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<document id="FCEE7547E705CE3FE4A662FC07B8543E" ID-DOI="10.5281/zenodo.200811" ID-GBIF-Dataset="76b2912f-8856-4c85-842d-c3312bc2650f" ID-ISSN="1175-5326" ID-Zenodo-Dep="200811" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460250427582" checkinUser="plazi" docAuthor="Mcdonald, Andrew T." docDate="2011" docId="E80287E9B119FFF9CDB95727FC10F979" docLanguage="en" docName="zt02783p068.pdf" docOrigin="Zootaxa 2783" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Osmakasaurus depressus Gilmore 1909, comb. nov." docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="143BFF91B110FFF5CD2E537DFFE3FF95" masterDocTitle="The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)" masterLastPageNumber="68" masterPageNumber="52" pageNumber="61" updateTime="1698242557538" updateUser="plazi">
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<mods:title id="418EB8730E7548BBF6212C5AA486C2CE">The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)</mods:title>
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<mods:namePart id="7727320D36B276EB0CFFE7E45C63C8C2">Mcdonald, Andrew T.</mods:namePart>
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<mods:date id="983FE1312F7650BA2A72A4DD04B9C767">2011</mods:date>
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<mods:identifier id="A005D942734C78097E2A9EBF4484E271" type="DOI">10.5281/zenodo.200811</mods:identifier>
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<treatment id="E80287E9B119FFF9CDB95727FC10F979" ID-DOI="http://doi.org/10.5281/zenodo.6185217" ID-GBIF-Taxon="119391377" ID-Zenodo-Dep="6185217" LSID="urn:lsid:plazi:treatment:E80287E9B119FFF9CDB95727FC10F979" httpUri="http://treatment.plazi.org/id/E80287E9B119FFF9CDB95727FC10F979" lastPageId="12" lastPageNumber="64" pageId="9" pageNumber="61">
<subSubSection id="28B16574B119FFFCCDB95727FD35FB63" pageId="9" pageNumber="61" type="nomenclature">
<paragraph id="601436FFB119FFFCCDB95727FD1AFBE6" blockId="9.[151,761,1114,1139]" box="[151,761,1114,1139]" pageId="9" pageNumber="61">
<heading id="3B5C8193B119FFFCCDB95727FD1AFBE6" bold="true" box="[151,761,1114,1139]" fontSize="10" level="2" pageId="9" pageNumber="61" reason="2">
<emphasis id="52DFEAEDB119FFFCCDB95727FD1AFBE6" bold="true" box="[151,761,1114,1139]" pageId="9" pageNumber="61">
<taxonomicName id="A7AB4D7CB119FFFCCDB95727FD88FBE6" authority="Gilmore, 1909" authorityName="Gilmore" authorityYear="1909" box="[151,619,1114,1139]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus" status="comb. nov.">
<emphasis id="52DFEAEDB119FFFCCDB95727FE59FBE6" bold="true" box="[151,442,1114,1139]" italics="true" pageId="9" pageNumber="61">Osmakasaurus depressus</emphasis>
<bibRefCitation id="043A4B0EB119FFFCCCEF5727FD88FBE6" author="Gilmore" box="[449,619,1114,1139]" pageId="9" pageNumber="61" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="49EC5796B119FFFCCF565726FD1AFBE6" box="[632,761,1115,1139]" pageId="9" pageNumber="61">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="601436FFB119FFFCCDB957DEFD75FB2D" blockId="9.[151,726,1186,1270]" box="[151,662,1186,1209]" pageId="9" pageNumber="61">
1909
<taxonomicName id="A7AB4D7CB119FFFCCDC557DEFDA6FB2D" authority="Gilmore" authorityName="Gilmore" box="[235,581,1186,1209]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCDC557DEFE0BFB2C" box="[235,488,1187,1209]" italics="true" pageId="9" pageNumber="61">Camptosaurus depressus</emphasis>
Gilmore
</taxonomicName>
, p. 292.
</paragraph>
<paragraph id="601436FFB119FFFCCDB957BFFD7CFB42" blockId="9.[151,726,1186,1270]" box="[151,671,1217,1239]" pageId="9" pageNumber="61">
2004?
<taxonomicName id="A7AB4D7CB119FFFCCDD857BFFDACFB42" authority="Norman" authorityName="Norman" box="[246,591,1217,1239]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCDD857BFFE10FB42" box="[246,499,1217,1239]" italics="true" pageId="9" pageNumber="61">Camptosaurus depressus</emphasis>
Norman
</taxonomicName>
, p. 415.
</paragraph>
<paragraph id="601436FFB119FFFCCDB9579DFD35FB63" blockId="9.[151,726,1186,1270]" box="[151,726,1248,1270]" pageId="9" pageNumber="61">
2008
<taxonomicName id="A7AB4D7CB119FFFCCDC5579DFD65FB63" authority="Carpenter &amp; Wilson" authorityName="Carpenter &amp; Wilson" box="[235,646,1248,1270]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressa">
<emphasis id="52DFEAEDB119FFFCCDC5579DFE52FB63" box="[235,433,1248,1270]" italics="true" pageId="9" pageNumber="61">Planicoxa depressa</emphasis>
Carpenter &amp; Wilson
</taxonomicName>
, p. 257.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B119FFFCCDB9565EFF3AF952" pageId="9" pageNumber="61" type="materials_examined">
<paragraph id="601436FFB119FFFCCDB9565EFD32FAAF" blockId="9.[151,1437,1314,1986]" box="[151,721,1314,1339]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDB9565EFEE8FAAE" bold="true" box="[151,267,1315,1339]" pageId="9" pageNumber="61">
<typeStatus id="BF10885DB119FFFCCDB9565EFEE5FAAE" box="[151,262,1315,1339]" pageId="9" pageNumber="61" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="06BAAE3AB119FFFCCC3C565FFE80FAAF" LSID="urn:lsid:biocol.org:col:34871" box="[274,355,1314,1338]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="61">USNM</collectionCode>
4753, fragmentary postcranium.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB563BFF2BFA5F" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB563BFE4FFACA" bold="true" box="[197,428,1350,1375]" pageId="9" pageNumber="61">Generic etymology.</emphasis>
The generic name is derived from
<emphasis id="52DFEAEDB119FFFCCE19563AFC6FFACA" box="[823,908,1351,1375]" italics="true" pageId="9" pageNumber="61">ósmaka</emphasis>
, which means “canyon” in the language of the Lakota People, in reference to the name of the
<typeStatus id="BF10885DB119FFFCCF905616FD0DFA16" box="[702,750,1387,1411]" pageId="9" pageNumber="61">type</typeStatus>
locality in western South Dakota, an area inhabited by the Lakota.
<taxonomicName id="A7AB4D7CB119FFFCCDDA56F2FEA1FA33" box="[244,322,1423,1446]" family="Synodontidae" genus="Saurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Aulopiformes" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCCDDA56F2FEA1FA33" box="[244,322,1423,1446]" italics="true" pageId="9" pageNumber="61">Saurus</emphasis>
</taxonomicName>
comes from the transliterated Greek meaning “lizard”. The gender of the generic name is masculine.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB56ABFCDFF987" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB56ABFE33FA7A" bold="true" box="[197,464,1494,1519]" pageId="9" pageNumber="61">Locality and horizon.</emphasis>
Calico Canyon, near Buffalo Gap, Custer County, South Dakota (
<bibRefCitation id="043A4B0EB119FFFCC9C956AAFA6CFA7A" author="Gilmore" box="[1255,1423,1495,1519]" pageId="9" pageNumber="61" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
); Lakota Formation, Barremian–Aptian (
<bibRefCitation id="043A4B0EB119FFFCCF635687FCD3F987" author="Tschudy" box="[589,816,1530,1555]" pageId="9" pageNumber="61" refString="Tschudy, R. H., Tschudy, B. D. &amp; Craig, L. C. (1984) Palynological evaluation of Cedar Mountain and Burro Canyon formations, Colorado Plateau. United States Geological Survey Professional Paper, 1281, 1 - 24." type="journal article" year="1984">
Tschudy
<emphasis id="52DFEAEDB119FFFCCF9A5681FD07F986" box="[692,740,1531,1555]" italics="true" pageId="9" pageNumber="61">et al</emphasis>
. 1984
</bibRefCitation>
).
</paragraph>
<paragraph id="601436FFB119FFFCCDEB5563FCF3F9AD" blockId="9.[151,1437,1314,1986]" box="[197,784,1566,1592]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB5563FE43F9A2" bold="true" box="[197,416,1566,1591]" pageId="9" pageNumber="61">Generic diagnosis.</emphasis>
<collectingCountry id="18BC766FB119FFFCCC895562FE25F9A2" box="[423,454,1567,1591]" name="American Samoa" pageId="9" pageNumber="61">As</collectingCountry>
for the species by monotypy.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB553FFF3AF952" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB553FFE43F9CE" bold="true" box="[197,416,1602,1627]" pageId="9" pageNumber="61">Specific diagnosis.</emphasis>
Basal styracosternan distinguished from all other iguanodontians by the following unique combination of characters: cranial end of preacetabular process of ilium modified into horizontal boot, straight dorsal margin of ilium, dorsal margin of ilium thickens mediolaterally towards the
<emphasis id="52DFEAEDB119FFFCC93D55F1FBC9F936" box="[1043,1066,1676,1699]" italics="true" pageId="9" pageNumber="61">M</emphasis>
.
<emphasis id="52DFEAEDB119FFFCC91655F6FB5FF936" box="[1080,1212,1675,1699]" italics="true" pageId="9" pageNumber="61">iliocaudalis</emphasis>
platform (
<figureCitation id="F8902A7AB119FFFCC81D55F7FA63F937" box="[1331,1408,1674,1698]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="9" pageNumber="61">Figs. 8</figureCitation>
A, B, E).
</paragraph>
</subSubSection>
<subSubSection id="28B16574B119FFF9CDEB55AEFC10F979" lastPageId="12" lastPageNumber="64" pageId="9" pageNumber="61" type="discussion">
<paragraph id="601436FFB119FFFCCDEB55AEFC11F857" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB55AEFEDAF97E" bold="true" box="[197,313,1747,1771]" pageId="9" pageNumber="61">Remarks.</emphasis>
<taxonomicName id="A7AB4D7CB119FFFCCC6F55AEFDB9F97E" box="[321,602,1747,1771]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCC6F55AEFDB9F97E" box="[321,602,1747,1771]" italics="true" pageId="9" pageNumber="61">Camptosaurus depressus</emphasis>
</taxonomicName>
was originally distinguished from other species of
<taxonomicName id="A7AB4D7CB119FFFCC98D55AEFAA5F97F" box="[1187,1350,1747,1770]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCC98D55AEFAA5F97F" box="[1187,1350,1747,1770]" italics="true" pageId="9" pageNumber="61">Camptosaurus</emphasis>
</taxonomicName>
chiefly by the “narrowness or depressed nature of the ilia” (
<bibRefCitation id="043A4B0EB119FFFCCFF5558AFC4CF89A" author="Gilmore" box="[731,943,1783,1807]" pageId="9" pageNumber="61" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore 1909:293</bibRefCitation>
). This condition is exhibited by only the left ilium of USNM 4753, on which the dorsal margin of the ilium above the acetabulum faces dorsolaterally and the postacetabular process is nearly horizontal (
<figureCitation id="F8902A7AB119FFFCCFAF5442FD27F8C2" box="[641,708,1855,1879]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="9" pageNumber="61">Fig. 8</figureCitation>
A).
<bibRefCitation id="043A4B0EB119FFFCCFC15442FBC0F8C2" author="Carpenter" box="[751,1059,1855,1879]" pageId="9" pageNumber="61" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
considered the horizontal postacetabular process of USNM 4753 similar to that of DMNH 42504, the
<typeStatus id="BF10885DB119FFFCCECC541FFBA7F8EF" box="[994,1092,1890,1914]" pageId="9" pageNumber="61" type="holotype">holotype</typeStatus>
ilium of
<taxonomicName id="A7AB4D7CB119FFFCC995541EFE38F80A" authority="DiCroce &amp; Carpenter 2001" authorityName="DiCroce &amp; Carpenter" authorityYear="2001" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB119FFFCC995541EFA7FF8EE" box="[1211,1436,1891,1915]" italics="true" pageId="9" pageNumber="61">Planicoxa venenica</emphasis>
(
<bibRefCitation id="043A4B0EB119FFFCCDB054FAFE31F80A" author="DiCroce" box="[158,466,1927,1951]" pageId="9" pageNumber="61" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter 2001</bibRefCitation>
)
</taxonomicName>
; thus, they referred
<taxonomicName id="A7AB4D7CB119FFFCCF9954F5FCA7F80A" box="[695,836,1927,1951]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCF9954F5FD29F80A" box="[695,714,1928,1951]" italics="true" pageId="9" pageNumber="61">C</emphasis>
.
<emphasis id="52DFEAEDB119FFFCCFF954FAFCA7F80A" box="[727,836,1927,1951]" italics="true" pageId="9" pageNumber="61">depressus</emphasis>
</taxonomicName>
to
<taxonomicName id="A7AB4D7CB119FFFCCE4954FAFC3AF80A" box="[871,985,1927,1951]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCCE4954FAFC3AF80A" box="[871,985,1927,1951]" italics="true" pageId="9" pageNumber="61">Planicoxa</emphasis>
</taxonomicName>
, as
<taxonomicName id="A7AB4D7CB119FFFCC92A54F5FB62F80A" box="[1028,1153,1927,1951]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressa">
<emphasis id="52DFEAEDB119FFFCC92A54F5FBF1F80A" box="[1028,1042,1928,1951]" italics="true" pageId="9" pageNumber="61">P</emphasis>
.
<emphasis id="52DFEAEDB119FFFCC93154FAFB62F80A" box="[1055,1153,1927,1951]" italics="true" pageId="9" pageNumber="61">depressa</emphasis>
</taxonomicName>
. However, the horizontal postacetabular process of USNM 4753 is more likely a product of distortion.
</paragraph>
<paragraph id="601436FFB11AFFFFCDE953EAFB9CFE61" blockId="10.[151,1436,151,500]" pageId="10" pageNumber="62">
The blade of the left ilium of USNM 4753 is heavily fractured by corresponding cracks on the lateral and medial surfaces (
<figureCitation id="F8902A7AB11AFFFFCC7953C1FE47FF41" box="[343,420,188,212]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62">Figs. 8</figureCitation>
A, B). Four roughly vertical cracks divide the blade of the ilium into four segments; these four cracks all merge with a roughly horizontal elongate crack extending craniocaudally dorsal to the acetabulum and peduncles. These cracks mean that the three more cranial segments are suspended in filling substance and that their orientations relative to the rest of the ilium are not certain. Furthermore, there is a point of offset on the lateral surface of the ilium at which one of the blade fragments overhangs the acetabular portion (
<figureCitation id="F8902A7AB11AFFFFC9BB5231FB3BFEF1" box="[1173,1240,332,356]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62">Fig. 8</figureCitation>
C); this overhang is due to filler between the two pieces and does not reflect the actual morphology of the ilium. Notwithstanding uncertainty regarding the reconstruction of the iliac blade fragments, the postacetabular process does project far dorsolaterally relative to the ischial peduncle (
<figureCitation id="F8902A7AB11AFFFFCF9E52CAFD15FE5A" box="[688,758,439,463]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62">Fig. 8</figureCitation>
D). However, comparing the case of USNM 4753 to other basal iguanodonts, it becomes clear that this is probably a product of plastic deformation.
</paragraph>
<caption id="34D46677B11AFFFFCDB95619FD67FA22" httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62" targetBox="[151,1436,537,1358]" targetPageId="10">
<paragraph id="601436FFB11AFFFFCDB95619FD67FA22" blockId="10.[151,1436,1380,1463]" pageId="10" pageNumber="62">
<emphasis id="52DFEAEDB11AFFFFCDB95619FEF1FAEC" bold="true" box="[151,274,1380,1403]" pageId="10" pageNumber="62">FIGURE 8.</emphasis>
Left ilium of USNM 4753 (holotype of
<taxonomicName id="A7AB4D7CB11AFFFFCF835619FC4EFAEF" box="[685,941,1380,1402]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="62" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11AFFFFCF835619FC4EFAEF" box="[685,941,1380,1402]" italics="true" pageId="10" pageNumber="62">Osmakasaurus depressus</emphasis>
</taxonomicName>
) in (
<emphasis id="52DFEAEDB11AFFFFCEF05619FC13FAEC" bold="true" box="[990,1008,1380,1401]" pageId="10" pageNumber="62">A</emphasis>
) lateral, (
<emphasis id="52DFEAEDB11AFFFFC97A5618FB86FAEF" bold="true" box="[1108,1125,1381,1402]" pageId="10" pageNumber="62">B</emphasis>
) medial, (
<emphasis id="52DFEAEDB11AFFFFC9E05619FB03FAEE" bold="true" box="[1230,1248,1380,1403]" pageId="10" pageNumber="62">C</emphasis>
) caudolateral, and (
<emphasis id="52DFEAEDB11AFFFFCDB156FEFF52FA0D" bold="true" box="[159,177,1411,1432]" pageId="10" pageNumber="62">D</emphasis>
) caudal views. Right ilium of USNM 4753 in (
<emphasis id="52DFEAEDB11AFFFFCF8756FEFD59FA0D" bold="true" box="[681,698,1411,1432]" pageId="10" pageNumber="62">E</emphasis>
) lateral and (
<emphasis id="52DFEAEDB11AFFFFCE6756FEFCBBFA0D" bold="true" box="[841,856,1411,1432]" pageId="10" pageNumber="62">F</emphasis>
) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm.
</paragraph>
</caption>
<paragraph id="601436FFB11AFFFFCDE95698FD64F889" blockId="10.[151,1436,1508,2036]" pageId="10" pageNumber="62">
Among the skeletons of
<taxonomicName id="A7AB4D7CB11AFFFFCCF05699FCEAFA69" box="[478,777,1508,1532]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="62" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11AFFFFCCF05699FCEAFA69" box="[478,777,1508,1532]" italics="true" pageId="10" pageNumber="62">Iguanodon bernissartensis</emphasis>
</taxonomicName>
excavated from the Bernissart Mine in
<collectingCountry id="18BC766FB11AFFFFC9E05699FACDFA69" box="[1230,1326,1508,1532]" name="Belgium" pageId="10" pageNumber="62">Belgium</collectingCountry>
(
<bibRefCitation id="043A4B0EB11AFFFFC8105699FF30F9B5" author="Norman" pageId="10" pageNumber="62" refString="Norman, D. B. (1980) On the ornithischian dinosaur Iguanodon bernissartensis from Belgium. Memoires Institut Royal des Sciences Naturelles de Belgique, 178, 1 - 103." type="journal article" year="1980">Norman 1980</bibRefCitation>
), there is variation in the forms of the ilia that can be attributed to deformation. The left ilium of the
<typeStatus id="BF10885DB11AFFFFC8005575FA74F9B5" box="[1326,1431,1544,1568]" pageId="10" pageNumber="62" type="lectotype">lectotype</typeStatus>
, IRSNB 1534, is well preserved with a nearly intact, gently convex dorsal margin along which there extends a thickened, laterally everted rim (
<figureCitation id="F8902A7AB11AFFFFCCE1552DFDF0F9FD" box="[463,531,1616,1640]" captionStart="FIGURE 9" captionStartId="11.[151,250,1766,1789]" captionTargetBox="[152,1390,343,1723]" captionTargetId="figure@11.[151,1436,307,1745]" captionTargetPageId="11" captionText="FIGURE 9. Left ilia of Iguanodon bernissartensis in lateral view. (A) IRSNB 1534 (lectotype), dorsal margin outlined in red; (B) IRSNB 1536, dorsal margin outlined in red; (C) IRSNB 1561." httpUri="https://zenodo.org/record/200820/files/figure.png" pageId="10" pageNumber="62">Fig. 9</figureCitation>
A). The postacetabular process tapers towards its caudal end. This is taken to be the undistorted form of the ilium. In contrast, the left ilia of IRSNB
<date id="1415103FB11AFFFFCEBF5509FBDBF919" box="[913,1080,1652,1676]" pageId="10" pageNumber="62" value="1536" valueMax="1561">1536 and 1561</date>
resemble those of USNM
<quantity id="A7539B1AB11AFFFFC84A5509FF4EF925" metricMagnitude="2" metricUnit="m" metricValue="1.207262" pageId="10" pageNumber="62" unit="in" value="4753.0">4753 in</quantity>
their dorsolaterally projecting dorsal margins dorsal to the ischial peduncle (
<figureCitation id="F8902A7AB11AFFFFC93F55E5FB83F925" box="[1041,1120,1688,1712]" captionStart="FIGURE 9" captionStartId="11.[151,250,1766,1789]" captionTargetBox="[152,1390,343,1723]" captionTargetId="figure@11.[151,1436,307,1745]" captionTargetPageId="11" captionText="FIGURE 9. Left ilia of Iguanodon bernissartensis in lateral view. (A) IRSNB 1534 (lectotype), dorsal margin outlined in red; (B) IRSNB 1536, dorsal margin outlined in red; (C) IRSNB 1561." httpUri="https://zenodo.org/record/200820/files/figure.png" pageId="10" pageNumber="62">Figs. 9</figureCitation>
B, C), giving the ilia a sinuous appearance in lateral view. That this morphology is likely a result of deformation is evinced in IRSNB 1536 by the sacral neural spines being inclined noticeably to the left side of the animal (
<figureCitation id="F8902A7AB11AFFFFC922559DFBBCF96D" box="[1036,1119,1760,1784]" captionStart="FIGURE 10" captionStartId="12.[151,250,1212,1235]" captionTargetBox="[501,1087,298,1189]" captionTargetId="figure@12.[496,1091,294,1191]" captionTargetPageId="12" captionText="FIGURE 10. Sacrum and ilia of IRSNB 1536 (Iguanodon bernissartensis) in dorsal view." httpUri="https://zenodo.org/record/200821/files/figure.png" pageId="10" pageNumber="62">Fig. 10</figureCitation>
), an obvious result of deformation that could have also affected the ilia.
</paragraph>
<paragraph id="601436FFB11AFFFECDEB5455FBB6FE89" blockId="10.[151,1436,1508,2036]" lastBlockId="11.[151,1436,151,284]" lastPageId="11" lastPageNumber="63" pageId="10" pageNumber="62">
Furthermore, the
<typeStatus id="BF10885DB11AFFFFCCA75455FE08F8D5" box="[393,491,1832,1856]" pageId="10" pageNumber="62" type="holotype">holotype</typeStatus>
skeleton of
<taxonomicName id="A7AB4D7CB11AFFFFCF585455FC94F8D5" box="[630,887,1832,1856]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="62" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB11AFFFFCF585455FC94F8D5" box="[630,887,1832,1856]" italics="true" pageId="10" pageNumber="62">Uteodon aphanoecetes</emphasis>
</taxonomicName>
, CM 11337, exhibits a left ilium strikingly similar to that of USNM
<quantity id="A7539B1AB11AFFFFCCA95431FE3EF8F1" box="[391,477,1868,1892]" metricMagnitude="2" metricUnit="m" metricValue="1.207262" pageId="10" pageNumber="62" unit="in" value="4753.0">4753 in</quantity>
lateral and caudal views, with the dorsal margin of the ilium facing almost laterally and the blade of the ilium projecting dorsolaterally far past the ischial peduncle (
<figureCitation id="F8902A7AB11AFFFFC90C540DFB92F81D" box="[1058,1137,1904,1928]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="10" pageNumber="62">Figs. 6</figureCitation>
B, C). That this is a consequence of plastic deformation in CM 11337 is indicated by the condition of the sacral neural spines, which incline sharply towards the left side of the skeleton, as documented above in IRSNB 1536 (
<figureCitation id="F8902A7AB11AFFFFC96754C5FB6DF845" box="[1097,1166,1976,2000]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="10" pageNumber="62">Fig. 6</figureCitation>
D; see also
<bibRefCitation id="043A4B0EB11AFFFFC83E54C5FEC8F861" author="Carpenter" pageId="10" pageNumber="62" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson 2008</bibRefCitation>
, fig. 13E); the blade of the left ilium likely was subject to the same force that distorted the neural spines. Furthermore, while the blade of the right ilium of CM 11337 is incomplete, enough is preserved to indicate that the iliac blade was nearly vertical and projected dorsally relative to the ischial peduncle, as in ilia of
<taxonomicName id="A7AB4D7CB11BFFFEC81453C0FED2FF6D" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB11BFFFEC81453C0FED2FF6D" italics="true" pageId="11" pageNumber="63">Camptosaurus dispar</emphasis>
</taxonomicName>
(e.g., YPM 1877, YPM 8622, USNM 5473;
<figureCitation id="F8902A7AB11BFFFECE1F53A2FC9CFF62" box="[817,895,223,247]" captionStart="FIGURE 4" captionStartId="6.[151,250,720,743]" captionTargetBox="[151,1436,193,698]" captionTargetId="figure@6.[151,1436,193,699]" captionTargetPageId="6" captionText="FIGURE 4. Ilia of Camptosaurus dispar in lateral view. (A) right ilium of YPM 1877 (holotype) from Quarry 13; (B) right ilium of YPM 8622 from Quarry 13; (C) left ilium of USNM 5473 from Quarry 13; (D) left ilium of YPM PU 14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in A, B, and D are copyright YPM. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="11" pageNumber="63">Figs. 4</figureCitation>
A–C;
<figureCitation id="F8902A7AB11BFFFECEE253A2FBECFF62" box="[972,1039,223,247]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="11" pageNumber="63">Fig. 6</figureCitation>
E). The situation in USNM 4753 is akin to that of CM 11337, i.e. one ilium is plastically deformed while the other is not.
</paragraph>
<caption id="34D46677B11BFFFECDB9559BFCD6F88E" httpUri="https://zenodo.org/record/200820/files/figure.png" pageId="11" pageNumber="63" targetBox="[152,1390,343,1723]" targetPageId="11">
<paragraph id="601436FFB11BFFFECDB9559BFCD6F88E" blockId="11.[151,1436,1766,1820]" pageId="11" pageNumber="63">
<emphasis id="52DFEAEDB11BFFFECDB9559BFEF0F969" bold="true" box="[151,275,1766,1789]" pageId="11" pageNumber="63">FIGURE 9.</emphasis>
Left ilia of
<taxonomicName id="A7AB4D7CB11BFFFECCA3559AFD78F968" box="[397,667,1767,1789]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11BFFFECCA3559AFD78F968" box="[397,667,1767,1789]" italics="true" pageId="11" pageNumber="63">Iguanodon bernissartensis</emphasis>
</taxonomicName>
in lateral view. (
<emphasis id="52DFEAEDB11BFFFECE69559AFCBAF969" bold="true" box="[839,857,1767,1788]" pageId="11" pageNumber="63">A</emphasis>
) IRSNB 1534 (lectotype), dorsal margin outlined in red; (
<emphasis id="52DFEAEDB11BFFFECDB1547BFF53F88E" bold="true" box="[159,176,1798,1819]" pageId="11" pageNumber="63">B</emphasis>
) IRSNB 1536, dorsal margin outlined in red; (
<emphasis id="52DFEAEDB11BFFFECFA45478FD7FF889" bold="true" box="[650,668,1797,1820]" pageId="11" pageNumber="63">C</emphasis>
) IRSNB 1561.
</paragraph>
</caption>
<paragraph id="601436FFB11BFFF9CDEB5435FCAFFF62" blockId="11.[151,1436,1864,2032]" lastBlockId="12.[151,1436,151,248]" lastPageId="12" lastPageNumber="64" pageId="11" pageNumber="63">
The preserved morphology of USNM 4753 is itself evidence against the supposed horizontal postacetabular process. The preserved part of the right ilium of USNM 4753 does not exhibit any hint of a folded, horizontally projecting iliac blade (
<emphasis id="52DFEAEDB11BFFFECCB654ECFE02F83D" box="[408,481,1937,1960]" italics="true" pageId="11" pageNumber="63">contra</emphasis>
<bibRefCitation id="043A4B0EB11BFFFECCC454EDFCF1F83D" author="Carpenter" box="[490,786,1936,1960]" pageId="11" pageNumber="63" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson 2008</bibRefCitation>
) (
<figureCitation id="F8902A7AB11BFFFECE0454EDFC8DF83D" box="[810,878,1936,1960]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="11" pageNumber="63">Fig. 8</figureCitation>
E). Indeed, the cross-section of the right ilium in caudal view indicates that the iliac blade was nearly vertical, as in
<taxonomicName id="A7AB4D7CB11BFFFECE5054C8FB8CF859" box="[894,1135,1972,1996]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB11BFFFECE5054C8FB8CF859" box="[894,1135,1972,1996]" italics="true" pageId="11" pageNumber="63">Camptosaurus dispar</emphasis>
</taxonomicName>
(
<figureCitation id="F8902A7AB11BFFFEC95054C9FB22F859" box="[1150,1217,1972,1996]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="11" pageNumber="63">Fig. 8</figureCitation>
F). Moreover, comparing the left ilium with the preserved portion of the right reveals that if the morphology of the left ilium were genuine, the right ilium should exhibit the cranial region of the folded iliac blade (
<figureCitation id="F8902A7AB11CFFF9C91453EAFB6EFF3A" box="[1082,1165,151,175]" captionStart="FIGURE 11" captionStartId="13.[151,250,1103,1126]" captionTargetBox="[151,1436,327,1082]" captionTargetId="figure@13.[151,1436,327,1082]" captionTargetPageId="13" captionText="FIGURE 11. Left (A) and right (B, reversed) ilia of USNM 4753 (holotype of Osmakasaurus depressus) in lateral view, set to the same scale and with the notch between the preacetabular process and pubic peduncle lined up. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200822/files/figure.png" pageId="12" pageNumber="64">Fig. 11</figureCitation>
). Thus, the morphology used by
<bibRefCitation id="043A4B0EB11CFFF9CDDD53C1FE42FF41" author="Gilmore" box="[243,417,188,212]" pageId="12" pageNumber="64" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore (1909)</bibRefCitation>
to demarcate
<taxonomicName id="A7AB4D7CB11CFFF9CF1353C0FD29FF41" box="[573,714,188,212]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CF1353C0FDB3FF41" box="[573,592,189,212]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CF7353C1FD29FF41" box="[605,714,188,212]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
and by
<bibRefCitation id="043A4B0EB11CFFF9CE0C53C1FBB0FF41" author="Carpenter" box="[802,1107,188,212]" pageId="12" pageNumber="64" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
to assign the species to
<taxonomicName id="A7AB4D7CB11CFFF9C87053C1FF37FF6D" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB11CFFF9C87053C1FF37FF6D" italics="true" pageId="12" pageNumber="64">Planicoxa</emphasis>
</taxonomicName>
is the product of plastic deformation and fragmentation.
</paragraph>
<caption id="34D46677B11CFFF9CDB957C1FBCAFB46" box="[151,1065,1212,1235]" httpUri="https://zenodo.org/record/200821/files/figure.png" pageId="12" pageNumber="64" targetBox="[501,1087,298,1189]" targetPageId="12">
<paragraph id="601436FFB11CFFF9CDB957C1FBCAFB46" blockId="12.[151,1065,1212,1235]" box="[151,1065,1212,1235]" pageId="12" pageNumber="64">
<emphasis id="52DFEAEDB11CFFF9CDB957C1FEFCFB47" bold="true" box="[151,287,1212,1235]" pageId="12" pageNumber="64">FIGURE 10.</emphasis>
Sacrum and ilia of IRSNB 1536 (
<taxonomicName id="A7AB4D7CB11CFFF9CF5657C0FC66FB46" box="[632,901,1213,1235]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11CFFF9CF5657C0FC66FB46" box="[632,901,1213,1235]" italics="true" pageId="12" pageNumber="64">Iguanodon bernissartensis</emphasis>
</taxonomicName>
) in dorsal view.
</paragraph>
</caption>
<paragraph id="601436FFB11CFFF9CDEB5782FC10F979" blockId="12.[151,1436,1279,1773]" pageId="12" pageNumber="64">
Although crushed and broken, a unique combination of characters can be discerned in the ilia of USNM 4753 (see “Specific diagnosis” above). The dorsal margin of the left ilium faces dorsolaterally due to distortion of the iliac blade (
<figureCitation id="F8902A7AB11CFFF9CC34563AFE85FACA" box="[282,358,1351,1375]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="12" pageNumber="64">Figs. 8</figureCitation>
A, C), making the ilium appear dorsally convex; however, the dorsal margin itself lacks any curvature and is straight. The dorsal margin of the ilium thickens mediolaterally towards the postacetabular process (
<figureCitation id="F8902A7AB11CFFF9CDB156F2FF08FA32" box="[159,235,1423,1447]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="12" pageNumber="64">Figs. 8</figureCitation>
A, C).
<taxonomicName id="A7AB4D7CB11CFFF9CC1656EDFE25FA3D" box="[312,454,1424,1448]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CC1656EDFEA8FA32" box="[312,331,1424,1447]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CC7756EDFE25FA3D" box="[345,454,1424,1448]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
is more derived than
<taxonomicName id="A7AB4D7CB11CFFF9CF9756EDFC49FA3D" box="[697,938,1424,1448]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB11CFFF9CF9756EDFC49FA3D" box="[697,938,1424,1448]" italics="true" pageId="12" pageNumber="64">Camptosaurus dispar</emphasis>
</taxonomicName>
in exhibiting a horizontal boot at the cranial end of the preacetabular process (
<figureCitation id="F8902A7AB11CFFF9CF3056C9FD80FA59" box="[542,611,1460,1484]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="12" pageNumber="64">Fig. 8</figureCitation>
E), a feature shared with other more derived basal iguanodonts such as
<taxonomicName id="A7AB4D7CB11CFFF9CDB956A5FE6CFA65" box="[151,399,1496,1520]" class="Reptilia" genus="Cedrorestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="crichtoni">
<emphasis id="52DFEAEDB11CFFF9CDB956A5FE6CFA65" box="[151,399,1496,1520]" italics="true" pageId="12" pageNumber="64">Cedrorestes crichtoni</emphasis>
</taxonomicName>
(DMNH 47994),
<taxonomicName id="A7AB4D7CB11CFFF9CF4256A5FC7FFA65" box="[620,924,1496,1520]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11CFFF9CF4256A5FC7FFA65" box="[620,924,1496,1520]" italics="true" pageId="12" pageNumber="64">Iguanodon bernissartensis</emphasis>
</taxonomicName>
(IRSNB 1534), and
<emphasis id="52DFEAEDB11CFFF9C9B056A5FA7FFA65" box="[1182,1436,1496,1520]" italics="true" pageId="12" pageNumber="64">Eolambia caroljonesa</emphasis>
(CEUM 52090). It might seem acceptable to refer
<taxonomicName id="A7AB4D7CB11CFFF9CFFA5680FC81F981" box="[724,866,1532,1556]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CFFA5680FD04F981" box="[724,743,1533,1556]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CFDB5681FC81F981" box="[757,866,1532,1556]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
to
<taxonomicName id="A7AB4D7CB11CFFF9CEA65681FBE6F981" box="[904,1029,1532,1556]" class="Reptilia" family="Iguanodontidae" genus="Dakotadon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB11CFFF9CEA65681FBE6F981" box="[904,1029,1532,1556]" italics="true" pageId="12" pageNumber="64">Dakotadon</emphasis>
</taxonomicName>
because USNM 4753 and the
<typeStatus id="BF10885DB11CFFF9C84F5681FF24F9AC" pageId="12" pageNumber="64" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A7AB4D7CB11CFFF9CDDE555CFE71F9AD" box="[240,402,1568,1592]" class="Reptilia" family="Iguanodontidae" genus="Dakotadon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="lakotaensis">
<emphasis id="52DFEAEDB11CFFF9CDDE555CFEE7F9AD" box="[240,260,1569,1592]" italics="true" pageId="12" pageNumber="64">D</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CC3A555DFE71F9AD" box="[276,402,1568,1592]" italics="true" pageId="12" pageNumber="64">lakotaensis</emphasis>
</taxonomicName>
, SDSM 8656, both came from the Lakota Formation of South Dakota. However, the only overlapping elements between the two specimens are poorly preserved caudal vertebrae that display no features that would positively link “
<emphasis id="52DFEAEDB11CFFF9CCE95515FE39F9EA" box="[455,474,1640,1663]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.”
<taxonomicName id="A7AB4D7CB11CFFF9CCDA5515FD81F915" box="[500,610,1640,1664]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CCDA5515FD81F915" box="[500,610,1640,1664]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
with
<taxonomicName id="A7AB4D7CB11CFFF9CFB15514FCDCF915" box="[671,831,1640,1664]" class="Reptilia" family="Iguanodontidae" genus="Dakotadon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="lakotaensis">
<emphasis id="52DFEAEDB11CFFF9CFB15514FD50F915" box="[671,691,1641,1664]" italics="true" pageId="12" pageNumber="64">D</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CFEF5515FCDCF915" box="[705,831,1640,1664]" italics="true" pageId="12" pageNumber="64">lakotaensis</emphasis>
</taxonomicName>
.
<collectingCountry id="18BC766FB11CFFF9CE635515FC8FF915" box="[845,876,1640,1664]" name="American Samoa" pageId="12" pageNumber="64">As</collectingCountry>
the specimen clearly does not pertain to the genus
<taxonomicName id="A7AB4D7CB11CFFF9CDB955F0FED9F931" box="[151,314,1677,1700]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB11CFFF9CDB955F0FED9F931" box="[151,314,1677,1700]" italics="true" pageId="12" pageNumber="64">Camptosaurus</emphasis>
</taxonomicName>
, cannot be confidently referred to another existing genus, and exhibits a unique combination of characters, a new genus should be created to accommodate “
<emphasis id="52DFEAEDB11CFFF9CE6F55CDFCB7F952" box="[833,852,1712,1735]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.”
<taxonomicName id="A7AB4D7CB11CFFF9CE4155CDFC3FF95D" box="[879,988,1712,1736]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CE4155CDFC3FF95D" box="[879,988,1712,1736]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
and achieve taxonomic stability. Hence the novel genus and combination proposed herein,
<taxonomicName id="A7AB4D7CB11CFFF9CFFC55A9FC0FF979" box="[722,1004,1748,1772]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CFFC55A9FC0FF979" box="[722,1004,1748,1772]" italics="true" pageId="12" pageNumber="64">Osmakasaurus depressus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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