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<document id="2CC8115AD64FA4B96FA569FF9510769D" ID-CLB-Dataset="22128" ID-DOI="10.5852/ejt.2013.69" ID-GBIF-Dataset="a2a819a6-f7a9-4d53-b426-09dde497f788" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827691" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589478069293" checkinUser="carolina" docAuthor="Cheek, Martin &amp; Jebb, Matthew" docDate="2013" docId="03D787F9FF8EFC51FDFA1432FC0B71C4" docLanguage="en" docName="ejt-2013-69.pdf.imf" docOrigin="European Journal of Taxonomy 69" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nepenthes kurata Jebb &amp; Cheek 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="FFEEFF81FF8BFC59FFDD165CFF93762F" masterDocTitle="Recircumscription of the Nepenthes alata group (Caryophyllales: Nepenthaceae), in the Philippines, with four new species" masterLastPageNumber="23" masterPageNumber="1" pageNumber="6" updateTime="1698839761783" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="4446A2518CF27061517F98202CB229E7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F1F7B9E54259DF04908F33051CDD5CEC">
<mods:title id="1F1EFCFE65EAD09FAE6BDA620EA89245">Recircumscription of the Nepenthes alata group (Caryophyllales: Nepenthaceae), in the Philippines, with four new species</mods:title>
</mods:titleInfo>
<mods:name id="39B7D73DF5F0A1ECAB9A707BE4FE1EBB" type="personal">
<mods:role id="6A1746F0B031A438777B702E1BEB0F3E">
<mods:roleTerm id="A8447DFF514B3C75D67372EEE6C02FF7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6D8ECE796DA349F616DC68665214830">Cheek, Martin</mods:namePart>
<mods:affiliation id="A48D4FDFF76E6CE787B990D576810582">Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, U. K. Email: m. cheek @ kew. org (corresponding author)</mods:affiliation>
</mods:name>
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<mods:role id="511A96853402EA6D69CBCE2D7C41F6BF">
<mods:roleTerm id="D5F3AAAFEF72B3DB05CCF0D95145DB75">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5128F5705491CC4EBB7611AF68733F56">Jebb, Matthew</mods:namePart>
<mods:affiliation id="AFA2D707067E7C481C87A0A1C323AD1F">National Botanic Garden, Glasnevin, Dublin 9, Ireland Email: matthew. jebb @ opw. ie</mods:affiliation>
</mods:name>
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<mods:title id="E51F7D352AE17BD7888CE7104C128521">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="BC7BF45A9228CB5222609F755CF6C7A3">
<mods:date id="A9614B2EECFF82C53061E822F11CE194">2013</mods:date>
<mods:detail id="BE2EAE0869C2A017B052E7DEC643C50A" type="pubDate">
<mods:number id="41FD8FD042C7EF670C571CE7962F73F2">2013-12-12</mods:number>
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<mods:detail id="B62D2784E97F92EE1FE6B5E1F602341F" type="issue">
<mods:number id="A8963C9172435D292B6A69945C01807B">69</mods:number>
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<mods:start id="14D401326357B00C52EF2FA992C659B7">1</mods:start>
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<mods:classification id="24205745DFC5AE47596CFDDCC6752AAD">journal article</mods:classification>
<mods:identifier id="FAD6BD404F1B1144A88B55BBCC4B3E51" type="CLB-Dataset">22128</mods:identifier>
<mods:identifier id="DF659A3492B41645E40C8778522ED1FC" type="DOI">10.5852/ejt.2013.69</mods:identifier>
<mods:identifier id="CD47936B5FC1405C8668EDF633F2723B" type="GBIF-Dataset">a2a819a6-f7a9-4d53-b426-09dde497f788</mods:identifier>
<mods:identifier id="F636F2AAEA5C101E42A08427F860D14B" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="59F23863C342C20D6780FA9BF31306AC" type="Zenodo-Dep">3827691</mods:identifier>
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<treatment id="03D787F9FF8EFC51FDFA1432FC0B71C4" ID-DOI="http://doi.org/10.5281/zenodo.3846735" ID-GBIF-Taxon="164152690" ID-Zenodo-Dep="3846735" LSID="urn:lsid:plazi:treatment:03D787F9FF8EFC51FDFA1432FC0B71C4" httpUri="http://treatment.plazi.org/id/03D787F9FF8EFC51FDFA1432FC0B71C4" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
<subSubSection id="C3646564FF8EFC5CFDFA1432FB9E74A6" box="[551,1037,622,649]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BC136EFFF8EFC5CFDFA1432FB9E74A6" blockId="5.[551,1037,622,720]" box="[551,1037,622,649]" pageId="5" pageNumber="6">
<heading id="D0898183FF8EFC5CFDFA1432FB9E74A6" box="[551,1037,622,649]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="4C7E4D6CFF8EFC5CFDFA1432FC3874A6" ID-CoL="73X8D" authority="Jebb &amp; Cheek" authorityName="Jebb &amp; Cheek" authorityYear="2013" box="[551,939,622,649]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="kurata" status="sp. nov.">
<emphasis id="B90AEAFDFF8EFC5CFDFA1432FD6E74A6" bold="true" box="[551,765,622,649]" italics="true" pageId="5" pageNumber="6">Nepenthes kurata</emphasis>
Jebb &amp; Cheek
</taxonomicName>
<taxonomicNameLabel id="A2395786FF8EFC5CFC6F1433FB9E74A6" box="[946,1037,623,649]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF8EFC5CFD9A14CEFCAF74FF" pageId="5" pageNumber="6" type="description">
<paragraph id="8BC136EFFF8EFC5CFD9A14CEFC7E7483" blockId="5.[551,1037,622,720]" box="[583,1005,658,684]" pageId="5" pageNumber="6">urn:lsid:ipni.org:names:77134486-1</paragraph>
<paragraph id="8BC136EFFF8EFC5CFD2A14EAFCAF74FF" blockId="5.[551,1037,622,720]" box="[759,828,694,720]" pageId="5" pageNumber="6">
<figureCitation id="13452A6AFF8EFC5CFD2A14EAFCAF74FF" box="[759,828,694,720]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF8EFC5CFF6014ABFB2075A7" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BC136EFFF8EFC5CFF6014ABFEA5753E" blockId="5.[189,310,759,785]" box="[189,310,759,785]" pageId="5" pageNumber="6">
<heading id="D0898183FF8EFC5CFF6014ABFEA5753E" bold="true" box="[189,310,759,785]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B90AEAFDFF8EFC5CFF6014ABFEA5753E" bold="true" box="[189,310,759,785]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8EFC5CFF60157AFB2075A7" blockId="5.[189,1398,806,904]" pageId="5" pageNumber="6">
Differs from
<taxonomicName id="4C7E4D6CFF8EFC5CFE87157BFD3F756F" authority="Sh. Kurata" authorityName="Sh. Kurata" authorityYear="2011" box="[346,684,806,832]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="mindanaoensis">
<emphasis id="B90AEAFDFF8EFC5CFE87157BFDBE756F" box="[346,557,806,832]" italics="true" pageId="5" pageNumber="6">N. mindanaoensis</emphasis>
Sh.Kurata
</taxonomicName>
in the petiole wings patent (not involute), the hairs of stem, midrib and leaf-edge bushy,
<quantity id="4C869B0AFF8EFC5CFDD21516FDF9754B" box="[527,618,842,868]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="mm" value="0.1">0.1 mm</quantity>
long, not bristle-like
<quantity id="4C869B0AFF8EFC5CFCB71515FC71754B" box="[874,994,841,868]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="6" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long; the lid about half as long as the mouth, lid base rounded or truncate (not about as long as the mouth, base cordate).
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF8EFC5CFF6015F3FD3A7211" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BC136EFFF8EFC5CFF6015F3FED775E6" blockId="5.[189,324,943,969]" box="[189,324,943,969]" pageId="5" pageNumber="6">
<heading id="D0898183FF8EFC5CFF6015F3FED775E6" bold="true" box="[189,324,943,969]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B90AEAFDFF8EFC5CFF6015F3FED775E6" bold="true" box="[189,324,943,969]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8EFC5CFF601581FD3A7211" blockId="5.[189,1399,989,1087]" pageId="5" pageNumber="6">
Named as a noun in apposition for Shigeo Kurata, whose book on the
<taxonomicName id="4C7E4D6CFF8EFC5CFBD31582FB1B75D7" baseAuthorityName="Jebb &amp; Cheek" baseAuthorityYear="1997" box="[1038,1160,990,1016]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B90AEAFDFF8EFC5CFBD31582FB1B75D7" box="[1038,1160,990,1016]" italics="true" pageId="5" pageNumber="6">Nepenthes</emphasis>
</taxonomicName>
of Mount Kinabalu (
<bibRefCitation id="EFEF4B1EFF8EFC5CFF18125DFEC47233" author="Kurata S." box="[197,343,1025,1052]" pageId="5" pageNumber="6" refId="ref14360" refString="Kurata S. 1976. The Nepenthes of Mt Kinabalu. Sabah National Park Trustees, Kota Kinabalu, Sabah." type="book" year="1976">Kurata 1976</bibRefCitation>
) inspired interest in the genus among its many readers, and whose descriptions of
<taxonomicName id="4C7E4D6CFF8EFC5CFB21125DFAE57234" baseAuthorityName="Jebb &amp; Cheek" baseAuthorityYear="1997" box="[1276,1398,1025,1051]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B90AEAFDFF8EFC5CFB21125DFAE57234" box="[1276,1398,1025,1051]" italics="true" pageId="5" pageNumber="6">Nepenthes</emphasis>
</taxonomicName>
are models of detail, precision and clarity.
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF8EFC5CFF60123AFF6E7382" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BC136EFFF8EFC5CFF60123AFF6972AF" blockId="5.[189,250,1126,1152]" box="[189,250,1126,1152]" pageId="5" pageNumber="6">
<heading id="D0898183FF8EFC5CFF60123AFF6972AF" bold="true" box="[189,250,1126,1152]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B90AEAFDFF8EFC5CFF60123AFF6972AF" bold="true" box="[189,250,1126,1152]" pageId="5" pageNumber="6">
<typeStatus id="54C5884DFF8EFC5CFF60123AFF6972AF" box="[189,250,1126,1152]" pageId="5" pageNumber="6">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8EFC5CFF6012C9FCAE72FC" blockId="5.[189,1399,1173,1235]" pageId="5" pageNumber="6">
<collectingCountry id="F369767FFF8EFC5CFF6012C9FEFD7280" box="[189,366,1173,1199]" name="Philippines" pageId="5" pageNumber="6">PHILIPPINES</collectingCountry>
. Mindanao, “Prov. of Misamis, Mount Malindang”.
<date id="FFC0102FFF8EFC5CFC1212C9FBDB7280" box="[975,1096,1173,1199]" pageId="5" pageNumber="6" value="1906-05">May 1906</date>
,
<emphasis id="B90AEAFDFF8EFC5CFB8912CAFE3472FD" italics="true" pageId="5" pageNumber="6">Mearns &amp; Hutchinson in Forest Bureau 4632</emphasis>
(
<typeStatus id="54C5884DFF8EFC5CFE6B12E4FD8C72FD" box="[438,543,1208,1234]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
<collectionCode id="ED6FAE2AFF8EFC5CFDFB12E5FDD072FC" box="[550,579,1209,1235]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="6" type="Herbarium">K</collectionCode>
!; isotype
<collectingCountry id="F369767FFF8EFC5CFD6C12E5FD7372FC" box="[689,736,1209,1235]" name="Philippines" pageId="5" pageNumber="6">PH</collectingCountry>
!).
<figureCitation id="13452A6AFF8EFC5CFD2512E4FCA472FC" box="[760,823,1208,1235]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="5" pageNumber="6">Fig 1</figureCitation>
.
</paragraph>
<paragraph id="8BC136EFFF8EFC5CFF6012A6FEA1733B" blockId="5.[189,306,1274,1300]" box="[189,306,1274,1300]" pageId="5" pageNumber="6">
<heading id="D0898183FF8EFC5CFF6012A6FEA1733B" bold="true" box="[189,306,1274,1300]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B90AEAFDFF8EFC5CFF6012A6FEA1733B" bold="true" box="[189,306,1274,1300]" pageId="5" pageNumber="6">Synonym</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8EFC5CFF601375FF6E7382" blockId="5.[189,1399,1321,1453]" pageId="5" pageNumber="6">
<emphasis id="B90AEAFDFF8EFC5CFF601375FEE9736C" box="[189,378,1321,1347]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C7E4D6CFF8EFC5CFF601375FEA4736C" baseAuthorityName="Jebb &amp; Cheek" baseAuthorityYear="1997" box="[189,311,1321,1347]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">Nepenthes</taxonomicName>
alata
</emphasis>
Blanco
<taxonomicName id="4C7E4D6CFF8EFC5CFE011375FC96736C" authority="Macfarlane" authorityName="Macfarlane" box="[476,773,1321,1347]" pageId="5" pageNumber="6" rank="variety" variety="ecristata">
var.
<emphasis id="B90AEAFDFF8EFC5CFDD31376FDE6736C" box="[526,629,1322,1347]" italics="true" pageId="5" pageNumber="6">ecristata</emphasis>
Macfarlane
</taxonomicName>
,
<taxonomicName id="4C7E4D6CFF8EFC5CFCCD1375FC2B736C" box="[784,952,1321,1347]" class="Magnoliopsida" family="Nepenthaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Nepenthaceae</taxonomicName>
.
<emphasis id="B90AEAFDFF8EFC5CFC1E1376FC48736C" box="[963,987,1322,1347]" italics="true" pageId="5" pageNumber="6">In</emphasis>
: Engler
<collectionCode id="ED6FAE2AFF8EFC5CFBE31375FBC4736C" box="[1086,1111,1321,1347]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="6" type="Herbarium">A</collectionCode>
. (ed.)
<emphasis id="B90AEAFDFF8EFC5CFB7D1376FAE5736C" box="[1184,1398,1321,1347]" italics="true" pageId="5" pageNumber="6">Das Pflanzenreich</emphasis>
Heft 36, 4, 3: 72 (1908). – Type:
<typeStatus id="54C5884DFF8EFC5CFDE71310FD3A7349" box="[570,681,1356,1382]" pageId="5" pageNumber="6" type="lectotype">lectotype</typeStatus>
,
<emphasis id="B90AEAFDFF8EFC5CFD6E1310FCE57348" bold="true" box="[691,886,1356,1383]" pageId="5" pageNumber="6">designated here</emphasis>
:
<collectingCountry id="F369767FFF8EFC5CFC591310FB987349" box="[900,1035,1356,1382]" name="Philippines" pageId="5" pageNumber="6">Philippines</collectingCountry>
, Mindanao, “Prov. of Misamis, Mount Malindang”,
<date id="FFC0102FFF8EFC5CFE69132CFDA173A5" box="[436,562,1392,1418]" pageId="5" pageNumber="6" value="1906-05">May 1906</date>
,
<emphasis id="B90AEAFDFF8EFC5CFD9F132DFBFE73A5" box="[578,1133,1392,1418]" italics="true" pageId="5" pageNumber="6">Mearns &amp; Hutchinson in Forest Bureau 4632</emphasis>
(lecto-:
<collectionCode id="ED6FAE2AFF8EFC5CFB0A132CFB6773A5" box="[1239,1268,1392,1418]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="5" pageNumber="6" type="Herbarium">K</collectionCode>
!; isolecto-:
<collectingCountry id="F369767FFF8EFC5CFF6013CFFF7F7382" box="[189,236,1427,1453]" name="Philippines" pageId="5" pageNumber="6">PH</collectingCountry>
!).
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF8EFC5EFF601389FC0673CE" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="8BC136EFFF8EFC5CFF601389FEDD73C0" blockId="5.[189,334,1493,1519]" box="[189,334,1493,1519]" pageId="5" pageNumber="6">
<heading id="D0898183FF8EFC5CFF601389FEDD73C0" bold="true" box="[189,334,1493,1519]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B90AEAFDFF8EFC5CFF601389FEDD73C0" bold="true" box="[189,334,1493,1519]" pageId="5" pageNumber="6">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8EFC5EFF601058FA8B7366" blockId="5.[189,1399,1540,2027]" lastBlockId="7.[189,1399,267,1354]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
Terrestrial shrub-climber, height unknown. Climbing stems terete to slightly angular,
<quantity id="4C869B0AFF8EFC5CFB621058FAB07031" box="[1215,1315,1540,1566]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="5" pageNumber="6" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
diam.; internodes
<quantity id="4C869B0AFF8EFC5CFE9E107BFE56706D" box="[323,453,1575,1602]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 mm</quantity>
long; axillary buds not evident; indumentum inconspicuous, persistent to the fifth internode from the apex, hairs translucent brown, simple or 2–3-armed from the base, hairs straight, variously angled from the horizontal, ca.
<quantity id="4C869B0AFF8EFC5CFCCD1032FCF870A6" box="[784,875,1646,1673]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="mm" value="0.1">0.1 mm</quantity>
long, covering ca. 5% of the surface except the axils (100% coverage) surface brown-black, matt. Leaves of rosette shoots thinly coriaceous, blade narrowly elliptic, 8–9 ×
<quantity id="4C869B0AFF8EFC5CFDF410E9FD0870FF" box="[553,667,1717,1744]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="5" pageNumber="6" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
; apex and base acute; longitudinal nerves 1–2 pairs, within
<quantity id="4C869B0AFF8EFC5CFAB510E9FF7870DC" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="2.0">2 mm</quantity>
of the margin, moderately conspicuous on both surfaces; pennate nerves at 90° from the midrib, numerous and moderately conspicuous; upper surface drying glossy pale brown, lower surface matt, mid-brown. Leaves of climbing stems as the rosette leaves, but blades suboblong or oblong-lanceolate 10–12.5 ×
<quantity id="4C869B0AFF8EFC5CFE9E111FFE5E7172" box="[323,461,1859,1885]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="5" pageNumber="6" unit="cm" value="3.5" valueMax="3.8" valueMin="3.2">3.2–3.8 cm</quantity>
; apex obtuse or acute; base obtuse; lower surface with sessile red glands ca.
<quantity id="4C869B0AFF8EFC5CFF60113AFE8971AE" box="[189,282,1894,1921]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
diam.; midrib 40–60% covered in patent, brown, simple or basally bifurcate-trifurcate hairs 0.1–0.3(–0.5) mm long; margin fringed, in young leaves, with hairs
<quantity id="4C869B0AFF8EFC5CFBFF11D6FB01718B" box="[1058,1170,1930,1957]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="6" unit="mm" value="0.25">0.25 mm</quantity>
long, pale-brown, 1–4-armed from the base. Petiole winged-canaliculate, 4–5 ×
<quantity id="4C869B0AFF8EFC5CFC7D11F1FC6771E7" box="[928,1012,1965,1992]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="cm" value="0.7">0.7 cm</quantity>
, wings patent; base clasping the stem for ⅓ to ½ its circumference, sometimes decurrent as an obtuse ridge to the node below. Lower pitchers unknown. Intermediate pitchers (tendrils uncoiled:
<emphasis id="B90AEAFDFF8CFC5EFCA61750FB56770A" box="[891,1221,267,294]" italics="true" pageId="7" pageNumber="8">Mearns &amp; Hutchinson 4632</emphasis>
)
<quantity id="4C869B0AFF8CFC5EFB0B1757FAE47709" box="[1238,1399,267,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.72" metricValueMin="1.25" pageId="7" pageNumber="8" unit="cm" value="14.85" valueMax="17.2" valueMin="12.5">12.5–17.2 cm</quantity>
long, ellipsoid in the basal third to half,
<quantity id="4C869B0AFF8CFC5EFD79176CFC877765" box="[676,788,304,331]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="5.7" metricValueMin="4.0" pageId="7" pageNumber="8" unit="cm" value="4.85" valueMax="5.7" valueMin="4.0">4–5.7 cm</quantity>
wide, constricted, more or less abruptly,
<quantity id="4C869B0AFF8CFC5EFADA176CFAE47765" box="[1287,1399,304,331]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="7" pageNumber="8" unit="cm" value="6.25" valueMax="7.5" valueMin="5.0">5–7.5 cm</quantity>
from the base into the subcylindrical upper part,
<quantity id="4C869B0AFF8CFC5EFCCA1709FC1B7740" box="[791,904,341,367]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="7" pageNumber="8" unit="cm" value="2.55" valueMax="3.0" valueMin="2.1">2.1–3 cm</quantity>
diam. dilating slightly towards the apex
<quantity id="4C869B0AFF8CFC5EFF601725FE8577BB" box="[189,278,377,404]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="7" pageNumber="8" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 cm</quantity>
diam.; outer surface strongly reticulated with raised nerves when dry, 2–5% covered in hairs of two
<typeStatus id="54C5884DFF8CFC5EFEC917C2FEC17797" box="[276,338,414,440]" pageId="7" pageNumber="8">types</typeStatus>
(
<figureCitation id="13452A6AFF8CFC5EFEBE17C1FE527797" box="[355,449,413,440]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="7" pageNumber="8">Fig. 1D</figureCitation>
), (1) large erect hairs
<quantity id="4C869B0AFF8CFC5EFD1017C1FCF97797" box="[717,874,413,440]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="7.5" metricValueMin="3.0" pageId="7" pageNumber="8" unit="mm" value="0.525" valueMax="0.75" valueMin="0.3">0.3–0.75 mm</quantity>
long, with a single, major, curved arm, and 1–2 much smaller erect arms, and (2) minute, 3–6-armed stellate hairs
<quantity id="4C869B0AFF8CFC5EFBD7179EFB3B77F2" box="[1034,1192,450,477]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="7" pageNumber="8" unit="mm" value="0.075" valueMax="0.1" valueMin="0.05">0.05–0.1 mm</quantity>
diam., which are more frequent, (ca. 4 per mm
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); surface covered throughout (6–10 per mm
<superScript id="7C0B9BA7FF8CFC5EFB9C17B9FBD977DC" attach="left" box="[1089,1098,485,499]" fontSize="6" pageId="7" pageNumber="8">2</superScript>
) with sessile, depressedglobose glands
<quantity id="4C869B0AFF8CFC5EFEA81457FD977409" box="[373,516,523,550]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="8" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
diam.; fringed wings reduced to ridges except in the ca.
<quantity id="4C869B0AFF8CFC5EFB7A1457FB687409" box="[1191,1275,523,550]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="25.0">25 mm</quantity>
below the peristome, widening to
<quantity id="4C869B0AFF8CFC5EFE0C146CFD877465" box="[465,532,560,586]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="mm" value="3.0">3 mm</quantity>
broad, with fringed elements
<quantity id="4C869B0AFF8CFC5EFCAF146CFC587465" box="[882,971,560,587]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="2.5">2.5 mm</quantity>
long,
<quantity id="4C869B0AFF8CFC5EFBC9146CFBE67465" box="[1044,1141,560,587]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="7" pageNumber="8" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
apart; mouth oblique, suborbicular, ovate, 3–4.8 ×
<quantity id="4C869B0AFF8CFC5EFDC61408FD367440" box="[539,677,596,623]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="4.5" metricValueMin="2.7" pageId="7" pageNumber="8" unit="cm" value="3.6" valueMax="4.5" valueMin="2.7">2.7–4.5 cm</quantity>
; apex with a column
<quantity id="4C869B0AFF8CFC5EFC6C1408FBB77440" box="[945,1060,596,623]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="7" pageNumber="8" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
long; peristome rounded to slightly flattened,
<quantity id="4C869B0AFF8CFC5EFE481425FD9D74BC" box="[405,526,633,660]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="7" pageNumber="8" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
wide, more or less even in width, ribs
<quantity id="4C869B0AFF8CFC5EFC361425FB1A74BC" box="[1003,1161,633,660]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="7" pageNumber="8" unit="mm" value="0.375" valueMax="0.5" valueMin="0.25">0.25–0.5 mm</quantity>
apart, conspicuous, about
<quantity id="4C869B0AFF8CFC5EFEDA14C2FEF17497" box="[263,354,670,696]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="0.1">0.1 mm</quantity>
high, outer edge lacking lobes, inner edge with very short teeth and conspicuous holes, teeth &lt;
<quantity id="4C869B0AFF8CFC5EFEC5149EFEE774F2" box="[280,372,706,733]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="0.1">0.1 mm</quantity>
long. Lid much smaller than the mouth, ovate, or broadly ovate, 25–35 ×
<quantity id="4C869B0AFF8CFC5EFB32149EFAE174F2" box="[1263,1394,706,733]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="7" pageNumber="8" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
, apex rounded to obtuse, base rounded to truncate; lower surface with a low basal ridge ca.
<quantity id="4C869B0AFF8CFC5EFB3214BBFAA0752E" box="[1263,1331,743,769]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="1.0">1 mm</quantity>
high,
<quantity id="4C869B0AFF8CFC5EFF601557FEBD7509" box="[189,302,779,806]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="7" pageNumber="8" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
long, either lacking a protruding appendage entirely (
<figureCitation id="13452A6AFF8CFC5EFC731557FB96750A" box="[942,1029,779,805]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="7" pageNumber="8">Fig. 1F</figureCitation>
) or with a modestly developed appendage
<quantity id="4C869B0AFF8CFC5EFE98156CFE3A7565" box="[325,425,816,842]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="8" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
high (
<figureCitation id="13452A6AFF8CFC5EFE27156CFDC97565" box="[506,602,816,842]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="7" pageNumber="8">Fig. 1H</figureCitation>
); nectar glands only slightly dimorphic, (1) midline nectar glands sparse, longitudinally elliptic, 0.5–0.7 ×
<quantity id="4C869B0AFF8CFC5EFD481508FCA07540" box="[661,819,852,879]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="7" pageNumber="8" unit="mm" value="0.175" valueMax="0.25" valueMin="0.1">0.1–0.25 mm</quantity>
, with a thin marginal rim (
<figureCitation id="13452A6AFF8CFC5EFBBB1508FB247541" box="[1126,1207,852,878]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="7" pageNumber="8">Fig. 1J</figureCitation>
), (2) outside the midline nectar glands circular (
<figureCitation id="13452A6AFF8CFC5EFDF31525FD1F75BC" box="[558,652,889,915]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="7" pageNumber="8">Fig. 1K</figureCitation>
), sparse, &lt;1 per mm
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, only 35–50 on each side of the midline, the largest scattered in the distal half,
<quantity id="4C869B0AFF8CFC5EFD5D15C1FD487597" box="[640,731,925,952]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="mm" value="0.5">0.5 mm</quantity>
diam., grading down to those of the marginal equatorial areas ca.
<quantity id="4C869B0AFF8CFC5EFEF6159EFE0675F3" box="[299,405,962,989]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="0.25">0.25 mm</quantity>
diam., and those at the attachment point with the peristome and the basal ridge and appendage,
<quantity id="4C869B0AFF8CFC5EFE9515BAFE3C722E" box="[328,431,998,1025]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="mm" value="0.15">0.15 mm</quantity>
diam.; sessile depressed-globose minute red glands
<quantity id="4C869B0AFF8CFC5EFBD615BBFB04722E" box="[1035,1175,999,1025]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="8" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
diam. are scattered over the surface at a density of 3–8 glands per mm
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; minute inconspicuous stellate hairs ca.
<quantity id="4C869B0AFF8CFC5EFB221257FAE47209" box="[1279,1399,1035,1062]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="8" unit="mm" value="0.075">0.075 mm</quantity>
diam. occur in an uneven,
<quantity id="4C869B0AFF8CFC5EFE201273FDE57265" box="[509,630,1071,1098]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="7" pageNumber="8" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
wide band, near the margin widening to
<quantity id="4C869B0AFF8CFC5EFBB81273FB527265" box="[1125,1217,1071,1098]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="mm" value="1.5">1.5 mm</quantity>
wide at the lid apex. Spur unbranched, curving downwards, stout at base and tapering to a slender apex, ca.
<quantity id="4C869B0AFF8CFC5EFB2F1208FAA67240" box="[1266,1333,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="mm" value="5.0">5 mm</quantity>
long, with scattered long, subpatent hairs
<quantity id="4C869B0AFF8CFC5EFDBB1225FD6772BC" box="[614,756,1145,1171]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="7" pageNumber="8" unit="mm" value="0.5" valueMax="0.7" valueMin="0.3">0.3–0.7 mm</quantity>
long (
<figureCitation id="13452A6AFF8CFC5EFC9D1225FC0272BC" box="[832,913,1145,1171]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="7" pageNumber="8">Fig. 1I</figureCitation>
). Upper pitchers (tendril coiled,
<emphasis id="B90AEAFDFF8CFC5EFAC81225FE3A7297" italics="true" pageId="7" pageNumber="8">Gaerlan et al. in PPI 10914</emphasis>
) resembling the intermediate pitchers, but fringed wings
<quantity id="4C869B0AFF8CFC5EFBBA12C1FB5F7297" box="[1127,1228,1181,1208]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="8" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
wide, fringed elements
<quantity id="4C869B0AFF8CFC5EFEF1129EFE1572F3" box="[300,390,1218,1245]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="2.5">2.5 mm</quantity>
long, (2–)
<quantity id="4C869B0AFF8CFC5EFE22129EFDF272F3" box="[511,609,1218,1245]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="7" pageNumber="8" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
apart, dilating to
<quantity id="4C869B0AFF8CFC5EFCED129EFC1372F3" box="[816,896,1218,1245]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="8" unit="cm" value="4.5">4.5 cm</quantity>
below the mouth; pitcher green, peristome maroon. Lid broadly ovate to suborbicular 32 ×
<quantity id="4C869B0AFF8CFC5EFD3112BAFCD2732E" box="[748,833,1254,1281]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="8" unit="mm" value="35.0">35 mm</quantity>
, lower surface with a basal ridge
<quantity id="4C869B0AFF8CFC5EFB1812BBFAA6732E" box="[1221,1333,1255,1281]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="7" pageNumber="8" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
long, ca.
<quantity id="4C869B0AFF8CFC5EFF391357FEB47309" box="[228,295,1291,1318]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="mm" value="2.0">2 mm</quantity>
high, bearing a central, symmetrical, protruding appendage 2 ×
<quantity id="4C869B0AFF8CFC5EFBD11357FBC27309" box="[1036,1105,1291,1318]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="mm" value="3.0">3 mm</quantity>
; nectar glands denser, ca. 110 on each side of the midline. Male and female inflorescences and infructescences unknown.
</paragraph>
<caption id="DF016667FF8DFC5FFF6010EBFAE671C4" ID-DOI="http://doi.org/10.5281/zenodo.3827693" ID-Zenodo-Dep="3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1719,1745]" targetBox="[296,1287,351,1645]" targetPageId="6">
<paragraph id="8BC136EFFF8DFC5FFF6010EBFAE671C4" blockId="6.[189,1399,1719,2027]" pageId="6" pageNumber="7">
<emphasis id="B90AEAFDFF8DFC5FFF6010EBFE9970FE" bold="true" box="[189,266,1719,1745]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
<taxonomicName id="4C7E4D6CFF8DFC5FFECD10EBFE4D70FE" authority="Cheek &amp; Jebb, 2013" authorityName="Cheek &amp; Jebb" authorityYear="2013" box="[272,478,1719,1745]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="kurata" status="sp. nov.">
<emphasis id="B90AEAFDFF8DFC5FFECD10EBFE4D70FE" box="[272,478,1719,1745]" italics="true" pageId="6" pageNumber="7">Nepenthes kurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2395786FF8DFC5FFE3910E4FDAD70FD" box="[484,574,1720,1746]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B90AEAFDFF8DFC5FFD9910EBFDCA70FE" bold="true" box="[580,601,1719,1745]" pageId="6" pageNumber="7">A</emphasis>
. Habit, climbing stem with upper pitcher.
<emphasis id="B90AEAFDFF8DFC5FFB9910EBFBCB70FE" bold="true" box="[1092,1112,1719,1745]" pageId="6" pageNumber="7">B</emphasis>
. Indumentum of midrib, lower surface of leaf-blade.
<emphasis id="B90AEAFDFF8DFC5FFDCA1086FDBF70DB" bold="true" box="[535,556,1754,1780]" pageId="6" pageNumber="7">C</emphasis>
. Indumentum of tendril.
<emphasis id="B90AEAFDFF8DFC5FFCBE1086FCEB70DB" bold="true" box="[867,888,1754,1780]" pageId="6" pageNumber="7">D</emphasis>
. Indumentum of outer pitcher surface.
<emphasis id="B90AEAFDFF8DFC5FFA811086FAE370DB" bold="true" box="[1372,1392,1754,1780]" pageId="6" pageNumber="7">E</emphasis>
. Lid, lower surface, showing nectar gland distribution on right (upper pitcher).
<emphasis id="B90AEAFDFF8DFC5FFBB010A2FB377138" bold="true" box="[1133,1188,1789,1816]" pageId="6" pageNumber="7">F–G</emphasis>
. Profiles of basal lid ridges, without appendage (
<emphasis id="B90AEAFDFF8DFC5FFDEC117DFDD07114" bold="true" box="[561,579,1825,1851]" pageId="6" pageNumber="7">F</emphasis>
), with appendage weakly developed (
<emphasis id="B90AEAFDFF8DFC5FFBD1117CFBB07115" bold="true" box="[1036,1059,1824,1850]" pageId="6" pageNumber="7">G</emphasis>
), and moderately developed (
<emphasis id="B90AEAFDFF8DFC5FFF1A1118FF4D7171" bold="true" box="[199,222,1860,1886]" pageId="6" pageNumber="7">H</emphasis>
).
<emphasis id="B90AEAFDFF8DFC5FFF2B1118FE927171" bold="true" box="[246,257,1860,1886]" pageId="6" pageNumber="7">I</emphasis>
Spur of intermediate pitcher.
<emphasis id="B90AEAFDFF8DFC5FFDBC1118FDE37171" bold="true" box="[609,624,1860,1886]" pageId="6" pageNumber="7">J</emphasis>
. Longitudinally elliptic nectar glands of lid midline.
<emphasis id="B90AEAFDFF8DFC5FFB3D1118FB647171" bold="true" box="[1248,1271,1860,1886]" pageId="6" pageNumber="7">K</emphasis>
. Orbicular nectar glands of lid, outside the midline.
<emphasis id="B90AEAFDFF8DFC5FFD47113BFD3D71AE" bold="true" box="[666,686,1895,1921]" pageId="6" pageNumber="7">L</emphasis>
. Peristome from above, short teeth and holes discernible.
<emphasis id="B90AEAFDFF8DFC5FFA8E113BFAFC71AE" bold="true" box="[1363,1391,1895,1921]" pageId="6" pageNumber="7">M</emphasis>
. Peristome viewed from inside pitcher.
<emphasis id="B90AEAFDFF8DFC5FFD5E11D6FD0B718B" bold="true" box="[643,664,1930,1956]" pageId="6" pageNumber="7">N</emphasis>
. Peristome transverse section, outer surface to right. A–E, J &amp; K from
<emphasis id="B90AEAFDFF8DFC5FFEC011F1FDD871E7" box="[285,587,1965,1992]" italics="true" pageId="6" pageNumber="7">Gaerlan et al. PPI 10911</emphasis>
; F–I, L–N from
<emphasis id="B90AEAFDFF8DFC5FFCCA11F2FBF771E8" box="[791,1124,1965,1992]" italics="true" pageId="6" pageNumber="7">Mearns &amp; Hutchinson 4632</emphasis>
. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown.
</paragraph>
</caption>
<paragraph id="8BC136EFFF8CFC5EFF60132EFE2773A3" blockId="7.[189,436,1394,1420]" box="[189,436,1394,1420]" pageId="7" pageNumber="8">
<heading id="D0898183FF8CFC5EFF60132EFE2773A3" bold="true" box="[189,436,1394,1420]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B90AEAFDFF8CFC5EFF60132EFE2773A3" bold="true" box="[189,436,1394,1420]" pageId="7" pageNumber="8">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8CFC5EFF6013FEFC0673CE" blockId="7.[189,1398,1442,1505]" pageId="7" pageNumber="8">
<collectingCountry id="F369767FFF8CFC5EFF6013FEFEFD7393" box="[189,366,1442,1468]" name="Philippines" pageId="7" pageNumber="8">PHILIPPINES</collectingCountry>
. Mindanao, Prov.
<collectingRegion id="49BAF80DFF8CFC5EFD9913FEFCA07393" box="[580,819,1442,1468]" country="Philippines" name="Misamis Occidental" pageId="7" pageNumber="8">Misamis Occidental</collectingRegion>
, S.E. slopes of Mt. Malindang, Lake Duminagat,
<date id="FFC0102FFF8CFC5EFF60139BFEA473CE" box="[189,311,1479,1505]" pageId="7" pageNumber="8" value="1993-05">May 1993</date>
,
<emphasis id="B90AEAFDFF8CFC5EFE9E139BFCB573CE" box="[323,806,1479,1505]" italics="true" pageId="7" pageNumber="8">Gaerlan, Sagcal &amp; Romero in PPI 10911</emphasis>
(BRIT!).
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF8CFC5EFF601055FB0B707B" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BC136EFFF8CFC5EFF601055FDF4700B" blockId="7.[189,615,1545,1572]" box="[189,615,1545,1572]" pageId="7" pageNumber="8">
<heading id="D0898183FF8CFC5EFF601055FDF4700B" bold="true" box="[189,615,1545,1572]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B90AEAFDFF8CFC5EFF601055FDF4700B" bold="true" box="[189,615,1545,1572]" pageId="7" pageNumber="8">Distribution, habitat &amp; phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8CFC5EFF601065FB0B707B" blockId="7.[189,1176,1593,1620]" box="[189,1176,1593,1620]" pageId="7" pageNumber="8">
<collectingCountry id="F369767FFF8CFC5EFF601065FED7707C" box="[189,324,1593,1619]" name="Philippines" pageId="7" pageNumber="8">Philippines</collectingCountry>
, Mindanao; evergreen forest, volcanic substrates. Elevation: ca.
<quantity id="4C869B0AFF8CFC5EFBE51066FB07707B" box="[1080,1172,1594,1620]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="8" unit="m" value="1400.0">1400 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF8CFC51FF601020FBD777B0" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="conservation">
<paragraph id="8BC136EFFF8CFC5EFF601020FE2670B8" blockId="7.[189,437,1660,1687]" box="[189,437,1660,1687]" pageId="7" pageNumber="8">
<heading id="D0898183FF8CFC5EFF601020FE2670B8" bold="true" box="[189,437,1660,1687]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B90AEAFDFF8CFC5EFF601020FE2670B8" bold="true" box="[189,437,1660,1687]" pageId="7" pageNumber="8">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF8CFC5EFF6010F0FDE971C4" blockId="7.[189,1399,1708,2027]" pageId="7" pageNumber="8">
<taxonomicName id="4C7E4D6CFF8CFC5EFF6010F0FE1F70E9" authority="Cheek &amp; Jebb, 2013" authorityName="Cheek &amp; Jebb" authorityYear="2013" box="[189,396,1708,1734]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="kurata" status="sp. nov.">
<emphasis id="B90AEAFDFF8CFC5EFF6010F0FE1F70E9" box="[189,396,1708,1734]" italics="true" pageId="7" pageNumber="8">Nepenthes kurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2395786FF8CFC5EFE4E10F1FE7D70E8" box="[403,494,1709,1735]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
is here assessed as Critically Endangered under Criterion D of
<bibRefCitation id="EFEF4B1EFF8CFC5EFB0410F0FAE570E9" author="IUCN" box="[1241,1398,1708,1734]" pageId="7" pageNumber="8" refId="ref14235" refString="IUCN 2012. IUCN Red List Categories and Criteria: Version 3.1. Prepared by the IUCN Species Survival Commission. IUCN, Gland, Switzerland / Cambridge, U. K." type="book" year="2012">IUCN (2012)</bibRefCitation>
since currently only two individuals, probably at a single location (as currently defined by IUCN) are known. This site, the ca. 6 ha crater Lake Duminagat, is within the ca. 50,000 ha Mt Malindang Range Natural Park of which at least 20,000 ha has been cleared for cultivation purposes, but which is a tentative World Heritage Site (http://whc.unesco.org/en/tentativelists/5029/, downloaded
<date id="FFC0102FFF8CFC5EFAC01162FF6E7152" pageId="7" pageNumber="8" value="2013-07-16">16 July 2013</date>
). In 2012 the Park was designated as an ASEAN Heritage Park (http://news.pia.gov.ph/index. php?article=1451343449808, downloaded
<date id="FFC0102FFF8CFC5EFD6E11DBFCDD718D" box="[691,846,1927,1954]" pageId="7" pageNumber="8" value="2013-07-16">16 July 2013</date>
). It is to be hoped that further investigation will discover additional individuals and locations for this species, decreasing its threat status, and increasing the likelihood that it can be protected.
</paragraph>
<paragraph id="8BC136EFFF83FC51FF601757FBD777B0" blockId="8.[189,1398,267,415]" pageId="8" pageNumber="9">
Since the terrain of Mt Malindang is reported as being rugged, with much forest surviving, there is every reason to hope that the species survives there, unlike
<taxonomicName id="4C7E4D6CFF83FC51FCE71768FAAD7761" authority="Cheek" authorityName="Cheek" box="[826,1342,308,334]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="robcantleyi">
<emphasis id="B90AEAFDFF83FC51FCE71768FBD17761" box="[826,1090,308,334]" italics="true" pageId="8" pageNumber="9">Nepenthes robcantleyi</emphasis>
Cheek (
<bibRefCitation id="EFEF4B1EFF83FC51FB781768FAA57761" author="Cheek M." box="[1189,1334,308,334]" pageId="8" pageNumber="9" pagination="677 - 681" refId="ref13249" refString="Cheek M. 2011. Nepenthes robcantleyi sp. nov. (Nepenthaceae) from Mindanao, Philippines. Nordic Journal of Botany 29: 677 - 681. http: // dx. doi. org / 10.1111 / j. 1756 - 1051.2011.01449. x" type="journal article" year="2011">Cheek 2011</bibRefCitation>
)
</taxonomicName>
also from Mindanao, which is already suspected to be extinct in the wild due to the almost total clearance of forest habitat at the single known wild location due to logging (
<bibRefCitation id="EFEF4B1EFF83FC51FC7817D9FBA477B0" author="Cheek M." box="[933,1079,389,415]" pageId="8" pageNumber="9" pagination="677 - 681" refId="ref13249" refString="Cheek M. 2011. Nepenthes robcantleyi sp. nov. (Nepenthaceae) from Mindanao, Philippines. Nordic Journal of Botany 29: 677 - 681. http: // dx. doi. org / 10.1111 / j. 1756 - 1051.2011.01449. x" type="journal article" year="2011">Cheek 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3646564FF83FC51FF601790FC0B71C4" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BC136EFFF83FC51FF601790FEBC77C9" blockId="8.[189,303,460,486]" box="[189,303,460,486]" pageId="8" pageNumber="9">
<heading id="D0898183FF83FC51FF601790FEBC77C9" bold="true" box="[189,303,460,486]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B90AEAFDFF83FC51FF601790FEBC77C9" bold="true" box="[189,303,460,486]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136EFFF83FC51FF60145CFB817444" blockId="8.[189,1399,512,619]" pageId="8" pageNumber="9">
The first
<taxonomicName id="4C7E4D6CFF83FC51FEF5145CFE317435" baseAuthorityName="Jebb &amp; Cheek" baseAuthorityYear="1997" box="[296,418,512,538]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="B90AEAFDFF83FC51FEF5145CFE317435" box="[296,418,512,538]" italics="true" pageId="8" pageNumber="9">Nepenthes</emphasis>
</taxonomicName>
taxa described from Mindanao, both of the
<emphasis id="B90AEAFDFF83FC51FC74145CFB9B7435" box="[937,1032,512,538]" italics="true" pageId="8" pageNumber="9">N. alata</emphasis>
group, (
<bibRefCitation id="EFEF4B1EFF83FC51FBB6145CFAFA7434" author="Cheek M. &amp; Jebb M." box="[1131,1385,512,539]" pageId="8" pageNumber="9" pagination="616 - 622" refId="ref13523" refString="Cheek M. &amp; Jebb M. 2013 d. Typification and redelimitation of Nepenthes alata Blanco with notes on the N. alata group, and N. negros sp. nov. from the Philippines. Nordic Journal of Botany 31: 616 - 622. http: // dx. doi. org / 10.1111 / j. 1756 - 1051.2012.00099. x" type="journal article" year="2013">Cheek &amp; Jebb 2013d</bibRefCitation>
), were
<emphasis id="B90AEAFDFF83FC51FF201475FEC8746D" box="[253,347,552,578]" italics="true" pageId="8" pageNumber="9">N. alata</emphasis>
<taxonomicName id="4C7E4D6CFF83FC51FEBF1475FCD8746D" authority="Macfarl." authorityName="Macfarl." box="[354,843,552,579]" pageId="8" pageNumber="9" rank="variety" variety="ecristata">
var.
<emphasis id="B90AEAFDFF83FC51FE481475FE6F746D" box="[405,508,553,578]" italics="true" pageId="8" pageNumber="9">ecristata</emphasis>
Macfarl. (
<bibRefCitation id="EFEF4B1EFF83FC51FDA81474FCD1746D" author="Macfarlane J. M." box="[629,834,552,578]" pageId="8" pageNumber="9" pagination="1 - 92" refId="ref14571" refString="Macfarlane J. M. 1908. Nepenthaceae. In: Engler A. (ed.) Das Pflanzenreich Heft 36, 4, 3: 1 - 92." type="journal article" year="1908">Macfarlane 1908</bibRefCitation>
)
</taxonomicName>
, based on
<emphasis id="B90AEAFDFF83FC51FC181475FA9E746D" box="[965,1293,552,579]" italics="true" pageId="8" pageNumber="9">Mearns &amp; Hutchinson 4632</emphasis>
from Mt Malindang, and
<taxonomicName id="4C7E4D6CFF83FC51FEA1140EFCFB7444" authority="Macfarl." authorityName="Macfarl." box="[380,872,593,619]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="copelandii">
<emphasis id="B90AEAFDFF83FC51FEA1140EFD887444" box="[380,539,593,619]" italics="true" pageId="8" pageNumber="9">N. copelandii</emphasis>
Macfarl. (
<bibRefCitation id="EFEF4B1EFF83FC51FD48140DFCF37444" author="Macfarlane J. M." box="[661,864,593,619]" pageId="8" pageNumber="9" pagination="1 - 92" refId="ref14571" refString="Macfarlane J. M. 1908. Nepenthaceae. In: Engler A. (ed.) Das Pflanzenreich Heft 36, 4, 3: 1 - 92." type="journal article" year="1908">Macfarlane 1908</bibRefCitation>
)
</taxonomicName>
from Mt Apo.
</paragraph>
<paragraph id="8BC136EFFF83FC51FF6014CBFB7F7231" blockId="8.[189,1399,663,1055]" pageId="8" pageNumber="9">
The first of these we here elevate to species level as
<taxonomicName id="4C7E4D6CFF83FC51FCE414C4FBCD749E" authority="Jebb &amp; Cheek" authorityName="Jebb &amp; Cheek" authorityYear="2013" box="[825,1118,663,690]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="kurata" status="sp. nov.">
<emphasis id="B90AEAFDFF83FC51FCE414C4FC38749E" box="[825,939,663,689]" italics="true" pageId="8" pageNumber="9">N. kurata</emphasis>
Jebb &amp; Cheek
</taxonomicName>
<taxonomicNameLabel id="A2395786FF83FC51FBB514C4FB57749D" box="[1128,1220,664,690]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
Previously we had considered this taxon to be synonymous with
<taxonomicName id="4C7E4D6CFF83FC51FCC8149DFA8B74F5" authority="Sh. Kurata" authorityName="Sh. Kurata" authorityYear="2011" box="[789,1304,704,730]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="mindanaoensis">
<emphasis id="B90AEAFDFF83FC51FCC8149DFC7A74F5" box="[789,1001,704,730]" italics="true" pageId="8" pageNumber="9">N. mindanaoensis</emphasis>
Sh.Kurata (
<bibRefCitation id="EFEF4B1EFF83FC51FBA4149CFA8374F5" author="Kurata S." box="[1145,1296,704,730]" pageId="8" pageNumber="9" pagination="30 - 35" refId="ref14381" refString="Kurata S. 2001. Two new species, Nepenthes pyriformis from West Sumatra (Indonesia) and Nepenthes mindanaoensis from Mindanao (Philippines). The Journal of the Insectivorous Plant Society. 52 (2): 30 - 35." type="journal article" year="2001">Kurata 2001</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="EFEF4B1EFF83FC51FAF7149CFEF5752C" author="Cheek M. &amp; Jebb M." pageId="8" pageNumber="9" pagination="616 - 622" refId="ref13523" refString="Cheek M. &amp; Jebb M. 2013 d. Typification and redelimitation of Nepenthes alata Blanco with notes on the N. alata group, and N. negros sp. nov. from the Philippines. Nordic Journal of Botany 31: 616 - 622. http: // dx. doi. org / 10.1111 / j. 1756 - 1051.2012.00099. x" type="journal article" year="2013">Cheek &amp; Jebb 2013d</bibRefCitation>
). The two taxa do have similarities in the overall shape of the upper pitchers, the weakly to moderately developed basal lid appendage and the sparse nectar glands of the lower lid surface. However they can be distinguished using the characters in
<tableCitation id="C6FC0354FF83FC51FC571566FC76757B" box="[906,997,826,852]" captionStart="Table 1" captionStartId="9.[189,258,268,294]" captionTargetId="graphics@9.[188,1399,327,894]" captionTargetPageId="9" captionText="Table 1. Diagnostic characters separating N. kurata sp. nov. and N. mindanaoensis." httpUri="http://table.plazi.org/id/DF016667FF82FC50FF601750FB1D7709" pageId="8" pageNumber="9" tableUuid="DF016667FF82FC50FF601750FB1D7709">Table 1</tableCitation>
. The number and extent of these features merit elevation from varietal to specific-level recognition in our opinion. Although the
<typeStatus id="54C5884DFF83FC51FA99153FFAE47552" box="[1348,1399,867,893]" pageId="8" pageNumber="9">type</typeStatus>
specimen has rosette stems and intermediate pitchers only, a second specimen, with climbing stems and upper pitchers,
<emphasis id="B90AEAFDFF83FC51FEB215EFFD9A75E2" box="[367,521,947,973]" italics="true" pageId="8" pageNumber="9">Gaerlan et al</emphasis>
.
<emphasis id="B90AEAFDFF83FC51FDCB15E8FD3F75E2" box="[534,684,947,973]" italics="true" pageId="8" pageNumber="9">in PPI 10911</emphasis>
came to light recently. It is from the
<typeStatus id="54C5884DFF83FC51FB8D15E8FB1075E1" box="[1104,1155,948,974]" pageId="8" pageNumber="9">type</typeStatus>
locality and matches the
<typeStatus id="54C5884DFF83FC51FF351580FE8875D9" box="[232,283,988,1014]" pageId="8" pageNumber="9">type</typeStatus>
in essential details.
<taxonomicName id="4C7E4D6CFF83FC51FDD81580FD4075D9" authority="Cheek &amp; Jebb, 2013" authorityName="Cheek &amp; Jebb" authorityYear="2013" box="[517,723,988,1014]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="kurata" status="sp. nov.">
<emphasis id="B90AEAFDFF83FC51FDD81580FD4075D9" box="[517,723,988,1014]" italics="true" pageId="8" pageNumber="9">Nepenthes kurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2395786FF83FC51FD071580FCA775D9" box="[730,820,988,1014]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
has the spot character within the
<emphasis id="B90AEAFDFF83FC51FB641580FAE575D9" box="[1209,1398,988,1014]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C7E4D6CFF83FC51FB641580FAA075D9" baseAuthorityName="Jebb &amp; Cheek" baseAuthorityYear="1997" box="[1209,1331,988,1014]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">Nepenthes</taxonomicName>
alata
</emphasis>
group of a small, more or less orbicular lid, only about half the length of the pitcher mouth.
</paragraph>
<paragraph id="8BC136EFFF83FC51FF601217FD9073FC" blockId="8.[189,1399,1099,1491]" pageId="8" pageNumber="9">
<bibRefCitation id="EFEF4B1EFF83FC51FF601217FE09724A" author="Macfarlane J. M." box="[189,410,1099,1125]" pageId="8" pageNumber="9" pagination="1 - 92" refId="ref14571" refString="Macfarlane J. M. 1908. Nepenthaceae. In: Engler A. (ed.) Das Pflanzenreich Heft 36, 4, 3: 1 - 92." type="journal article" year="1908">Macfarlane (1908)</bibRefCitation>
characterised his
<emphasis id="B90AEAFDFF83FC51FDAE1210FD40724A" box="[627,723,1099,1125]" italics="true" pageId="8" pageNumber="9">N. alata</emphasis>
<taxonomicName id="4C7E4D6CFF83FC51FD011210FCEB724A" box="[732,888,1100,1126]" pageId="8" pageNumber="9" rank="variety" variety="ecristata">
var.
<emphasis id="B90AEAFDFF83FC51FCCC1210FCEB724A" box="[785,888,1100,1125]" italics="true" pageId="8" pageNumber="9">ecristata</emphasis>
</taxonomicName>
by the lid appendage being either reduced or absent; the nectar glands being few, medium to large in size, and irregularly dispersed. Of the single specimen cited (
<emphasis id="B90AEAFDFF83FC51FE5A12C1FD4D7299" box="[391,734,1180,1207]" italics="true" pageId="8" pageNumber="9">Mearns &amp; Hutchinson 4632</emphasis>
), only two sheets (
<collectingCountry id="F369767FFF83FC51FC1212C0FC657299" box="[975,1014,1180,1206]" name="Philippines" pageId="8" pageNumber="9">PH</collectingCountry>
and K) have been found, both annotated in Macfarlane’s hand, each with two intermediate pitchers. Although all four pitchers share a basal ridge (
<figureCitation id="13452A6AFF83FC51FE9612B1FE197328" box="[331,394,1261,1287]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
F–H), only one of the four has an appendage, and that is only moderately developed as a convex emergence from the basal ridge (
<figureCitation id="13452A6AFF83FC51FD6D134AFC9E731F" box="[688,781,1302,1328]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="8" pageNumber="9">Fig. 1H</figureCitation>
). However a recent collection (
<emphasis id="B90AEAFDFF83FC51FBA2134AFA8F731F" box="[1151,1308,1302,1328]" italics="true" pageId="8" pageNumber="9">Gaerlan et al</emphasis>
.
<emphasis id="B90AEAFDFF83FC51FAF6134BFE957376" italics="true" pageId="8" pageNumber="9">in PPI 10977</emphasis>
) with upper pitchers, does show a developed appendage (
<figureCitation id="13452A6AFF83FC51FC7D1362FC647377" box="[928,1015,1342,1368]" captionStart="Fig" captionStartId="6.[189,232,1719,1745]" captionTargetBox="[296,1287,351,1645]" captionTargetId="figure@6.[272,1314,261,1668]" captionTargetPageId="6" captionText="Fig. 1. Nepenthes kurata sp. nov. A. Habit, climbing stem with upper pitcher. B. Indumentum of midrib, lower surface of leaf-blade. C. Indumentum of tendril. D. Indumentum of outer pitcher surface. E. Lid, lower surface, showing nectar gland distribution on right (upper pitcher). F–G. Profiles of basal lid ridges, without appendage (F), with appendage weakly developed (G), and moderately developed (H). I Spur of intermediate pitcher. J. Longitudinally elliptic nectar glands of lid midline. K. Orbicular nectar glands of lid, outside the midline. L. Peristome from above, short teeth and holes discernible. M. Peristome viewed from inside pitcher. N. Peristome transverse section, outer surface to right. A–E, J &amp; K from Gaerlan et al. PPI 10911; F–I, L–N from Mearns &amp; Hutchinson 4632. Scale bars: single = 1 mm; graduated single = 2 mm; double = 1 cm; graduated double = 5 cm. All drawn by Andrew Brown." figureDoi="http://doi.org/10.5281/zenodo.3827693" httpUri="https://zenodo.org/record/3827693/files/figure.png" pageId="8" pageNumber="9">Fig. 1E</figureCitation>
), suggesting the epithet
<emphasis id="B90AEAFDFF83FC51FACD1363FAE47377" box="[1296,1399,1343,1368]" italics="true" pageId="8" pageNumber="9">ecristata</emphasis>
“lacking a crest” is inappropriate. In any case, the Code demands priority only at one rank, so there is no requirement to adopt the varietal epithet at specific level, for which reason Macfarlane’s taxon is renamed as
<taxonomicName id="4C7E4D6CFF83FC51FEEF13E5FE3273FD" authority="Cheek &amp; Jebb, 2013" authorityName="Cheek &amp; Jebb" authorityYear="2013" box="[306,417,1464,1490]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="kurata" status="sp. nov.">
<emphasis id="B90AEAFDFF83FC51FEEF13E5FE3273FD" box="[306,417,1464,1490]" italics="true" pageId="8" pageNumber="9">N. kurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2395786FF83FC51FE7513E5FD9073FC" box="[424,515,1465,1491]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC136EFFF83FC51FF6013A3FE867045" blockId="8.[189,1399,1535,1642]" pageId="8" pageNumber="9">The upper pitchers also differ from the intermediate pitchers in the greater density of the nectar glands on the lower surface of the lid. However the shape, distribution and size of the nectar glands remain similar.</paragraph>
<paragraph id="8BC136EFFF83FC51FF6010CBFDE57105" blockId="8.[189,1398,1687,1835]" pageId="8" pageNumber="9">
This is the only known species of
<taxonomicName id="4C7E4D6CFF83FC51FD9B10CBFD53709E" baseAuthorityName="Jebb &amp; Cheek" baseAuthorityYear="1997" box="[582,704,1687,1713]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="B90AEAFDFF83FC51FD9B10CBFD53709E" box="[582,704,1687,1713]" italics="true" pageId="8" pageNumber="9">Nepenthes</emphasis>
</taxonomicName>
from Mt Malindang at this time, and it is therefore the most westerly known species of the genus in Mindanao.
<taxonomicName id="4C7E4D6CFF83FC51FCDA10E3FC4070F6" authority="Cheek &amp; Jebb, 2013" authorityName="Cheek &amp; Jebb" authorityYear="2013" box="[775,979,1727,1753]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="kurata" status="sp. nov.">
<emphasis id="B90AEAFDFF83FC51FCDA10E3FC4070F6" box="[775,979,1727,1753]" italics="true" pageId="8" pageNumber="9">Nepenthes kurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2395786FF83FC51FC05109CFBA270F5" box="[984,1073,1728,1754]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
is still incompletely known, full details on its ecology, altitudinal range, population density, inflorescences and infructescences, and ethnobotany remain to be discovered.
</paragraph>
<paragraph id="8BC136EFFF83FC51FF60110BFC0B71C4" blockId="8.[189,1399,1879,2027]" pageId="8" pageNumber="9">
The type specimens were collected by Major E.A. Mearns and W.J. Hutchinson in 1906 on the first recorded ascent of Mt Malindang, a volcanic mountain in the NW of Mindanao. Both sheets are annotated in the hand of Macfarlane as “
<emphasis id="B90AEAFDFF83FC51FD4F11F5FD7C71ED" box="[658,751,1960,1986]" italics="true" pageId="8" pageNumber="9">N. alata</emphasis>
<taxonomicName id="4C7E4D6CFF83FC51FD2B11F5FBB271ED" authority="Macfarlane" authorityName="Macfarlane" box="[758,1057,1960,1987]" pageId="8" pageNumber="9" rank="variety" variety="ecristata">
var.
<emphasis id="B90AEAFDFF83FC51FCF511F5FC1C71ED" box="[808,911,1961,1986]" italics="true" pageId="8" pageNumber="9">ecristata</emphasis>
Macfarlane
</taxonomicName>
”, and either could be selected as
<typeStatus id="54C5884DFF83FC51FF00118DFEDA71C4" box="[221,329,2001,2027]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
of that name. The K sheet is accordingly selected.
</paragraph>
</subSubSection>
</treatment>
</document>
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