<documentid="8280D0343C1C769530742E1B98526B7B"ID-DOI="10.26028/cybium/2017-411-005"ID-ISSN="2101-0315"ID-Zenodo-Dep="13485593"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="guilherme"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="guilherme"checkinTime="1723043260833"checkinUser="felipe"docAuthor="Keith, Philippe, Lord, Clara & Larson, Helen K."docDate="2017"docId="2F75C925CE49FB26FEDC2F2329E8C9F3"docLanguage="en"docName="Cybium.41.1.45-66.pdf"docOrigin="Cybium 41 (1)"docSource="http://dx.doi.org/10.26028/cybium/2017-411-005"docStyle="DocumentStyle:73DAD84611431DC6606F8B63B2C6039C.1:Cybium.2013-2018.journal_article.type1"docStyleId="73DAD84611431DC6606F8B63B2C6039C"docStyleName="Cybium.2013-2018.journal_article.type1"docStyleVersion="1"docTitle="Schismatogobius tiola Keith, Lord & Larson, 2017, new species"docType="treatment"docVersion="3"lastPageNumber="57"masterDocId="D34CB15DCE4EFB2AFFD92A3A2A4DC802"masterDocTitle="Review of Schismatogobius (Gobiidae) from Papua New Guinea to Samoa, with description of seven new species"masterLastPageNumber="66"masterPageNumber="45"pageNumber="52"updateTime="1724938464977"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="EFAF2AED3379FAB88B20BB866BC05559">Review of Schismatogobius (Gobiidae) from Papua New Guinea to Samoa, with description of seven new species</mods:title>
<taxonomicNameid="60DC03B0CE49FB2DFEDC2F232BBACD33"ID-CoL="6XW3R"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[261,503,1305,1329]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="7"pageNumber="52"phylum="Chordata"rank="species"species="tiola"status="sp. nov.">
<figureCitationid="3FE764B6CE49FB2DFEE62F032BE6CD53"box="[319,427,1337,1361]"captionStart-0="Figure 1"captionStart-1="Figure 2"captionStartId-0="3.[132,196,1837,1859]"captionStartId-1="4.[699,763,1861,1883]"captionTargetBox-0="[173,1407,922,1824]"captionTargetBox-1="[132,1455,1176,1907]"captionTargetId-0="figure-298@3.[169,1418,921,1827]"captionTargetId-1="figure-740@4.[699,1455,1176,1844]"captionTargetPageId-0="3"captionTargetPageId-1="4"captionText-0="Figure 1. – Bayesian phylogenetic reconstruction, based on the partial COI barcode region (636 bp). The posterior probability values are given at each node. Following each species name is the country the specimen comes from; the numbers following the species are the numbers given on the field."captionText-1="Figure 2. – A-F: Schematic drawings of the ventral surface of head of Schis- matogobius species from Papua New Guinea to Samoa. A: 13-14 pectoral rays."figureDoi-0="http://doi.org/10.5281/zenodo.13485601"figureDoi-1="http://doi.org/10.5281/zenodo.13485609"httpUri-0="https://zenodo.org/record/13485601/files/figure.png"httpUri-1="https://zenodo.org/record/13485609/files/figure.png"pageId="7"pageNumber="52">Figs 1-2C</figureCitation>
<figureCitationid="3FE764B6CE49FB2DFE6F2F032B91CD53"box="[438,476,1337,1361]"captionStart-0="Figure 4"captionStart-1="Figure 5"captionStartId-0="10.[132,195,966,988]"captionStartId-1="11.[132,195,1022,1044]"captionTargetBox-0="[132,1455,215,949]"captionTargetBox-1="[132,775,215,1005]"captionTargetId-0="figure-279@10.[132,1455,215,949]"captionTargetId-1="figure-646@11.[132,775,488,747]"captionTargetPageId-0="10"captionTargetPageId-1="11"captionText-0="Figure 4. – Distribution area of Schismatogobius species from Papua New Guinea to Samoa."captionText-1="Figure 5. – A: Schismatogobius tiola n. sp., male, 23.1 mm SL; B: Female, 30 mm SL (Photos C. Lord). C: Schismatogobius tiola, holotype, MNHN 2016-0265, male 25.5 mm SL (Photo P. Keith)."figureDoi-0="http://doi.org/10.5281/zenodo.13485625"figureDoi-1="http://doi.org/10.5281/zenodo.13485639"httpUri-0="https://zenodo.org/record/13485625/files/figure.png"httpUri-1="https://zenodo.org/record/13485639/files/figure.png"pageId="7"pageNumber="52">4-5</figureCitation>
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<specimenCodeid="F77AD048CE49FB2DFEE12F80285ECDD3"box="[312,531,1465,1489]"collectionCode="MNHN"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="7"pageNumber="52">MNHN 2016-0265</specimenCode>
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<specimenCodeid="F77AD048CE49FB2DFE942C00287CCE53"box="[333,561,1593,1617]"collectionCode="MNHN"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="7"pageNumber="52">MNHN 2016-0266</specimenCode>
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<specimenCodeid="F77AD048CE49FB2DFDF62CE0294FCEF3"box="[559,770,1753,1777]"collectionCode="MNHN"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="7"pageNumber="52">MNHN 2016-0292</specimenCode>
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<specimenCodeid="F77AD048CE49FB2DFEEA2D00284CCF53"box="[307,513,1849,1873]"collectionCode="MNHN"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="7"pageNumber="52">MNHN 2016-0294</specimenCode>
<taxonomicNameid="60DC03B0CE49FB2DFF5D2D632BD1CF70"authority="River, Ranongga"authorityName="River, Ranongga"box="[132,412,1881,1906]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="7"pageNumber="52"phylum="Chordata"rank="species"species="mondo">Mondo River, Ranongga</taxonomicName>
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<paragraphid="A7637833CE49FB2DFC8A2BA32E7FCA93"blockId="7.[813,1456,377,658]"pageId="7"pageNumber="52">Usually 15 pectoral rays; pectoral fins with a large triangular dorsal (or transverse) black band and a smaller one posteriorly. First dorsal fin membrane posterior to spine 6 not connected to base of spine of second dorsal fin. Anal fin I,8-9. Ventral surface of head in male blackish with a white mentum. Ventral surface of head in female whitish or slightly pigmented with dusky blotches and with usually a blackish ring around mentum.</paragraph>
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SL). Body naked, slender, almost circular in cross-section. Head rounded, snout rather pointed, cheeks bulbous in male. Mouth oblique, tip of lower lip anteriormost. Jaw lengths in males much greater than in females; jaw length 48-55% of HL in male and 28-35% of HL in females. Lower jaw reaching vertical of 1/3 to 1/2 of the eye in female and exceeding (for half eye diameter) a vertical of posterior margin of eye in male. Eyes high on head, close together with interorbital width about equal to half eye diameter. Anterior nostril short and tube-like.
Dorsal fins VI-I,9, membrane in first dorsal fin posterior to spine 6 not connected to base of spine of second dorsal fin. D1 with all spines about equal in length. Anal fin I,8(2)- I,9(5), slightly behind or approximately below second dorsal fin origin. Caudal fin with 11-12 branched rays, posterior margin rounded. Pectoral fins oblong with posterior margin pointed and 14(1)-15(6) rays (
<tableCitationid="EA5E4D88CE49FB2DFB0D2EC02F59CD10"box="[1236,1300,1274,1298]"captionStart="Table I"captionStartId="6.[813,866,1319,1341]"captionText="Table I. – Number of pectoral rays of Schismatogobius species from Papua New Guinea to Samoa"pageId="7"pageNumber="52">Tab. I</tableCitation>
), ventralmost ray unbranched. Pelvic fins always I,5, with both fins joined together for their entire length between fifth rays to form a strong cup-like disc and a well developed and lobed frenum between spines, fins not extending beyond anus. Morphomeristic data given in table III.
</paragraph>
<paragraphid="A7637833CE49FB2DFC8A2F812F1ACE50"blockId="7.[813,1456,697,1907]"pageId="7"pageNumber="52">Tongue (anterior tip) bilobed. Teeth in upper jaw (15- 20) usually in two rows, teeth conical and slightly recurved. Teeth in lower jaw (6-13) usually in one or two rows anteriorly and single row laterally, all teeth conical with outer row teeth only slightly enlarged and somewhat recurved.</paragraph>
Cephalic sensory pore system always with pores B, D, F, K, L, N and O, pore D singular with all other pores paired; oculoscapular canal absent between pores F and K. Anterior interorbital extension of anterior oculoscapular canal with double terminal pores B slightly posterior to posterior nostril. D pore at rear of interorbital. Posterior extension of anterior oculoscapular canal terminating laterally on each side of head at pore F, just behind posterior edge of eye. Posterior oculoscapular canal with 2 terminal pores, K and L; preopercular canal with 2 pores, N and O. Cutaneous sensory
<figureCitationid="3FE764B6CE46FB22FB5C283E2E9BCA1E"box="[1157,1238,516,540]"captionStart="Figure 5"captionStartId="11.[132,195,1022,1044]"captionTargetBox="[132,775,215,1005]"captionTargetId="figure-646@11.[132,775,488,747]"captionTargetPageId="11"captionText="Figure 5. – A: Schismatogobius tiola n. sp., male, 23.1 mm SL; B: Female, 30 mm SL (Photos C. Lord). C: Schismatogobius tiola, holotype, MNHN 2016-0265, male 25.5 mm SL (Photo P. Keith)."figureDoi="http://doi.org/10.5281/zenodo.13485639"httpUri="https://zenodo.org/record/13485639/files/figure.png"pageId="8"pageNumber="53">Fig. 5C</figureCitation>
(a whitish pigmented Fig more. 2C. or First), or less usually mostly dorsal white with fin brown isthmus with blackish with large; belly a ring black white around whitish median mentum mentum; breast band and paired 7 6.76 6.92 7.08 6.92
and with black a black margin median. Second band. Caudal dorsal fin fin black mostly and cream white,
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vic with black white pigmented disc black. spots mostly. blotch Base posteriorly whitish of at pectoral centre. over Anal of rays fin hypural fin white, mostly sometimes crease and cream distal and slightly. Pelpart two - between 4 3
band black female with band. In male. a Between bright, presence blue vertical of border bright black, both blue bands spots in, male orange on each and to of 1 0.00 0.31 0.31
rose rose with with many many large small blue blue spots spots.. Male Female head head orange darker to percentage)) Caledonia and Lower mottled half of, mostly the body black in lateral at nape view and between mottled, eyes with.
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<taxonomicNameid="60DC03B0CE46FB22FD252C1A295ECE45"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[764,787,1568,1607]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="8"pageNumber="53"phylum="Chordata"rank="species"species="tiola">
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from black superior, with and half reddish of the body blotches. First, clearly dorsal fin different trans- distance 06948 Solomon 11939 Solomon Solomon 12065 11932 Solomon
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gin dark lucent. Second spots with on large dorsal rays black. fin Caudal translucent median fin marbled band with and or rows barred black of small mar- with
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and black lucent two and. Pelvic large white white disc, with mostly spots black posteriorly whitish spot over at. hypural Anal rays fin. Frenum crease trans-. II
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<taxonomicNameid="60DC03B0CE47FB23FF23287F2B59CA39"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[250,276,510,596]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="hoesei">
<taxonomicNameid="60DC03B0CE47FB23FF23291C2B59CB1E"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[250,276,711,821]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="baitabag">
<taxonomicNameid="60DC03B0CE47FB23FF232E8D2B59CCAC"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[250,276,1162,1222]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="essi">
<taxonomicNameid="60DC03B0CE47FB23FF232FAA2B59CD85"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[250,276,1371,1439]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="tiola">
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<tdid="5A3D130DCE4704D5FEFE2CFC2936CF49"box="[295,891,1734,1867]"gridcol="1"gridrow="14"pageId="9"pageNumber="54">P D A PDL PAL HL male JL male JL / HL male HL female JL female JL / HL female CPD Pect L BDa SDFL AFL CFL</td>
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<taxonomicNameid="60DC03B0CE47FB23FC1C2AC32EF0C913"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[965,1213,249,273]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="tiola">
<paragraphid="A7637833CE47FB23FBAD2B232EC0C953"blockId="9.[927,1456,249,1875]"pageId="9"pageNumber="54">with moderate to fast flow in shallow areas of rocks</paragraph>
<taxonomicNameid="60DC03B0CE47FB23FC462B402E09C993"authorityName="Keith, Marquet & Watson"authorityYear="2004"box="[927,1092,378,401]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="vanuatuensis">
<paragraphid="A7637833CE47FB23FC1C28A02E36CAF0"blockId="9.[927,1456,249,1875]"pageId="9"pageNumber="54">In the story of Tiola, a stone dog protector of Nusa Roviana village, Tiola would turn its body to face the direction</paragraph>
<paragraphid="A7637833CE47FB23FC2928C02ED4CB10"blockId="9.[927,1456,249,1875]"box="[1008,1177,762,786]"pageId="9"pageNumber="54">in the direction</paragraph>
<paragraphid="A7637833CE47FB23FB6729802F1BCBD3"blockId="9.[927,1456,249,1875]"box="[1214,1366,954,978]"pageId="9"pageNumber="54">and place it on</paragraph>
<paragraphid="A7637833CE47FB23FA8229802EE5CBF3"blockId="9.[927,1456,249,1875]"pageId="9"pageNumber="54">the bow of the canoe. The dog sat</paragraph>
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differs from the other species sequenced and present in the area studied by having a high% of divergence in COI gene (10.9- 18.9%) (Tab. II) and from these species, except
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<taxonomicNameid="60DC03B0CE47FB23FC462F612E50CD70"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[927,1053,1370,1394]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="baitabag"status="n. sp.">
<figureCitationid="3FE764B6CE47FB23FC712C602E6ECE70"box="[936,1059,1626,1650]"captionStart="Figure 2"captionStartId="4.[699,763,1861,1883]"captionTargetBox="[132,1455,1176,1907]"captionTargetId="figure-740@4.[699,1455,1176,1844]"captionTargetPageId="4"captionText="Figure 2. – A-F: Schematic drawings of the ventral surface of head of Schis- matogobius species from Papua New Guinea to Samoa. A: 13-14 pectoral rays."figureDoi="http://doi.org/10.5281/zenodo.13485609"httpUri="https://zenodo.org/record/13485609/files/figure.png"pageId="9"pageNumber="54">Fig. 2B, C</figureCitation>
<taxonomicNameid="60DC03B0CE47FB23FB382C612F64CE70"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[1249,1321,1627,1650]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="essi">
<paragraphid="A7637833CE47FB23FBC12C412F33CE90"blockId="9.[927,1456,249,1875]"box="[1048,1406,1658,1683]"pageId="9"pageNumber="54">(average adult size more than 26</paragraph>
<paragraphid="A7637833CE47FB23FC382CA02ECCCEB0"blockId="9.[927,1456,249,1875]"box="[993,1153,1690,1714]"pageId="9"pageNumber="54">less), a smaller</paragraph>
<paragraphid="A7637833CE47FB23FB512CA02F7DCEB0"blockId="9.[927,1456,249,1875]"box="[1160,1328,1690,1714]"pageId="9"pageNumber="54">anal fin length (</paragraph>
Table; length mm and of pectoral fins. It differs from
<taxonomicNameid="60DC03B0CE47FB23FAF52D012FDECF51"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[1324,1427,1851,1875]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="9"pageNumber="54"phylum="Chordata"rank="species"species="mondo">
1 female (15.3 mm SL), Vage River, Kolombangara Island, frenum in male and female (Fig. 2B, C). It differs from
<taxonomicNameid="60DC03B0CE44FB20FF192E8D2B70CCCD"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[192,317,1207,1231]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="10"pageNumber="55"phylum="Chordata"rank="species"species="baitabag"status="n. sp.">
<paragraphid="A7637833CE44FB20FF5D2F2D28FCCD4D"blockId="10.[132,690,1303,1903]"pageId="10"pageNumber="55">SL), and a different colour pattern of ventral surface of head, pelvic disk and frenum in male and female</paragraph>
<taxonomicNameid="60DC03B0CE44FB20FB4D2F202EB6CD32"authorityName="Jenkins & Boseto"authorityYear="2005"box="[1172,1275,1306,1328]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="10"pageNumber="55"phylum="Chordata"rank="species"species="vitiensis">
<taxonomicNameid="60DC03B0CE44FB20FAD72F202FEECD32"authorityName="Keith, Marquet & Watson"authorityYear="2004"box="[1294,1443,1306,1328]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="10"pageNumber="55"phylum="Chordata"rank="species"species="vanuatuensis">
<figureCitationid="3FE764B6CE44FB20FF552F6D2AADCD6D"box="[140,224,1367,1391]"captionStart="Figure 2"captionStartId="4.[699,763,1861,1883]"captionTargetBox="[132,1455,1176,1907]"captionTargetId="figure-740@4.[699,1455,1176,1844]"captionTargetPageId="4"captionText="Figure 2. – A-F: Schematic drawings of the ventral surface of head of Schis- matogobius species from Papua New Guinea to Samoa. A: 13-14 pectoral rays."figureDoi="http://doi.org/10.5281/zenodo.13485609"httpUri="https://zenodo.org/record/13485609/files/figure.png"pageId="10"pageNumber="55">Fig. 2C</figureCitation>
<taxonomicNameid="60DC03B0CE44FB20FF732F8D2ABBCDCD"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[170,246,1463,1487]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="10"pageNumber="55"phylum="Chordata"rank="species"species="tiola">
<figureCitationid="3FE764B6CE44FB20FF552FED2A80CDED"box="[140,205,1495,1519]"captionStart="Figure 4"captionStartId="10.[132,195,966,988]"captionTargetBox="[132,1455,215,949]"captionTargetId="figure-279@10.[132,1455,215,949]"captionTargetPageId="10"captionText="Figure 4. – Distribution area of Schismatogobius species from Papua New Guinea to Samoa."figureDoi="http://doi.org/10.5281/zenodo.13485625"httpUri="https://zenodo.org/record/13485625/files/figure.png"pageId="10"pageNumber="55">Fig. 4</figureCitation>
<taxonomicNameid="60DC03B0CE44FB20FF3B2C0C2B86CE4D"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[226,459,1590,1615]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="10"pageNumber="55"phylum="Chordata"rank="species"species="essi"status="sp. nov.">
<figureCitationid="3FE764B6CE44FB20FECB2C6D2B33CE6D"box="[274,382,1623,1647]"captionStart-0="Figure 1"captionStart-1="Figure 2"captionStartId-0="3.[132,196,1837,1859]"captionStartId-1="4.[699,763,1861,1883]"captionTargetBox-0="[173,1407,922,1824]"captionTargetBox-1="[132,1455,1176,1907]"captionTargetId-0="figure-298@3.[169,1418,921,1827]"captionTargetId-1="figure-740@4.[699,1455,1176,1844]"captionTargetPageId-0="3"captionTargetPageId-1="4"captionText-0="Figure 1. – Bayesian phylogenetic reconstruction, based on the partial COI barcode region (636 bp). The posterior probability values are given at each node. Following each species name is the country the specimen comes from; the numbers following the species are the numbers given on the field."captionText-1="Figure 2. – A-F: Schematic drawings of the ventral surface of head of Schis- matogobius species from Papua New Guinea to Samoa. A: 13-14 pectoral rays."figureDoi-0="http://doi.org/10.5281/zenodo.13485601"figureDoi-1="http://doi.org/10.5281/zenodo.13485609"httpUri-0="https://zenodo.org/record/13485601/files/figure.png"httpUri-1="https://zenodo.org/record/13485609/files/figure.png"pageId="10"pageNumber="55">Figs 1-2B</figureCitation>
<figureCitationid="3FE764B6CE44FB20FE502C6D2BD5CE6D"box="[393,408,1623,1647]"captionStart="Figure 4"captionStartId="10.[132,195,966,988]"captionTargetBox="[132,1455,215,949]"captionTargetId="figure-279@10.[132,1455,215,949]"captionTargetPageId="10"captionText="Figure 4. – Distribution area of Schismatogobius species from Papua New Guinea to Samoa."figureDoi="http://doi.org/10.5281/zenodo.13485625"httpUri="https://zenodo.org/record/13485625/files/figure.png"pageId="10"pageNumber="55">4</figureCitation>
<specimenCodeid="F77AD048CE44FB20FEE92CCD284FCF0D"box="[304,514,1783,1807]"collectionCode="MNHN"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="10"pageNumber="55">MNHN 2016-0267</specimenCode>
<specimenCodeid="F77AD048CE45FB21FEE22EA12BBBCCB1"box="[315,502,1179,1203]"collectionCode="NTM"country="Australia"httpUri="http://grbio.org/cool/jg13-7fbc"name="Northern Territory Museum of Arts and Sciences"pageId="11"pageNumber="56">NTM 16447-002</specimenCode>
<specimenCodeid="F77AD048CE45FB21FED42F212BAECD31"box="[269,483,1307,1331]"collectionCode="MNHN"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="11"pageNumber="56">MNHN 2016-0296</specimenCode>
<taxonomicNameid="60DC03B0CE45FB21FC782BF22E38C9DC"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[929,1141,456,478]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="11"pageNumber="56"phylum="Chordata"rank="species"species="essi"status="sp. nov.">
<taxonomicNameid="60DC03B0CE45FB21FECB29C42BA7CC11"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[274,490,1022,1043]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="11"pageNumber="56"phylum="Chordata"rank="species"species="tiola"status="sp. nov.">
<taxonomicNameid="60DC03B0CE45FB21FDF82E2C294DCC29"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[545,768,1046,1067]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="11"pageNumber="56"phylum="Chordata"rank="species"species="tiola">
<paragraphid="A7637833CE45FB21FF732FE1280DCEF1"blockId="11.[132,775,1466,1779]"pageId="11"pageNumber="56">Usually 15 pectoral rays; pectoral fins with broad black triangular wedge dorsally usually with few rows of dark spots below it; dorsal fins VI-I,9, membrane in first dorsal fin posterior to spine 6 not connected to base of spine in second dorsal fin. Anal fin I,8-9. Ventral surface of head, frenum and pelvic disc in male entirely blackish or entirely brownish. In female, underside of head pale with chin and surrounding area blackish, with diffuse dark brown area on anterior part of branchiostegal membrane and isthmus.</paragraph>
<taxonomicNameid="60DC03B0CE45FB21FE9E2D01284FCF51"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[327,514,1851,1875]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="11"pageNumber="56"phylum="Chordata"rank="genus">
SL). Body naked, slender, almost circular in cross-section.
</paragraph>
<paragraphid="A7637833CE45FB21FCF42820292FCB30"blockId="11.[813,1456,538,1906]"pageId="11"pageNumber="56">Head rounded, snout rather pointed, cheeks bulbous in males. Mouth oblique, tip of lower lip anteriormost. Jaw lengths in males much greater than in females; jaw length 48-57% of HL in males and 31-34% of HL in females. Lower jaw reaching a vertical of anterior third of eye in female and reaching or exceed (for third of eye diameter) a vertical of posterior part of eye in male. Eyes high on head, close together with narrow interorbital, 9.2% of HL. Anterior nostril with short tube.</paragraph>
Dorsal fins VI-I,9, membrane in first dorsal fin posterior to spine 6 not connected to base of spine of second dorsal fin. D1 with all spines about equal in length. Anal fin I,8(2)- I,9(7), slightly behind or approximately below second dorsal fin origin. Caudal fin with 11-12 branched rays, posterior margin rounded. Pectoral fins oblong with posterior margin pointed and 14(2)-15(8)-16(1) rays (
<tableCitationid="EA5E4D88CE45FB21FB6D29C02EBFCC10"box="[1204,1266,1018,1042]"captionStart="Table I"captionStartId="6.[813,866,1319,1341]"captionText="Table I. – Number of pectoral rays of Schismatogobius species from Papua New Guinea to Samoa"pageId="11"pageNumber="56">Tab. I</tableCitation>
), ventralmost ray unbranched. Pelvic fins always I,5, with both fins joined together their entire length between rays 5 to form strong cup-like disc; a well developed frenum (small pointed lobe at each side) between spines, fins not extending beyond anus. Morphomeristic data given in table III.
</paragraph>
<paragraphid="A7637833CE45FB21FC8B2E8129DCCD70"blockId="11.[813,1456,538,1906]"pageId="11"pageNumber="56">Tongue (anterior tip) bilobed, not deeply. Teeth in upper jaw (12-23) usually in two rows, teeth conical and slightly recurved. Teeth in lower jaw (10-12) usually in one or two rows anteriorly and single row laterally, all teeth conical with outer row teeth only slightly enlarged and somewhat recurved.</paragraph>
<paragraphid="A7637833CE45FB21FC8A2F4029F5CEF0"blockId="11.[813,1456,538,1906]"pageId="11"pageNumber="56">Cephalic sensory pore system always with pores B, D, F, K, L, N and O, pore D singular with all other pores paired; oculoscapular canal absent between pores F and K. Anterior interorbital extension of anterior oculoscapular canal with double terminal pores B slightly posterior to posterior nostril. D pore at rear of interorbital. Posterior extension of anterior oculoscapular canal terminating laterally on each side of head at pore F, just behind posterior edge of eye. Posterior oculoscapular canal with 2 terminal pores, K and L; preopercular canal with 2 pores, N and O. Cutaneous sensory papillae not well developed and inconspicuous due to preservation.</paragraph>
<paragraphid="A7637833CE45FB21FC8A2CC02FEBCF70"blockId="11.[813,1456,538,1906]"pageId="11"pageNumber="56">Sexual dimorphism fairly well developed with males always having jaws longer than females and a different colour pattern on the ventral surface of head. Urogenital papilla broadly rounded in females and slightly triangular in males.</paragraph>
<taxonomicNameid="60DC03B0CE42FB26FEC029342840CB21"authorityName="Keith & Lord & Larson"authorityYear="2017"box="[281,525,782,803]"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="12"pageNumber="57"phylum="Chordata"rank="species"species="mondo"status="sp. nov.">
<taxonomicNameid="60DC03B0CE42FB26FD5D291C2ABECB51"authorityName="Keith & Lord & Larson"authorityYear="2017"family="Gobiidae"genus="Schismatogobius"kingdom="Animalia"order="Perciformes"pageId="12"pageNumber="57"phylum="Chordata"rank="species"species="mondo"status="sp. nov.">
Usually four vertical black bands over pale yellowish body in dorsal view; first band below first dorsal fin, second and third bands below the second dorsal fin (may coalesce to form single broad band, slightly paler toward centre) and fourth band at base of caudal fin. Side of body, between dark bands, with fine brown mottling and speckles, may form vermiculate patterns. Head slightly darker than body, side of head with darker brown mottling; indistinct darker brown bar from eye to jaw and another on opercle just behind rear preopercular margin; few small dark brown spots on middle of nape and another above corner of opercle. In male, ventral surface of head, frenum and pelvic disk entirely blackish or brownish (
<figureCitationid="3FE764B6CE42FB26FEE32F682BC2CD68"box="[314,399,1362,1386]"captionStart="Figure 2"captionStartId="4.[699,763,1861,1883]"captionTargetBox="[132,1455,1176,1907]"captionTargetId="figure-740@4.[699,1455,1176,1844]"captionTargetPageId="4"captionText="Figure 2. – A-F: Schematic drawings of the ventral surface of head of Schis- matogobius species from Papua New Guinea to Samoa. A: 13-14 pectoral rays."figureDoi="http://doi.org/10.5281/zenodo.13485609"httpUri="https://zenodo.org/record/13485609/files/figure.png"pageId="12"pageNumber="57">Fig. 2B</figureCitation>
). In female, underside of head pale with chin and surrounding area blackish, with diffuse dark brown area on anterior part of branchiostegal membrane and isthmus, most of isthmus and breast unpigmented (
<figureCitationid="3FE764B6CE42FB26FD7F2F8028B7CDD0"box="[678,762,1466,1490]"captionStart="Figure 2"captionStartId="4.[699,763,1861,1883]"captionTargetBox="[132,1455,1176,1907]"captionTargetId="figure-740@4.[699,1455,1176,1844]"captionTargetPageId="4"captionText="Figure 2. – A-F: Schematic drawings of the ventral surface of head of Schis- matogobius species from Papua New Guinea to Samoa. A: 13-14 pectoral rays."figureDoi="http://doi.org/10.5281/zenodo.13485609"httpUri="https://zenodo.org/record/13485609/files/figure.png"pageId="12"pageNumber="57">Fig. 2B</figureCitation>
). Belly pale anteriorly; posterior half of belly with brown pigment where black body bar extends. First dorsal fin with black basal band and black margin. Second dorsal fin translucent with several vertical blackish streaks and spots along rays. Caudal fin black and white, with broad black blotch (continuation of last body bar) extending in irregular pattern onto fin and becoming dusky posteriorly, partly surrounding two whitish spots posteriorly. Anal fin translucent to cream. Pectoral fins with broad black triangular wedge, with narrow black bar (formed by row of spots) ventrally and second short row of black spots posteriorly; a dark spot is generally visible on the posterior upper part.
<emphasisid="95A8A421CE42FB26FCF42AE3298FC8F3"bold="true"box="[813,962,217,241]"pageId="12"pageNumber="57">Colour in life</emphasis>
</paragraph>
<paragraphid="A7637833CE42FB26FC8B2AC329E8C9F3"blockId="12.[813,1456,217,497]"pageId="12"pageNumber="57">Similar to that in preservation, but lateral and dorsal parts of body between vertical black bands reddish. First dorsal fin translucent with small black basal band and black margin. Second dorsal fin translucent with black stripes. Caudal fin black and white, with black blotch at hypural and two white spots posteriorly. Anal fin translucent. Pectoral fins with broad black triangular band or wedge commencing from dorsal part.</paragraph>
