDADEC82F6028A4BCAB7BFB209">Zootaxa</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:part id="5B647E8D4D002BFACBDA679F48EB7DE8">
|
|||
|
|
<mods:date id="6A1D3414F68EBEE204568B0C6446B029">2011</mods:date>
|
|||
|
|
<mods:detail id="FBD65DCF6CB745E178DEA3C259BB3B10" type="volume">
|
|||
|
|
<mods:number id="344E71DCE8DD4E9BF7BCA6DFB1FE9258">2889</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:extent id="A11BC4AE6AF27C3FE9D8D7FE9DA939A9" unit="page">
|
|||
|
|
<mods:start id="FB9CDA9834B9D07C64466580CB9F77B1">49</mods:start>
|
|||
|
|
<mods:end id="B7D1EFFA2F69FA6BEF8B2305A1B00D1B">68</mods:end>
|
|||
|
|
</mods:extent>
|
|||
|
|
</mods:part>
|
|||
|
|
</mods:relatedItem>
|
|||
|
|
<mods:classification id="BB2DDE249D49FCD5B771973869D6B4E8">journal article</mods:classification>
|
|||
|
|
<mods:identifier id="1FC573514449876EB35346704589A1EF" type="CLB-Dataset">46838</mods:identifier>
|
|||
|
|
<mods:identifier id="59650C3831AD70FB9C211FCF9FFE2380" type="DOI">10.5281/zenodo.201061</mods:identifier>
|
|||
|
|
<mods:identifier id="6B80BE145240DA67329036D067571974" type="GBIF-Dataset">5e7d0d28-851a-429b-8e52-8fe8793095b4</mods:identifier>
|
|||
|
|
<mods:identifier id="1FF6A82A5DAFA5456D3A7F808C0DE623" type="ISSN">1175-5326</mods:identifier>
|
|||
|
|
<mods:identifier id="58261429FF031254A39E17C625181B08" type="Zenodo-Dep">201061</mods:identifier>
|
|||
|
|
</mods:mods>
|
|||
|
|
<treatment id="DE292E1AFFE7FFA8ADE23696F167C15A" ID-DOI="http://doi.org/10.5281/zenodo.6193226" ID-GBIF-Taxon="119409395" ID-Zenodo-Dep="6193226" LSID="urn:lsid:plazi:treatment:DE292E1AFFE7FFA8ADE23696F167C15A" httpUri="http://treatment.plazi.org/id/DE292E1AFFE7FFA8ADE23696F167C15A" lastPageId="12" lastPageNumber="61" pageId="7" pageNumber="56">
|
|||
|
|
<subSubSection id="1E9ACC87FFE7FFA3ADE23696F11AC0B7" pageId="7" pageNumber="56" type="nomenclature">
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADE23696F1E6C0D4" blockId="7.[151,502,151,210]" box="[151,502,151,177]" pageId="7" pageNumber="56">
|
|||
|
|
<heading id="0D772860FFE7FFA3ADE23696F1E6C0D4" bold="true" box="[151,502,151,177]" fontSize="11" level="1" pageId="7" pageNumber="56" reason="1">
|
|||
|
|
<emphasis id="64F4431EFFE7FFA3ADE23696F1E6C0D4" bold="true" box="[151,502,151,177]" pageId="7" pageNumber="56">
|
|||
|
|
<taxonomicName id="9180E48FFFE7FFA3ADE23696F181C0D4" box="[151,401,151,177]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFE7FFA3ADE23696F181C0D4" bold="true" box="[151,401,151,177]" italics="true" pageId="7" pageNumber="56">Byrathis penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFE7FFA3ACED3696F1E6C0D4" box="[408,502,151,177]" pageId="7" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADE236BBF11AC0B7" blockId="7.[151,502,151,210]" box="[151,266,186,210]" pageId="7" pageNumber="56">
|
|||
|
|
(
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3ADEA36BBF113C0B7" box="[159,259,186,210]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="8.[151,249,1915,1938]" captionStartId-1="9.[151,249,1883,1906]" captionStartId-2="10.[151,249,1884,1907]" captionStartId-3="11.[151,249,1453,1476]" captionStartId-4="12.[151,249,1440,1463]" captionTargetBox-0="[213,1372,208,1875]" captionTargetBox-1="[189,1419,195,1855]" captionTargetBox-2="[162,1421,234,1829]" captionTargetBox-3="[193,1387,190,1436]" captionTargetBox-4="[196,1396,368,1418]" captionTargetId-0="figure@8.[188,1398,193,1894]" captionTargetId-1="figure@9.[161,1425,193,1865]" captionTargetId-2="figure@10.[151,1436,193,1863]" captionTargetId-3="figure@11.[170,1417,167,1436]" captionTargetId-4="figure@12.[188,1398,347,1419]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 5. Byrathis penicillatus sp. nov. Female. A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, lateral view; D, rostrum, ventral view; E, posterior prosome and urosome, dorsal view; F, posterior prosome and urosome, lateral view; G, posterior prosome and genital double somite, lateral view. A – B, D, E – F, holotype; C, G, paratype. Scale bars: A, B, 0.5 mm, C – F, 0.1 mm." captionText-1="FIGURE 6. Byrathis penicillatus sp. nov. Female. A, antennule, articulating segments 1 – 17 (ancestral segments I – XX); B, antennule, articulating segments 18 – 24 (ancestral segments XXI – XXVIII); C, antennule, segment 24 (ancestral segment XXVII – XXVIII); D, antenna; E, mandible; F, mandible, gnathobase; G, mandible, gnathobase, cutting edge; H, maxillule; I, maxillule, praecoxal arthrite. D, E, H, holotype; A – C, F – G, I, paratype; G, for smaller size (3.35 mm) paratype. Scale bars: 0.1 mm." captionText-2="FIGURE 7. Byrathis penicillatus sp. nov. Female. A, maxilla (sensory setae are not figured); B, maxilla, sensory setae on endopod; C, maxilla, sensory setae on endopod, worm-like setae are not figured); D, maxilliped; E, P 5. A, D, holotype; B – C, E, paratype; C, for smaller size (3.35 mm) paratype. Scale bars 0.1 mm." captionText-3="FIGURE 8. Byrathis penicillatus sp. nov. Female, holotype. A, P 1; B, P 2; C, P 3; D, P 4. Scale bars 0.1 mm." captionText-4="FIGURE 9. Distribution of species of Byrathis in the World Ocean: black rhomboid B. macrocephalon (Grice & Hulsemann, 1970); circle outlined, B. penicillatus sp. nov.; black square, B. laurenae Markhaseva & Ferrari, 2005; big triangle outlined, B. arnei Schulz, 2006; black small triangle, B. divae sp. nov.; rhomboid outlined, B. laptevorum (Markhaseva, 1998); rhomboid outlined with crest, B. volcani Markhaseva & Ferrari, 2005." httpUri-0="https://zenodo.org/record/201066/files/figure.png" httpUri-1="https://zenodo.org/record/201067/files/figure.png" httpUri-2="https://zenodo.org/record/201068/files/figure.png" httpUri-3="https://zenodo.org/record/201069/files/figure.png" httpUri-4="https://zenodo.org/record/201070/files/figure.png" pageId="7" pageNumber="56">Figs 5–9</figureCitation>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1E9ACC87FFE7FFA3ADE236FCF334C250" pageId="7" pageNumber="56" type="materials_examined">
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADE236FCF41AC15C" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
<emphasis id="64F4431EFFE7FFA3ADE236FCF11BC170" bold="true" box="[151,267,253,277]" pageId="7" pageNumber="56">
|
|||
|
|
<typeStatus id="893B21AEFFE7FFA3ADE236FCF116C170" box="[151,262,253,277]" pageId="7" pageNumber="56" type="holotype">Holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Partly dissected adult female, body length
|
|||
|
|
<quantity id="917832E9FFE7FFA3AF8F36FCF372C170" box="[762,866,253,277]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" pageId="7" pageNumber="56" unit="mm" value="4.05">4.05 mm</quantity>
|
|||
|
|
.
|
|||
|
|
<collectionCode id="309107C9FFE7FFA3AE1A36FFF3BDC170" box="[879,941,254,277]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="7" pageNumber="56">ZMH</collectionCode>
|
|||
|
|
Reg. no. K–42157. Collected on
|
|||
|
|
<date id="223EB9CCFFE7FFA3A85B36FCF0DFC15C" pageId="7" pageNumber="56" value="2005-03-15">15 March 2005</date>
|
|||
|
|
by the
|
|||
|
|
<collectionCode id="309107C9FFE7FFA3AC573720F176C15C" box="[290,358,289,313]" pageId="7" pageNumber="56">DIVA</collectionCode>
|
|||
|
|
–II expedition above the sea bed at abyssal depths (
|
|||
|
|
<quantity id="917832E9FFE7FFA3AED63720F3EEC15C" box="[931,1022,289,313]" metricMagnitude="3" metricUnit="m" metricValue="5.058" pageId="7" pageNumber="56" unit="m" value="5058.0">5058 m</quantity>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB03744F2ABC274" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
<emphasis id="64F4431EFFE7FFA3ADB03744F12DC138" bold="true" box="[197,317,325,349]" pageId="7" pageNumber="56">
|
|||
|
|
<typeStatus id="893B21AEFFE7FFA3ADB03744F12DC138" box="[197,317,325,349]" pageId="7" pageNumber="56" type="paratype">Paratypes</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
, 1 partly dissected adult female, body length
|
|||
|
|
<quantity id="917832E9FFE7FFA3AE3F3744F3A3C138" box="[842,947,325,349]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" pageId="7" pageNumber="56" unit="mm" value="3.95">3.95 mm</quantity>
|
|||
|
|
.
|
|||
|
|
<collectionCode id="309107C9FFE7FFA3AECA3747F3EDC138" box="[959,1021,326,349]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="7" pageNumber="56">ZMH</collectionCode>
|
|||
|
|
Reg. no. K–42158. Collected on
|
|||
|
|
<date id="223EB9CCFFE7FFA3A8F53744F134C1E4" pageId="7" pageNumber="56" value="2005-03-15">15 March 2005</date>
|
|||
|
|
by the
|
|||
|
|
<collectionCode id="309107C9FFE7FFA3ACF33768F1DAC1E4" box="[390,458,361,385]" pageId="7" pageNumber="56">DIVA</collectionCode>
|
|||
|
|
–II expedition above the sea bed at abyssal depths (
|
|||
|
|
<quantity id="917832E9FFE7FFA3A9413768F481C1E4" box="[1076,1169,361,385]" metricMagnitude="3" metricUnit="m" metricValue="5.05" pageId="7" pageNumber="56" unit="m" value="5050.0">5050 m</quantity>
|
|||
|
|
) in the South Atlantic (
|
|||
|
|
<geoCoordinate id="33B4F9CBFFE7FFA3ADEA378CF11EC1C0" box="[159,270,397,421]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="92" value="-0.14166667">00º08.5’S</geoCoordinate>
|
|||
|
|
<geoCoordinate id="33B4F9CBFFE7FFA3AC63378CF184C1C0" box="[278,404,397,421]" direction="west" orientation="longitude" pageId="7" pageNumber="56" precision="92" value="-2.5033333">02º30.2’W</geoCoordinate>
|
|||
|
|
); 1 partly dissected adult female, body length
|
|||
|
|
<quantity id="917832E9FFE7FFA3AEE2378CF3EEC1C0" box="[919,1022,397,421]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" pageId="7" pageNumber="56" unit="mm" value="3.35">3.35 mm</quantity>
|
|||
|
|
.
|
|||
|
|
<collectionCode id="309107C9FFE7FFA3A97C378CF429C1C0" box="[1033,1081,397,421]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="56">ZIN</collectionCode>
|
|||
|
|
–91103. Collected
|
|||
|
|
<date id="223EB9CCFFE7FFA3A87E378CF58CC1C0" box="[1291,1436,397,421]" pageId="7" pageNumber="56" value="2009-07-23">23 July 2009</date>
|
|||
|
|
by the
|
|||
|
|
<collectionCode id="309107C9FFE7FFA3AD9037B0F139C1AC" box="[229,297,433,457]" pageId="7" pageNumber="56">DIVA</collectionCode>
|
|||
|
|
–III expedition in the South Atlantic (
|
|||
|
|
<geoCoordinate id="33B4F9CBFFE7FFA3AFA137B0F33FC1AC" box="[724,815,433,457]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="925" value="-26.583334">26º35’S</geoCoordinate>
|
|||
|
|
<geoCoordinate id="33B4F9CBFFE7FFA3AE4337B0F38EC1AC" box="[822,926,433,457]" direction="west" orientation="longitude" pageId="7" pageNumber="56" precision="925" value="-35.233334">35º14’W</geoCoordinate>
|
|||
|
|
), above the sea bed at abyssal depths (
|
|||
|
|
<quantity id="917832E9FFE7FFA3A82137B0F0FFC188" metricMagnitude="3" metricUnit="m" metricValue="4.4855" metricValueMax="4.489" metricValueMin="4.482" pageId="7" pageNumber="56" unit="m" value="4485.5" valueMax="4489.0" valueMin="4482.0">4482– 4489 m</quantity>
|
|||
|
|
). Additional material:
|
|||
|
|
<specimenCount id="40865485FFE7FFA3AC9F37D4F249C188" box="[490,601,469,493]" pageId="7" pageNumber="56" type="female">4 females</specimenCount>
|
|||
|
|
, body length
|
|||
|
|
<quantity id="917832E9FFE7FFA3AF9A37D4F383C188" box="[751,915,469,493]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.9" metricValueMin="3.2" pageId="7" pageNumber="56" unit="mm" value="4.05" valueMax="4.9" valueMin="3.2">3.20–4.90 mm</quantity>
|
|||
|
|
, collected in the South Atlantic from the equator to about 36ºS at depths from
|
|||
|
|
<quantity id="917832E9FFE7FFA3AF7637F8F2A7C274" box="[515,695,505,529]" metricMagnitude="3" metricUnit="m" metricValue="4.998" metricValueMax="5.395" metricValueMin="4.601" pageId="7" pageNumber="56" unit="m" value="4998.0" valueMax="5395.0" valueMin="4601.0">4601 to 5395 m</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB0341CF334C250" blockId="7.[151,1437,253,2031]" box="[197,804,541,566]" pageId="7" pageNumber="56">
|
|||
|
|
<emphasis id="64F4431EFFE7FFA3ADB0341CF173C253" bold="true" box="[197,355,541,566]" pageId="7" pageNumber="56">
|
|||
|
|
<typeStatus id="893B21AEFFE7FFA3ADB0341CF110C250" box="[197,256,541,565]" pageId="7" pageNumber="56">Type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
</emphasis>
|
|||
|
|
South Atlantic (
|
|||
|
|
<geoCoordinate id="33B4F9CBFFE7FFA3AF69341CF29DC250" box="[540,653,541,565]" direction="south" orientation="latitude" pageId="7" pageNumber="56" precision="92" value="-0.020000001">00º01.2’S</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="33B4F9CBFFE7FFA3AFEC341CF307C250" box="[665,791,541,565]" direction="west" orientation="longitude" pageId="7" pageNumber="56" precision="92" value="-2.4783335">02º28.7’W</geoCoordinate>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1E9ACC87FFE7FFAFADB23441F201C62A" lastPageId="11" lastPageNumber="60" pageId="7" pageNumber="56" type="description">
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB23441F4F8C2A0" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
<emphasis id="64F4431EFFE7FFA3ADB23441F149C23C" bold="true" box="[199,345,576,601]" pageId="7" pageNumber="56">Description.</emphasis>
|
|||
|
|
Female. Body length
|
|||
|
|
<quantity id="917832E9FFE7FFA3AF2D3440F310C23C" box="[600,768,577,601]" metricMagnitude="-3" metricUnit="m" metricValue="4.125" metricValueMax="4.9" metricValueMin="3.35" pageId="7" pageNumber="56" unit="mm" value="4.125" valueMax="4.9" valueMin="3.35">3.35–4.90 mm</quantity>
|
|||
|
|
. Prosome 3.7–4.1 times as long as urosome. Rostrum as a triangular plate with 2 filaments (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AF783464F25EC218" box="[525,590,613,637]" captionStart="FIGURE 5" captionStartId="8.[151,249,1915,1938]" captionTargetBox="[213,1372,208,1875]" captionTargetId="figure@8.[188,1398,193,1894]" captionTargetPageId="8" captionText="FIGURE 5. Byrathis penicillatus sp. nov. Female. A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, lateral view; D, rostrum, ventral view; E, posterior prosome and urosome, dorsal view; F, posterior prosome and urosome, lateral view; G, posterior prosome and genital double somite, lateral view. A – B, D, E – F, holotype; C, G, paratype. Scale bars: A, B, 0.5 mm, C – F, 0.1 mm." httpUri="https://zenodo.org/record/201066/files/figure.png" pageId="7" pageNumber="56">Fig. 5</figureCitation>
|
|||
|
|
C–D). Cephalosome and pedigerous somite 1 and pedigerous somites 4 and 5 partly separate; posterior corners sharply triangular in dorsal and lateral view (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3A95E3488F47DC2C4" box="[1067,1133,649,673]" captionStart="FIGURE 5" captionStartId="8.[151,249,1915,1938]" captionTargetBox="[213,1372,208,1875]" captionTargetId="figure@8.[188,1398,193,1894]" captionTargetPageId="8" captionText="FIGURE 5. Byrathis penicillatus sp. nov. Female. A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, lateral view; D, rostrum, ventral view; E, posterior prosome and urosome, dorsal view; F, posterior prosome and urosome, lateral view; G, posterior prosome and genital double somite, lateral view. A – B, D, E – F, holotype; C, G, paratype. Scale bars: A, B, 0.5 mm, C – F, 0.1 mm." httpUri="https://zenodo.org/record/201066/files/figure.png" pageId="7" pageNumber="56">Fig. 5</figureCitation>
|
|||
|
|
A–B, E–G). Urosome of 4 somites. Caudal rami with 4 terminal setae, 1 ventral seta and 1 small dorsolateral seta (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3A91B34ACF4B8C2A0" box="[1134,1192,685,709]" captionStart="FIGURE 5" captionStartId="8.[151,249,1915,1938]" captionTargetBox="[213,1372,208,1875]" captionTargetId="figure@8.[188,1398,193,1894]" captionTargetPageId="8" captionText="FIGURE 5. Byrathis penicillatus sp. nov. Female. A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, lateral view; D, rostrum, ventral view; E, posterior prosome and urosome, dorsal view; F, posterior prosome and urosome, lateral view; G, posterior prosome and genital double somite, lateral view. A – B, D, E – F, holotype; C, G, paratype. Scale bars: A, B, 0.5 mm, C – F, 0.1 mm." httpUri="https://zenodo.org/record/201066/files/figure.png" pageId="7" pageNumber="56">Fig.5</figureCitation>
|
|||
|
|
E–F).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB034D0F0F3C3F8" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
Antennule (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AC3934D0F181C28C" box="[332,401,721,745]" captionStart="FIGURE 6" captionStartId="9.[151,249,1883,1906]" captionTargetBox="[189,1419,195,1855]" captionTargetId="figure@9.[161,1425,193,1865]" captionTargetPageId="9" captionText="FIGURE 6. Byrathis penicillatus sp. nov. Female. A, antennule, articulating segments 1 – 17 (ancestral segments I – XX); B, antennule, articulating segments 18 – 24 (ancestral segments XXI – XXVIII); C, antennule, segment 24 (ancestral segment XXVII – XXVIII); D, antenna; E, mandible; F, mandible, gnathobase; G, mandible, gnathobase, cutting edge; H, maxillule; I, maxillule, praecoxal arthrite. D, E, H, holotype; A – C, F – G, I, paratype; G, for smaller size (3.35 mm) paratype. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/201067/files/figure.png" pageId="7" pageNumber="56">Fig. 6</figureCitation>
|
|||
|
|
A–C) extending to distal margin of pedigerous somites 4–5, of 24 articulating segments; armature as follows:
|
|||
|
|
<date id="223EB9CCFFE7FFA3ACF434F4F1B9C368" box="[385,425,757,781]" bridgedPair="-" pageId="7" pageNumber="56">I–3</date>
|
|||
|
|
s, II–
|
|||
|
|
<date id="223EB9CCFFE7FFA3AC9434F4F20BC368" box="[481,539,757,781]" bridgedPair="-" pageId="7" pageNumber="56">IV–6</date>
|
|||
|
|
s + 1ae,
|
|||
|
|
<date id="223EB9CCFFE7FFA3AF0334F4F2B8C368" box="[630,680,757,781]" bridgedPair="-" pageId="7" pageNumber="56">V–2</date>
|
|||
|
|
s + 1ae,
|
|||
|
|
<date id="223EB9CCFFE7FFA3AE7734F4F32EC368" box="[770,830,757,781]" bridgedPair="-" pageId="7" pageNumber="56">VI–2</date>
|
|||
|
|
s,
|
|||
|
|
<date id="223EB9CCFFE7FFA3AE2E34F4F38FC368" box="[859,927,757,781]" bridgedPair="-" pageId="7" pageNumber="56">VII–2</date>
|
|||
|
|
s + 1ae,
|
|||
|
|
<date id="223EB9CCFFE7FFA3AE8F34F4F45DC368" box="[1018,1101,757,781]" bridgedPair="-" pageId="7" pageNumber="56">VIII–1</date>
|
|||
|
|
s+1?,
|
|||
|
|
<date id="223EB9CCFFE7FFA3A9FC34F4F4D5C368" box="[1161,1221,757,781]" bridgedPair="-" pageId="7" pageNumber="56">IX–2</date>
|
|||
|
|
s + 1ae, X–
|
|||
|
|
<date id="223EB9CCFFE7FFA3A83734F4F56DC368" box="[1346,1405,757,781]" bridgedPair="-" pageId="7" pageNumber="56">XI–4</date>
|
|||
|
|
s + 1ae,
|
|||
|
|
<date id="223EB9CCFFE7FFA3ADB93518F102C354" box="[204,274,793,817]" bridgedPair="-" pageId="7" pageNumber="56">XII–1</date>
|
|||
|
|
?, XIII–1?, XIV–2s + 1ae, XV–1?, XVI–2s + 1ae, XVII–1?, XVIII–2?, XIX–1?, XX–2?, XXI–1s + 1ae, XXII–1?, XXIII–1s, XXIV–1s + 1?, XXV–2s, XXVI–2s, XXVII–XXVIII–4s + 1ae, (setation on
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3A870353CF559C330" box="[1285,1353,829,853]" captionStart="FIGURE 6" captionStartId="9.[151,249,1883,1906]" captionTargetBox="[189,1419,195,1855]" captionTargetId="figure@9.[161,1425,193,1865]" captionTargetPageId="9" captionText="FIGURE 6. Byrathis penicillatus sp. nov. Female. A, antennule, articulating segments 1 – 17 (ancestral segments I – XX); B, antennule, articulating segments 18 – 24 (ancestral segments XXI – XXVIII); C, antennule, segment 24 (ancestral segment XXVII – XXVIII); D, antenna; E, mandible; F, mandible, gnathobase; G, mandible, gnathobase, cutting edge; H, maxillule; I, maxillule, praecoxal arthrite. D, E, H, holotype; A – C, F – G, I, paratype; G, for smaller size (3.35 mm) paratype. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/201067/files/figure.png" pageId="7" pageNumber="56">Fig. 6</figureCitation>
|
|||
|
|
A–C is given for segments II–IV, VI, VII, IX–XI, XIV, XVI and XXIII after the
|
|||
|
|
<typeStatus id="893B21AEFFE7FFA3AEB63560F435C31C" box="[963,1061,865,889]" pageId="7" pageNumber="56" type="holotype">holotype</typeStatus>
|
|||
|
|
and supplemented from the
|
|||
|
|
<typeStatus id="893B21AEFFE7FFA3A8173563F0C7C3F8" pageId="7" pageNumber="56" type="paratype">paratypes</typeStatus>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB235ABF0C6C448" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
Antenna (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AC4335A8F16AC3A4" box="[310,378,937,961]" captionStart="FIGURE 6" captionStartId="9.[151,249,1883,1906]" captionTargetBox="[189,1419,195,1855]" captionTargetId="figure@9.[161,1425,193,1865]" captionTargetPageId="9" captionText="FIGURE 6. Byrathis penicillatus sp. nov. Female. A, antennule, articulating segments 1 – 17 (ancestral segments I – XX); B, antennule, articulating segments 18 – 24 (ancestral segments XXI – XXVIII); C, antennule, segment 24 (ancestral segment XXVII – XXVIII); D, antenna; E, mandible; F, mandible, gnathobase; G, mandible, gnathobase, cutting edge; H, maxillule; I, maxillule, praecoxal arthrite. D, E, H, holotype; A – C, F – G, I, paratype; G, for smaller size (3.35 mm) paratype. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/201067/files/figure.png" pageId="7" pageNumber="56">Fig. 6</figureCitation>
|
|||
|
|
D), coxa with 1 seta, basis with 2 setae; exopod of 7 free segments with 1, 1-1-1-1, 1, 1, 1, 1, and 3 setae, seta on proximal exopod segment rudimentary, following complex segment with 3 rudimentary setae, partly fused with first short segment bearing long seta; first endopodal segment with 2 setae, second with 8 + 7 setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB03238F2E1C4D8" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
Mandible (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AC353238F194C434" box="[320,388,1081,1105]" captionStart="FIGURE 6" captionStartId="9.[151,249,1883,1906]" captionTargetBox="[189,1419,195,1855]" captionTargetId="figure@9.[161,1425,193,1865]" captionTargetPageId="9" captionText="FIGURE 6. Byrathis penicillatus sp. nov. Female. A, antennule, articulating segments 1 – 17 (ancestral segments I – XX); B, antennule, articulating segments 18 – 24 (ancestral segments XXI – XXVIII); C, antennule, segment 24 (ancestral segment XXVII – XXVIII); D, antenna; E, mandible; F, mandible, gnathobase; G, mandible, gnathobase, cutting edge; H, maxillule; I, maxillule, praecoxal arthrite. D, E, H, holotype; A – C, F – G, I, paratype; G, for smaller size (3.35 mm) paratype. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/201067/files/figure.png" pageId="7" pageNumber="56">Fig. 6</figureCitation>
|
|||
|
|
E–G), gnathobase with crest, number of teeth on cutting edge of
|
|||
|
|
<typeStatus id="893B21AEFFE7FFA3A9103238F4D7C434" box="[1125,1223,1081,1105]" pageId="7" pageNumber="56" type="holotype">holotype</typeStatus>
|
|||
|
|
difficult to follow,
|
|||
|
|
<typeStatus id="893B21AEFFE7FFA3ADE2325FF0E7C413" box="[151,247,1118,1142]" pageId="7" pageNumber="56" type="paratype">paratype</typeStatus>
|
|||
|
|
bearing 4 large and 3 small teeth near dorsal seta, lateral tooth situated apart from the remaining teeth; basis with 3 setae; exopod 5-segmented with 1, 1, 1, 1, and 2 setae; endopod segment 1 with 2 setae, segment 2 with 9 setae and distal rows of small surface spinules.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB032C8F202C54C" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
Maxillule (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AC3732C8F195C484" box="[322,389,1225,1249]" captionStart="FIGURE 6" captionStartId="9.[151,249,1883,1906]" captionTargetBox="[189,1419,195,1855]" captionTargetId="figure@9.[161,1425,193,1865]" captionTargetPageId="9" captionText="FIGURE 6. Byrathis penicillatus sp. nov. Female. A, antennule, articulating segments 1 – 17 (ancestral segments I – XX); B, antennule, articulating segments 18 – 24 (ancestral segments XXI – XXVIII); C, antennule, segment 24 (ancestral segment XXVII – XXVIII); D, antenna; E, mandible; F, mandible, gnathobase; G, mandible, gnathobase, cutting edge; H, maxillule; I, maxillule, praecoxal arthrite. D, E, H, holotype; A – C, F – G, I, paratype; G, for smaller size (3.35 mm) paratype. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/201067/files/figure.png" pageId="7" pageNumber="56">Fig. 6</figureCitation>
|
|||
|
|
H–I), praecoxal arthrite with 9 terminal, 4 posterior, and 1 anterior setae; coxal endite with 2 setae; coxal epipodite with 9 setae; proximal basal endite with 3 setae, distal basal endite with 3 setae; endopod with 9 setae; exopod with 8 setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB03334F1C5C647" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
Maxilla (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AC5E3334F17EC528" box="[299,366,1333,1357]" captionStart="FIGURE 7" captionStartId="10.[151,249,1884,1907]" captionTargetBox="[162,1421,234,1829]" captionTargetId="figure@10.[151,1436,193,1863]" captionTargetPageId="10" captionText="FIGURE 7. Byrathis penicillatus sp. nov. Female. A, maxilla (sensory setae are not figured); B, maxilla, sensory setae on endopod; C, maxilla, sensory setae on endopod, worm-like setae are not figured); D, maxilliped; E, P 5. A, D, holotype; B – C, E, paratype; C, for smaller size (3.35 mm) paratype. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/201068/files/figure.png" pageId="7" pageNumber="56">Fig. 7</figureCitation>
|
|||
|
|
A–C), praecoxal endite (previously considered as proximal praecoxal endite) with 4 setae and a short attenuation; coxal endite (previously considered as distal praecoxal endite) with 3 setae; basal endites (previously considered as coxal endites) with 3 setae each, 1 seta on the proximal basal endite very short, 1 seta on distal basal endite very strong, spine-like, enditic-like lobe of proximal endopodal segment (previously considered as proximal basal endite) with 4 setae, 1 is thicker and 1 is sensory; all endites with a patch of long spinules at the base of the setae; endopod with 3 worm-like and 5 brush-like sensory setae: 2 setae shorter with large brushes, 3 setae longer with smaller brushes.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB0302CF26CC6B6" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
Maxilliped (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AC25302CF184C620" box="[336,404,1581,1605]" captionStart="FIGURE 7" captionStartId="10.[151,249,1884,1907]" captionTargetBox="[162,1421,234,1829]" captionTargetId="figure@10.[151,1436,193,1863]" captionTargetPageId="10" captionText="FIGURE 7. Byrathis penicillatus sp. nov. Female. A, maxilla (sensory setae are not figured); B, maxilla, sensory setae on endopod; C, maxilla, sensory setae on endopod, worm-like setae are not figured); D, maxilliped; E, P 5. A, D, holotype; B – C, E, paratype; C, for smaller size (3.35 mm) paratype. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/201068/files/figure.png" pageId="7" pageNumber="56">Fig. 7</figureCitation>
|
|||
|
|
D), syncoxa with 1 seta and a row of spinules on proximal praecoxal endite, 2 setae (1 sensory in distal part) and a patch of spinules on the middle endite and 2 sclerotized and 1 large brush-like seta on the distal praecoxal endite; coxal endite with 3 setae. Basis with 3 setae, proximal row of long spinules extended to the proximal basal seta and row of small spinules extended from the proximal to the distal-most basal seta. Endopod 6- segmented with 2, 4, 4, 3, 3+1, and 4 setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFE7FFA3ADB030DEF1AAC78A" blockId="7.[151,1437,253,2031]" pageId="7" pageNumber="56">
|
|||
|
|
Swimming legs. P1 (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3ACD830DEF1FDC692" box="[429,493,1759,1783]" captionStart="FIGURE 8" captionStartId="11.[151,249,1453,1476]" captionTargetBox="[193,1387,190,1436]" captionTargetId="figure@11.[170,1417,167,1436]" captionTargetPageId="11" captionText="FIGURE 8. Byrathis penicillatus sp. nov. Female, holotype. A, P 1; B, P 2; C, P 3; D, P 4. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/201069/files/figure.png" pageId="7" pageNumber="56">Fig. 8</figureCitation>
|
|||
|
|
A), coxa with anterior row of spinules along the distal margin; basis with medial distal seta strongly curved with setules; endopod 1-segmented with lateral lobe, its lateral margin with spinules; exopod segments 1 to 3 with 1 lateral spine each, spine of segment 1 reaching base of following spine; spine of exopod segment 2 not reaching the base of distal-most spine. P2–P4 (
|
|||
|
|
<figureCitation id="CEBB8389FFE7FFA3AE113148F3B8C704" box="[868,936,1865,1889]" captionStart="FIGURE 8" captionStartId="11.[151,249,1453,1476]" captionTargetBox="[193,1387,190,1436]" captionTargetId="figure@11.[170,1417,167,1436]" captionTargetPageId="11" captionText="FIGURE 8. Byrathis penicillatus sp. nov. Female, holotype. A, P 1; B, P 2; C, P 3; D, P 4. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/201069/files/figure.png" pageId="7" pageNumber="56">Fig. 8</figureCitation>
|
|||
|
|
B–D), coxa with 1 seta; basis without seta; endopod 2-segmented in P2, second segment on posterior surface with patch of long spinules; exopods 3-segmented. Endopods 3-segmented in P3–P4, segment 2 with a patch of long spinules on the posterior surface, segment 3 posterior surface densely spinulate, although with shorter spinules. Posterior surface of P4 coxa, basis and exopod densely spinulate.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="02FFCF84FFE8FFACADE2317AF0FCC788" httpUri="https://zenodo.org/record/201066/files/figure.png" pageId="8" pageNumber="57" targetBox="[213,1372,208,1875]" targetPageId="8">
|
|||
|
|
<paragraph id="563F9F0CFFE8FFACADE2317AF0FCC788" blockId="8.[151,1436,1915,2029]" pageId="8" pageNumber="57">
|
|||
|
|
<emphasis id="64F4431EFFE8FFACADE2317AF11CC7F4" bold="true" box="[151,268,1915,1938]" pageId="8" pageNumber="57">FIGURE 5</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="9180E48FFFE8FFACAC6D317AF1F9C7F4" box="[280,489,1915,1937]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFE8FFACAC6D317AF1F9C7F4" box="[280,489,1915,1937]" italics="true" pageId="8" pageNumber="57">Byrathis penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="64F4431EFFE8FFACAC85317DF22FC7F4" bold="true" box="[496,575,1916,1937]" pageId="8" pageNumber="57">
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFE8FFACAC85317DF22FC7F4" box="[496,575,1916,1937]" pageId="8" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Female. A, habitus, lateral view; B, habitus, dorsal view; C, rostrum, lateral view; D, rostrum, ventral view; E, posterior prosome and urosome, dorsal view; F, posterior prosome and urosome, lateral view; G, posterior prosome and genital double somite, lateral view. A–B, D, E–F, holotype; C, G, paratype. Scale bars: A, B, 0.5 mm, C–F, 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="02FFCF84FFE9FFADADE2315AF0D5C789" httpUri="https://zenodo.org/record/201067/files/figure.png" pageId="9" pageNumber="58" targetBox="[189,1419,195,1855]" targetPageId="9">
|
|||
|
|
<paragraph id="563F9F0CFFE9FFADADE2315AF0D5C789" blockId="9.[151,1436,1883,2028]" pageId="9" pageNumber="58">
|
|||
|
|
<emphasis id="64F4431EFFE9FFADADE2315AF11EC714" bold="true" box="[151,270,1883,1906]" pageId="9" pageNumber="58">FIGURE 6</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="9180E48FFFE9FFADAC68315DF1E2C717" box="[285,498,1884,1906]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFE9FFADAC68315DF1E2C717" box="[285,498,1884,1906]" italics="true" pageId="9" pageNumber="58">Byrathis penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="64F4431EFFE9FFADAC8E315DF25DC714" bold="true" box="[507,589,1884,1905]" pageId="9" pageNumber="58">
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFE9FFADAC8E315DF25DC714" box="[507,589,1884,1905]" pageId="9" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Female. A, antennule, articulating segments 1–17 (ancestral segments I–XX); B, antennule, articulating segments 18–24 (ancestral segments XXI–XXVIII); C, antennule, segment 24 (ancestral segment XXVII–XXVIII); D, antenna; E, mandible; F, mandible, gnathobase; G, mandible, gnathobase, cutting edge; H, maxillule; I, maxillule, praecoxal arthrite. D, E, H, holotype; A–C, F–G, I, paratype; G, for smaller size (3.35 mm) paratype. Scale bars: 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="02FFCF84FFEAFFAEADE2315DF349C7D5" httpUri="https://zenodo.org/record/201068/files/figure.png" pageId="10" pageNumber="59" targetBox="[162,1421,234,1829]" targetPageId="10">
|
|||
|
|
<paragraph id="563F9F0CFFEAFFAEADE2315DF349C7D5" blockId="10.[151,1436,1884,1968]" pageId="10" pageNumber="59">
|
|||
|
|
<emphasis id="64F4431EFFEAFFAEADE2315DF11EC717" bold="true" box="[151,270,1884,1907]" pageId="10" pageNumber="59">FIGURE 7</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="9180E48FFFEAFFAEAC6B315CF1E3C716" box="[286,499,1885,1907]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFEAFFAEAC6B315CF1E3C716" box="[286,499,1885,1907]" italics="true" pageId="10" pageNumber="59">Byrathis penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="64F4431EFFEAFFAEAC89315CF25FC717" bold="true" box="[508,591,1885,1906]" pageId="10" pageNumber="59">
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFEAFFAEAC89315CF25FC717" box="[508,591,1885,1906]" pageId="10" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Female. A, maxilla (sensory setae are not figured); B, maxilla, sensory setae on endopod; C, maxilla, sensory setae on endopod, worm-like setae are not figured); D, maxilliped; E, P5. A, D, holotype; B–C, E, paratype; C, for smaller size (3.35 mm) paratype. Scale bars 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="02FFCF84FFEBFFAFADE233ACF4F0C5A6" box="[151,1248,1453,1476]" httpUri="https://zenodo.org/record/201069/files/figure.png" pageId="11" pageNumber="60" targetBox="[193,1387,190,1436]" targetPageId="11">
|
|||
|
|
<paragraph id="563F9F0CFFEBFFAFADE233ACF4F0C5A6" blockId="11.[151,1248,1453,1476]" box="[151,1248,1453,1476]" pageId="11" pageNumber="60">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFADE233ACF11CC5A6" bold="true" box="[151,268,1453,1476]" pageId="11" pageNumber="60">FIGURE 8</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="9180E48FFFEBFFAFAC6C33AFF1FBC5A1" box="[281,491,1454,1476]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAC6C33AFF1FBC5A1" box="[281,491,1454,1476]" italics="true" pageId="11" pageNumber="60">Byrathis penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFEBFFAFAC8433AFF251C5A6" box="[497,577,1454,1475]" pageId="11" pageNumber="60" rank="species">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAC8433AFF251C5A6" bold="true" box="[497,577,1454,1475]" pageId="11" pageNumber="60">sp. nov.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
Female, holotype. A, P1; B, P2; C, P3; D, P4. Scale bars 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="563F9F0CFFEBFFAFADB033EEF201C62A" blockId="11.[151,1436,1519,2012]" pageId="11" pageNumber="60">
|
|||
|
|
P5 (
|
|||
|
|
<figureCitation id="CEBB8389FFEBFFAFAD8733EEF124C662" box="[242,308,1519,1543]" captionStart="FIGURE 7" captionStartId="10.[151,249,1884,1907]" captionTargetBox="[162,1421,234,1829]" captionTargetId="figure@10.[151,1436,193,1863]" captionTargetPageId="10" captionText="FIGURE 7. Byrathis penicillatus sp. nov. Female. A, maxilla (sensory setae are not figured); B, maxilla, sensory setae on endopod; C, maxilla, sensory setae on endopod, worm-like setae are not figured); D, maxilliped; E, P 5. A, D, holotype; B – C, E, paratype; C, for smaller size (3.35 mm) paratype. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/201068/files/figure.png" pageId="11" pageNumber="60">Fig. 7</figureCitation>
|
|||
|
|
E) 3-segmented, coxa and basis of equal length; coxa and basis with a patch of minor spinules distolaterally; exopod ornamented with surface spinules, 4 distal spines, medial terminal spine is curved, about 1.7 times longer than lateral terminal spine.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1E9ACC87FFEBFFAFADB2305DF106C6F2" pageId="11" pageNumber="60" type="etymology">
|
|||
|
|
<paragraph id="563F9F0CFFEBFFAFADB2305DF106C6F2" blockId="11.[151,1436,1519,2012]" pageId="11" pageNumber="60">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFADB2305DF15EC611" bold="true" box="[199,334,1628,1652]" pageId="11" pageNumber="60">Etymology.</emphasis>
|
|||
|
|
The species name “
|
|||
|
|
<taxonomicName id="9180E48FFFEBFFAFAF4C305DF2ACC611" box="[569,700,1628,1652]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="penicillatus">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAF4C305DF2ACC611" box="[569,700,1628,1652]" italics="true" pageId="11" pageNumber="60">penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
” refers to a large brush-like sensory seta on the syncoxa of the maxilliped.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1E9ACC87FFEBFFA8ADB030A5F167C15A" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60" type="discussion">
|
|||
|
|
<paragraph id="563F9F0CFFEBFFAFADB030A5F2F8C7F1" blockId="11.[151,1436,1519,2012]" pageId="11" pageNumber="60">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFADB030A5F129C6D9" bold="true" box="[197,313,1700,1724]" pageId="11" pageNumber="60">Remarks.</emphasis>
|
|||
|
|
<taxonomicName id="9180E48FFFEBFFAFAC4A30A4F1F1C6D9" box="[319,481,1700,1724]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAC4A30A4F1F1C6D9" box="[319,481,1700,1724]" italics="true" pageId="11" pageNumber="60">B. penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAC9230A5F250C6D9" bold="true" box="[487,576,1700,1724]" pageId="11" pageNumber="60">
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFEBFFAFAC9230A5F250C6D9" box="[487,576,1700,1724]" pageId="11" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is more closely related to its most geographically distant congener
|
|||
|
|
<taxonomicName id="9180E48FFFEBFFAFA85A30A4F0C2C685" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="laptevorum">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFA85A30A4F0C2C685" italics="true" pageId="11" pageNumber="60">B. laptevorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="CEBB8389FFEBFFAFAD9130C6F13AC6BA" box="[228,298,1735,1759]" captionStart="FIGURE 9" captionStartId="12.[151,249,1440,1463]" captionTargetBox="[196,1396,368,1418]" captionTargetId="figure@12.[188,1398,347,1419]" captionTargetPageId="12" captionText="FIGURE 9. Distribution of species of Byrathis in the World Ocean: black rhomboid B. macrocephalon (Grice & Hulsemann, 1970); circle outlined, B. penicillatus sp. nov.; black square, B. laurenae Markhaseva & Ferrari, 2005; big triangle outlined, B. arnei Schulz, 2006; black small triangle, B. divae sp. nov.; rhomboid outlined, B. laptevorum (Markhaseva, 1998); rhomboid outlined with crest, B. volcani Markhaseva & Ferrari, 2005." httpUri="https://zenodo.org/record/201070/files/figure.png" pageId="11" pageNumber="60">Fig. 9</figureCitation>
|
|||
|
|
) and shares with this species the following distinguishing characters: i) maxilla distal basal endite (previously considered as distal coxal endite) and enditic-like lobe of proximal endopodal segment (previously considered as proximal basal endite), each with 1 very strong, spine-like setal element (
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFA91B3110F49DC74D" box="[1134,1165,1809,1832]" italics="true" pageId="11" pageNumber="60">vs.</emphasis>
|
|||
|
|
these setal elements are neither strong nor spine-like in congeners); ii) each maxilla endite with a patch of long surface spinules (
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFA8533134F555C729" box="[1318,1349,1845,1868]" italics="true" pageId="11" pageNumber="60">vs.</emphasis>
|
|||
|
|
surface spinule patch is absent in the other species of the genus); iii) medial praecoxal endite of maxilliped syncoxa with a patch of spinules (
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAC10317CF193C7F1" box="[357,387,1917,1940]" italics="true" pageId="11" pageNumber="60">vs.</emphasis>
|
|||
|
|
no spinule patch in congeners).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFEBFFAFADB031A0F58CC7B9" blockId="11.[151,1436,1519,2012]" pageId="11" pageNumber="60">
|
|||
|
|
<taxonomicName id="9180E48FFFEBFFAFADB031A0F176C7DD" box="[197,358,1952,1976]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFADB031A0F176C7DD" box="[197,358,1952,1976]" italics="true" pageId="11" pageNumber="60">B. penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAC1831A1F1D6C7DD" bold="true" box="[365,454,1952,1976]" pageId="11" pageNumber="60">
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFEBFFAFAC1831A1F1D6C7DD" box="[365,454,1952,1976]" pageId="11" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="9180E48FFFEBFFAFAF2C31A0F2E7C7DD" box="[601,759,1952,1976]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="laptevorum">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAF2C31A0F2E7C7DD" box="[601,759,1952,1976]" italics="true" pageId="11" pageNumber="60">B. laptevorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in: i) large brush-like sensory seta on distal praecoxal endite (
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFADD531C4F0AFC7B9" box="[160,191,1989,2012]" italics="true" pageId="11" pageNumber="60">vs.</emphasis>
|
|||
|
|
poorly developed brush in
|
|||
|
|
<taxonomicName id="9180E48FFFEBFFAFAC8931C4F28DC7B9" box="[508,669,1988,2012]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="laptevorum">
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFAC8931C4F28DC7B9" box="[508,669,1988,2012]" italics="true" pageId="11" pageNumber="60">B. laptevorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
); ii) maxilliped basis with a patch of long spinules proximally (
|
|||
|
|
<emphasis id="64F4431EFFEBFFAFA80831C4F58CC7B9" box="[1405,1436,1989,2012]" italics="true" pageId="11" pageNumber="60">vs.</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="563F9F0CFFECFFA8ADE23696F167C15A" blockId="12.[151,1436,151,320]" pageId="12" pageNumber="61">
|
|||
|
|
spinules absent in
|
|||
|
|
<taxonomicName id="9180E48FFFECFFA8AC133698F215C0D5" box="[358,517,152,176]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="laptevorum">
|
|||
|
|
<emphasis id="64F4431EFFECFFA8AC133698F215C0D5" box="[358,517,152,176]" italics="true" pageId="12" pageNumber="61">B. laptevorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), and iii) P5 medial terminal spine curved, about 1.7 times longer than lateral terminal spine (
|
|||
|
|
<emphasis id="64F4431EFFECFFA8AC5C36BCF157C0B1" box="[297,327,189,212]" italics="true" pageId="12" pageNumber="61">vs.</emphasis>
|
|||
|
|
straight spine, nearly twice as long as lateral terminal spine in
|
|||
|
|
<taxonomicName id="9180E48FFFECFFA8A97C36BCF4B8C0B1" box="[1033,1192,188,212]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="laptevorum">
|
|||
|
|
<emphasis id="64F4431EFFECFFA8A97C36BCF4B8C0B1" box="[1033,1192,188,212]" italics="true" pageId="12" pageNumber="61">B. laptevorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
). Maxilla endopod of single known specimen of
|
|||
|
|
<taxonomicName id="9180E48FFFECFFA8ACB736E0F271C09D" box="[450,609,224,248]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="laptevorum">
|
|||
|
|
<emphasis id="64F4431EFFECFFA8ACB736E0F271C09D" box="[450,609,224,248]" italics="true" pageId="12" pageNumber="61">B. laptevorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is described as bearing 1 worm-like sensory seta (
|
|||
|
|
<emphasis id="64F4431EFFECFFA8A9E036E0F4A3C09D" box="[1173,1203,225,248]" italics="true" pageId="12" pageNumber="61">vs.</emphasis>
|
|||
|
|
3 brush-like setae in
|
|||
|
|
<taxonomicName id="9180E48FFFECFFA8ADE23704F129C179" box="[151,313,260,284]" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="penicillatus" status="sp. nov.">
|
|||
|
|
<emphasis id="64F4431EFFECFFA8ADE23704F129C179" box="[151,313,260,284]" italics="true" pageId="12" pageNumber="61">B. penicillatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="64F4431EFFECFFA8AC353705F189C179" bold="true" box="[320,409,260,284]" pageId="12" pageNumber="61">
|
|||
|
|
<taxonomicNameLabel id="7FC7FE65FFECFFA8AC353705F189C179" box="[320,409,260,284]" pageId="12" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
), but this character should be re-examined for the species when new specimens of
|
|||
|
|
<taxonomicName id="9180E48FFFECFFA8A84E3704F0CFC125" class="Maxillopoda" family="Scolecitrichidae" genus="Byrathis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Calanoida" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="laptevorum">
|
|||
|
|
<emphasis id="64F4431EFFECFFA8A84E3704F0CFC125" italics="true" pageId="12" pageNumber="61">B. laptevorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are obtained.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|