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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3690.1.1" ID-GBIF-Dataset="c8670273-a22c-4b4a-972d-38a43cd66f86" ID-ISSN="1175-5326" ID-Zenodo-Dep="6329811" ID-ZooBank="0874D336-BA8C-4266-AA50-633167C816F3" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1629874133667" checkinUser="felipe" docAuthor="PLANT, ADRIAN R." docDate="2013" docId="03D787D0FF8BFF9FFF05FD68FB3EFE24" docLanguage="en" docName="zt03690p098.pdf" docOrigin="Zootaxa 3690 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3690.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.10:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="10" docTitle="Hybos steatopygus PLANT 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="66" masterDocId="FFEEFFA8FFCBFFDDFF92FFB5FF99FF88" masterDocTitle="The genus Hybos Meigen (Diptera: Empidoidea: Hybotidae) in Thailand" masterLastPageNumber="98" masterPageNumber="1" pageNumber="65" updateTime="1679570708792" updateUser="felipe">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Hybos Meigen (Diptera: Empidoidea: Hybotidae) in Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>PLANT, ADRIAN R.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-07-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3690</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>98</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.3690.1.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3690.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">c8670273-a22c-4b4a-972d-38a43cd66f86</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">6329811</mods:identifier>
<mods:identifier type="ZooBank">0874D336-BA8C-4266-AA50-633167C816F3</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6339128" ID-GBIF-Taxon="193889248" ID-Zenodo-Dep="6339128" LSID="urn:lsid:plazi:treatment:03D787D0FF8BFF9FFF05FD68FB3EFE24" httpUri="http://treatment.plazi.org/id/03D787D0FF8BFF9FFF05FD68FB3EFE24" lastPageId="66" lastPageNumber="66" pageId="64" pageNumber="65">
<subSubSection box="[151,475,733,759]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph blockId="64.[150,475,733,792]" box="[151,475,733,759]" pageId="64" pageNumber="65">
<heading bold="true" box="[151,475,733,759]" fontSize="11" level="2" pageId="64" pageNumber="65" reason="4">
<emphasis bold="true" box="[151,475,733,759]" pageId="64" pageNumber="65">
<taxonomicName ID-CoL="8CDKJ" authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[151,375,733,759]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="steatopygus" status="sp. nov.">
<emphasis bold="true" box="[151,375,733,759]" italics="true" pageId="64" pageNumber="65">Hybos steatopygus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[381,475,733,759]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[150,435,768,792]" pageId="64" pageNumber="65" type="description">
<paragraph blockId="64.[150,475,733,792]" box="[150,435,768,792]" pageId="64" pageNumber="65">
(
<figureCitation box="[158,316,768,792]" captionStart="FIGURES 168–173" captionStartId="65.[151,263,1807,1830]" captionTargetBox="[170,1417,872,1763]" captionTargetId="figure-442@65.[151,1436,855,1786]" captionTargetPageId="65" captionText="FIGURES 168–173. Hybos steatopygus sp. nov. 168. ♂ terminalia, dorsal view. 169. hypandrium, ventral view. 170. left surstylus, lateral view. 171. right surstylus, oblique lateral view. 172–173. ♀ terminalia. 172. ventral view. 173. lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10." figureDoi="http://doi.org/10.5281/zenodo.6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="64" pageNumber="65">Figs 168–173</figureCitation>
,
<figureCitation box="[327,371,768,792]" captionStart="FIGURES 269–277" captionStartId="89.[151,264,1945,1968]" captionTargetBox="[244,1325,201,1898]" captionTargetId="figure-80@89.[219,1367,165,1924]" captionTargetPageId="89" captionText="FIGURES 269–277. Distribution of Hybos spp. in Thailand plotted at a resolution of 1° of latitude and longitude. 269. H. serratus Yang &amp; Yang. 270. H. shamshevi sp. nov. 271. H. sinclairi sp. nov. 272. H. songbai sp. nov. 273. H. steatopygus sp. nov. 274. H. stigmaticus sp. nov. 275. H. subapicalis sp. nov. 276. H. tetricus sp. nov. 277. H. thaosaeo sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6329911" httpUri="https://zenodo.org/record/6329911/files/figure.png" pageId="64" pageNumber="65">273</figureCitation>
,
<figureCitation box="[383,426,768,792]" captionStart="FIGURES 299–306" captionStartId="94.[151,263,1907,1930]" captionTargetBox="[218,1370,188,1882]" captionTargetId="figure-81@94.[177,1410,165,1918]" captionTargetPageId="94" captionText="FIGURES 299–306. Phenology of Hybos spp. monthly abundance (as % of total number of individuals captured) is plotted against month of capture. 299. H. steatopygus sp. nov. 300. H. stigmaticus sp. nov. 301. H. subapicalis sp. nov. 302. H. thaosaeo sp. nov. 303. H. thepkaisoni sp. nov. 304. H. tilokarati sp. nov. 305. H. xishuangbannaensis Yang &amp; Yang. 306. H. zhejiangensis Yang &amp; Yang." figureDoi="http://doi.org/10.5281/zenodo.6329919" httpUri="https://zenodo.org/record/6329919/files/figure.png" pageId="64" pageNumber="65">299</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="64" pageNumber="65" type="materials_examined">
<paragraph blockId="64.[151,1436,839,2005]" pageId="64" pageNumber="65">
<emphasis bold="true" box="[151,325,839,864]" pageId="64" pageNumber="65">Type material.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3698642361" collectedFrom="montane forest" collectingDate="2008-05-21" collectingDateMax="2008-05-28" collectingDateMin="2008-05-21" collectingMethod="Malaise trap" collectionCode="QSBG" collectorName="P. Wongchai" country="Thailand" elevation="2174" latitude="20.05915" location="Doi Phahompok National Park" longLatPrecision="1" longitude="99.14253" pageId="64" pageNumber="65" specimenCode="T6100" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province" typeStatus="holotype">
<emphasis bold="true" box="[332,491,839,864]" pageId="64" pageNumber="65">
<typeStatus box="[332,491,839,864]" pageId="64" pageNumber="65">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount box="[499,520,841,864]" pageId="64" pageNumber="65" type="male">♂</specimenCount>
:
<collectingCountry box="[535,682,840,864]" name="Thailand" pageId="64" pageNumber="65">THAILAND</collectingCountry>
,
<collectingRegion box="[694,938,840,864]" country="Thailand" name="Chiang Mai" pageId="64" pageNumber="65">Chiang Mai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03D787D0FF8BFF9FFF05FD68FB3EFE24:8EA1601DFF8BFF9DFC27FCFDFA8BFCE8" box="[949,1298,840,864]" country="Thailand" latitude="20.05915" longLatPrecision="1" longitude="99.14253" name="Doi Phahompok National Park" pageId="64" pageNumber="65" stateProvince="Chiang Mai Province">Doi Phahompok National Park</location>
,
<location LSID="urn:lsid:plazi:treatment:03D787D0FF8BFF9FFF05FD68FB3EFE24:8EA1601DFF8BFF9DFA8CFCFDFA0AFCE8" box="[1310,1427,840,864]" country="Thailand" latitude="20.05915" longLatPrecision="1" longitude="99.14253" name="Kiewlom 1" pageId="64" pageNumber="65" stateProvince="Chiang Mai Province">Kiewlom1</location>
/
<collectedFrom box="[151,327,876,900]" pageId="64" pageNumber="65">montane forest</collectedFrom>
,
<geoCoordinate box="[344,475,876,900]" degrees="20" direction="north" minutes="3.549" orientation="latitude" pageId="64" pageNumber="65" precision="1" value="20.05915">20°3.549'N</geoCoordinate>
,
<geoCoordinate box="[492,619,876,900]" degrees="99" direction="east" minutes="8.552" orientation="longitude" pageId="64" pageNumber="65" precision="1" value="99.14253">99°8.552'E</geoCoordinate>
,
<quantity box="[637,730,876,900]" metricMagnitude="3" metricUnit="m" metricValue="2.174" pageId="64" pageNumber="65" unit="m" value="2174.0">
<elevation box="[637,730,876,900]" metricMagnitude="3" metricUnit="m" metricValue="2.174" pageId="64" pageNumber="65" unit="m" value="2174.0">2174 m</elevation>
</quantity>
,
<date box="[748,903,876,900]" pageId="64" pageNumber="65" value="2008-05-21" valueMax="2008-05-28" valueMin="2008-05-21">
<collectingDate box="[748,903,876,900]" pageId="64" pageNumber="65" value="2008-05-21" valueMax="2008-05-28" valueMin="2008-05-21">21–28.v.2008</collectingDate>
</date>
,
<collectingMethod box="[920,1068,876,900]" pageId="64" pageNumber="65">Malaise trap</collectingMethod>
,
<collectorName box="[1085,1217,876,900]" pageId="64" pageNumber="65">P.Wongchai</collectorName>
[
<specimenCode box="[1237,1316,876,900]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="64" pageNumber="65">T6100</specimenCode>
] (
<collectionCode box="[1344,1424,876,900]" pageId="64" pageNumber="65">QSBG</collectionCode>
).
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3698642427" box="[151,1236,911,936]" pageId="64" pageNumber="65">
<emphasis bold="true" box="[151,315,911,936]" pageId="64" pageNumber="65">
<typeStatus box="[151,315,911,936]" pageId="64" pageNumber="65">PARATYPES</typeStatus>
</emphasis>
:
<specimenCount box="[329,378,912,936]" pageId="64" pageNumber="65" type="male">16♂</specimenCount>
,
<specimenCount box="[392,427,912,936]" pageId="64" pageNumber="65" type="female">6♀</specimenCount>
, same data as holotype;
<specimenCount box="[703,752,912,936]" pageId="64" pageNumber="65" type="male">20♂</specimenCount>
,
<specimenCount box="[766,801,912,936]" pageId="64" pageNumber="65" type="female">3♀</specimenCount>
,
<date box="[815,984,912,936]" pageId="64" pageNumber="65" value="2008-05-28" valueMax="2008-06-07" valueMin="2008-05-28">
<collectingDate box="[815,984,912,936]" pageId="64" pageNumber="65" value="2008-05-28" valueMax="2008-06-07" valueMin="2008-05-28">28.v.-7.vi.2008</collectingDate>
</date>
(
<collectionCode box="[999,1075,912,936]" pageId="64" pageNumber="65">QSBG</collectionCode>
and
<collectionCode box="[1129,1224,912,936]" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="64" pageNumber="65" type="Museum">NMWC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="64.[151,1436,839,2005]" pageId="64" pageNumber="65">
<emphasis bold="true" box="[199,451,948,973]" pageId="64" pageNumber="65">
<typeStatus box="[199,445,948,973]" isOtherTypeStatus="true" pageId="64" pageNumber="65">Additional material</typeStatus>
.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3698642386" box="[467,993,948,972]" pageId="64" pageNumber="65">
<specimenCount box="[467,502,948,972]" pageId="64" pageNumber="65" type="female">3♀</specimenCount>
, same data as holotype,
<date box="[821,993,948,972]" pageId="64" pageNumber="65" value="2008-05-28" valueMax="2008-06-07" valueMin="2008-05-28">
<collectingDate box="[821,993,948,972]" pageId="64" pageNumber="65" value="2008-05-28" valueMax="2008-06-07" valueMin="2008-05-28">28.v.-7.vi.2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642514" pageId="64" pageNumber="65">
<specimenCount box="[1014,1049,948,972]" pageId="64" pageNumber="65" type="male">3♂</specimenCount>
,
<specimenCount box="[1072,1107,948,972]" pageId="64" pageNumber="65" type="female">3♀</specimenCount>
, Doi Phaluang,
<geoCoordinate box="[1315,1431,948,972]" degrees="20" direction="north" minutes="1.06" orientation="latitude" pageId="64" pageNumber="65" precision="9" value="20.017666">20°1.06'N</geoCoordinate>
,
<geoCoordinate box="[151,278,984,1008]" degrees="99" direction="east" minutes="9.581" orientation="longitude" pageId="64" pageNumber="65" precision="1" value="99.15968">99°9.581'E</geoCoordinate>
,
<quantity box="[289,377,984,1008]" metricMagnitude="3" metricUnit="m" metricValue="1.449" pageId="64" pageNumber="65" unit="m" value="1449.0">1449 m</quantity>
,
<date box="[389,544,984,1008]" pageId="64" pageNumber="65" value="2008-05-21" valueMax="2008-05-28" valueMin="2008-05-21">21–28.v.2008</date>
,
<date box="[555,706,984,1008]" pageId="64" pageNumber="65" value="2008-06-07" valueMax="2008-06-14" valueMin="2008-06-07">7–14.vi.2008</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3698642555" pageId="64" pageNumber="65">
<specimenCount box="[718,753,984,1008]" pageId="64" pageNumber="65" type="male">2♂</specimenCount>
,
<specimenCount box="[767,802,984,1008]" pageId="64" pageNumber="65" type="female">5♀</specimenCount>
, Mae Fang Hotspring,
<geoCoordinate box="[1063,1207,984,1008]" degrees="19" direction="north" minutes="57.961" orientation="latitude" pageId="64" pageNumber="65" precision="1" value="19.966017">19°57.961'N</geoCoordinate>
,
<geoCoordinate box="[1219,1346,984,1008]" degrees="99" direction="east" minutes="9.355" orientation="longitude" pageId="64" pageNumber="65" precision="1" value="99.155914">99°9.355'E</geoCoordinate>
,
<quantity box="[1358,1431,984,1008]" metricMagnitude="2" metricUnit="m" metricValue="5.6899999999999995" pageId="64" pageNumber="65" unit="m" value="569.0">
<elevation box="[1358,1431,984,1008]" metricMagnitude="2" metricUnit="m" metricValue="5.6899999999999995" pageId="64" pageNumber="65" unit="m" value="569.0">569 m</elevation>
</quantity>
,
<date box="[151,299,1020,1044]" pageId="64" pageNumber="65" value="2008-06-07" valueMax="2008-06-14" valueMin="2008-06-07">
<collectingDate box="[151,299,1020,1044]" pageId="64" pageNumber="65" value="2008-06-07" valueMax="2008-06-14" valueMin="2008-06-07">7–14.vi.2008</collectingDate>
</date>
(
<collectionCode box="[313,389,1020,1044]" pageId="64" pageNumber="65">QSBG</collectionCode>
and
<collectionCode box="[444,539,1020,1044]" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="64" pageNumber="65" type="Museum">NMWC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="64" pageNumber="65" type="etymology">
<paragraph blockId="64.[151,1436,839,2005]" pageId="64" pageNumber="65">
<emphasis bold="true" box="[199,334,1056,1080]" pageId="64" pageNumber="65">Etymology.</emphasis>
A concatenation of Latin and Greek meaning ‘fat buttocks’, in reference to the globose male terminalia.
</paragraph>
</subSubSection>
<subSubSection pageId="64" pageNumber="65" type="diagnosis">
<paragraph blockId="64.[151,1436,839,2005]" pageId="64" pageNumber="65">
<emphasis bold="true" box="[199,321,1127,1152]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
A species with black thorax, hind femur black with a yellow apex, mid and front femora yellow but darkened dorsally. The mid tibia is clear yellow with only short hairs apart from very long and strong bristles dorsally near the middle. A dorsal seta is present on the postpedicel of the antenna. The male abdomen is constricted basally with segments 2 and 3 slightly elongated.
</paragraph>
</subSubSection>
<subSubSection lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="description">
<paragraph blockId="64.[151,1436,839,2005]" lastBlockId="65.[151,1436,151,802]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
<emphasis bold="true" box="[199,345,1271,1296]" pageId="64" pageNumber="65">Description.</emphasis>
<emphasis box="[356,413,1272,1296]" italics="true" pageId="64" pageNumber="65">Male</emphasis>
: body length 3.0–
<quantity box="[631,723,1272,1296]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="64" pageNumber="65" unit="mm" value="3.5">3.5 mm</quantity>
.
<emphasis box="[738,798,1272,1296]" italics="true" pageId="64" pageNumber="65">Head</emphasis>
subshining black, thinly dusted; face yellowish black, especially on lower part. All setae dark, with some paler hairs on lower occiput behind mouth. Antenna greyish black; postpedicel ovate in lateral view, ~2X long as wide, 1 fine dorsal seta present; stylus bare, 4–5X long as postpedicel, apical 0.25 narrower, paler in certain lights. Mouthparts dark yellowish brown, palpus very narrow, with 1 distinct dark apical seta and a few smaller setae below.
<emphasis box="[860,941,1416,1440]" italics="true" pageId="64" pageNumber="65">Thorax</emphasis>
with ground colour black with pleura often somewhat paler brownish black, postalar callus yellowish anterolaterally, scutellum narrowly yellowish basally; scutum, scutellum and pleura subshining, rather thickly and coarsely dusted greyish with faint indications of more shining stripes along line of dc near anterior margin of scutum; acr ~2–4-serial, dc uniserial, small, fine except 1 strong posterior dc, not continued over prescutellar depression; 1 strong (upper) and 1 weaker (lower) npl; pa weak; scutellum with pair of strong black bristles and several fine marginal hairs.
<emphasis box="[1045,1098,1597,1620]" italics="true" pageId="64" pageNumber="65">Legs</emphasis>
with coxae black; F
<subScript attach="left" box="[1342,1350,1611,1625]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
and F
<subScript attach="right" box="[1428,1436,1611,1625]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
yellowish to yellowish brown; F
<subScript attach="left" box="[535,543,1650,1664]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
usually rather darker than F
<subScript attach="left" box="[886,894,1650,1664]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
, sometimes blackish but both darker dorsally (especially F
<subScript attach="left" box="[294,302,1689,1703]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
); T
<subScript attach="left" box="[343,351,1689,1703]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
yellowish brown, T
<subScript attach="left" box="[575,583,1689,1703]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
and T
<subScript attach="left" box="[654,662,1689,1703]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
yellow; anterior tarsus brownish, mid and posterior tarsi with basal 2 segments yellow, distal segments brownish. Coxae with mostly pale hairs, conspicuously long behind C
<subScript attach="right" box="[1347,1355,1727,1741]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
; a few dark bristles anteroapically, especially on C
<subScript attach="left" box="[659,667,1766,1780]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
. F
<subScript attach="left" box="[700,708,1766,1780]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
and F
<subScript attach="left" box="[786,794,1766,1780]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
without distinct pv fringe, entirely short-haired, apical circlet with only small fine setae. F
<subScript attach="none" box="[546,554,1805,1819]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
rather slender, only slightly inflated, widest 0.6–0.8 from base where ~1.5–1.8 as deep as at base; ventral spines black, comprising av series of ~9–11 bristles (as long as limb is deep on basal 0.8, much shorter and spine-like on distal 0.2) behind which on basal 0.3 are 3–4 somewhat shorter bristles; pv fringe with 2–3 black bristles on distal 0.2, otherwise with fine pale hairs becoming shorter proximally; 1 rather proclinate, distinctly curved, long bristle anteriorly at ~0.8 and usually another at ~0.5 from base. T
<subScript attach="left" box="[1280,1288,1952,1966]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
with distinct dark bristles dorsally at 0.6 from base and near tip, pv fringe of fine hairs hardly longer than limb is deep. T
<subScript attach="left" box="[1351,1359,1991,2005]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
with 1 very strong, black bristle ~0.9X long as limb dorsally at 0.25–0.3 from base and 1 similar bristle ventrally slightly beyond; apical circlet of 5–6 black bristles rather short except 1 anterior ~0.7X long as MT
<subScript attach="left" box="[1199,1207,203,217]" fontSize="6" pageId="65" pageNumber="66">2</subScript>
. T
<subScript attach="left" box="[1239,1247,203,217]" fontSize="6" pageId="65" pageNumber="66">3</subScript>
slightly swollen apically, rather thinly covered with longish pubescent hairs; distinct black dorsal seta at 0.4 from base; no apical circlet apart from 1 distinct anterior and 1 pd setae. Front tarsi with short hairs apart from slightly longer paired dorsoapical seta on each segment and 1 short, rather stout ventrobasal on MT
<subScript attach="left" box="[1020,1028,314,328]" fontSize="6" pageId="65" pageNumber="66">1</subScript>
. MT
<subScript attach="left" box="[1084,1092,314,328]" fontSize="6" pageId="65" pageNumber="66">2</subScript>
with short erect ventral bristle near base with 2–3 similar but shorter setae beyond, otherwise with short hairs. MT
<subScript attach="left" box="[1095,1103,352,366]" fontSize="6" pageId="65" pageNumber="66">2</subScript>
short spinose ventrally.
<emphasis box="[1378,1435,338,361]" italics="true" pageId="65" pageNumber="66">Wing</emphasis>
membrane tinged brown, paler posteriorly; veins brown; stigma faint, brownish, reaching 0.8X distance between end of R
<subScript attach="left" box="[248,256,427,441]" fontSize="6" pageId="65" pageNumber="66">1</subScript>
and R
<subScript attach="left" box="[330,354,427,441]" fontSize="6" pageId="65" pageNumber="66">2+3</subScript>
. Squamae with white fringes. Halter white.
<emphasis box="[853,959,412,436]" italics="true" pageId="65" pageNumber="66">Abdomen</emphasis>
subshining brownish black, thinly dusted; pregenital segments with all setae pale; segments 2 and 3 slightly constricted, elongate (somewhat longer than wide). Terminalia (
<figureCitation box="[365,522,487,511]" captionStart="FIGURES 168–173" captionStartId="65.[151,263,1807,1830]" captionTargetBox="[170,1417,872,1763]" captionTargetId="figure-442@65.[151,1436,855,1786]" captionTargetPageId="65" captionText="FIGURES 168–173. Hybos steatopygus sp. nov. 168. ♂ terminalia, dorsal view. 169. hypandrium, ventral view. 170. left surstylus, lateral view. 171. right surstylus, oblique lateral view. 172–173. ♀ terminalia. 172. ventral view. 173. lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10." figureDoi="http://doi.org/10.5281/zenodo.6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="65" pageNumber="66">Figs 168–171</figureCitation>
) globose, much wider than abdomen, black with setae dark. Right surstylus (
<figureCitation captionStart="FIGURES 168–173" captionStartId="65.[151,263,1807,1830]" captionTargetBox="[170,1417,872,1763]" captionTargetId="figure-442@65.[151,1436,855,1786]" captionTargetPageId="65" captionText="FIGURES 168–173. Hybos steatopygus sp. nov. 168. ♂ terminalia, dorsal view. 169. hypandrium, ventral view. 170. left surstylus, lateral view. 171. right surstylus, oblique lateral view. 172–173. ♀ terminalia. 172. ventral view. 173. lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10." figureDoi="http://doi.org/10.5281/zenodo.6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="65" pageNumber="66">Fig. 171</figureCitation>
) with long, narrow outer process and broader, internally directed inner process. Left surstylus (
<figureCitation box="[1255,1350,523,547]" captionStart="FIGURES 168–173" captionStartId="65.[151,263,1807,1830]" captionTargetBox="[170,1417,872,1763]" captionTargetId="figure-442@65.[151,1436,855,1786]" captionTargetPageId="65" captionText="FIGURES 168–173. Hybos steatopygus sp. nov. 168. ♂ terminalia, dorsal view. 169. hypandrium, ventral view. 170. left surstylus, lateral view. 171. right surstylus, oblique lateral view. 172–173. ♀ terminalia. 172. ventral view. 173. lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10." figureDoi="http://doi.org/10.5281/zenodo.6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="65" pageNumber="66">Fig. 170</figureCitation>
) broad, with short, curving, spine-like, dorsal process and longer, blunt, curving, ventral process. Hypandrium (
<figureCitation box="[1331,1428,559,583]" captionStart="FIGURES 168–173" captionStartId="65.[151,263,1807,1830]" captionTargetBox="[170,1417,872,1763]" captionTargetId="figure-442@65.[151,1436,855,1786]" captionTargetPageId="65" captionText="FIGURES 168–173. Hybos steatopygus sp. nov. 168. ♂ terminalia, dorsal view. 169. hypandrium, ventral view. 170. left surstylus, lateral view. 171. right surstylus, oblique lateral view. 172–173. ♀ terminalia. 172. ventral view. 173. lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10." figureDoi="http://doi.org/10.5281/zenodo.6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="65" pageNumber="66">Fig. 169</figureCitation>
) narrow, with narrow strongly curved process arising from left margin, with long curving bristles arising from base.
<emphasis box="[151,235,631,655]" italics="true" pageId="65" pageNumber="66">Female</emphasis>
. Differing from male as follows. Slightly smaller, body length
<quantity box="[937,1075,631,655]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="65" pageNumber="66" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.1 mm</quantity>
. F
<subScript attach="left" box="[1101,1109,646,660]" fontSize="6" pageId="65" pageNumber="66">3</subScript>
with ventral spines uniserial, longer (1–2 about middle 2X long as limb is deep). Abdomen not constricted basally, segments 2 and 3 wider than long; shorter-haired but sternite 7 with long pale bristles on posterior margin. Terminalia (
<figureCitation box="[1204,1367,706,730]" captionStart="FIGURES 168–173" captionStartId="65.[151,263,1807,1830]" captionTargetBox="[170,1417,872,1763]" captionTargetId="figure-442@65.[151,1436,855,1786]" captionTargetPageId="65" captionText="FIGURES 168–173. Hybos steatopygus sp. nov. 168. ♂ terminalia, dorsal view. 169. hypandrium, ventral view. 170. left surstylus, lateral view. 171. right surstylus, oblique lateral view. 172–173. ♀ terminalia. 172. ventral view. 173. lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10." figureDoi="http://doi.org/10.5281/zenodo.6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="65" pageNumber="66">Figs 172, 173</figureCitation>
) with tergite 8 broad, almost encircling abdomen, with long setae on posterior margin. Sternite 8 (
<figureCitation box="[1207,1305,742,766]" captionStart="FIGURES 168–173" captionStartId="65.[151,263,1807,1830]" captionTargetBox="[170,1417,872,1763]" captionTargetId="figure-442@65.[151,1436,855,1786]" captionTargetPageId="65" captionText="FIGURES 168–173. Hybos steatopygus sp. nov. 168. ♂ terminalia, dorsal view. 169. hypandrium, ventral view. 170. left surstylus, lateral view. 171. right surstylus, oblique lateral view. 172–173. ♀ terminalia. 172. ventral view. 173. lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10." figureDoi="http://doi.org/10.5281/zenodo.6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="65" pageNumber="66">Fig. 172</figureCitation>
) bare, very narrow, irregularly upcurved distally. Sternite 10, small, Y-shaped, rather densely pilose.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6329879" ID-Zenodo-Dep="6329879" httpUri="https://zenodo.org/record/6329879/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,263,1807,1830]" targetBox="[170,1417,872,1763]" targetPageId="65">
<paragraph blockId="65.[151,1437,1807,1891]" pageId="65" pageNumber="66">
<emphasis bold="true" box="[151,368,1807,1830]" pageId="65" pageNumber="66">FIGURES 168–173.</emphasis>
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[378,571,1808,1830]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="steatopygus" status="sp. nov.">
<emphasis box="[378,571,1808,1830]" italics="true" pageId="65" pageNumber="66">Hybos steatopygus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[581,720,1807,1830]" pageId="65" pageNumber="66">
<taxonomicNameLabel box="[581,665,1808,1829]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
168.
</emphasis>
♂ terminalia, dorsal view.
<emphasis bold="true" box="[1012,1056,1807,1830]" pageId="65" pageNumber="66">169.</emphasis>
hypandrium, ventral view.
<emphasis bold="true" box="[1348,1393,1807,1830]" pageId="65" pageNumber="66">170.</emphasis>
left surstylus, lateral view.
<emphasis bold="true" box="[388,433,1838,1861]" pageId="65" pageNumber="66">171.</emphasis>
right surstylus, oblique lateral view.
<emphasis bold="true" box="[819,914,1838,1861]" pageId="65" pageNumber="66">172–173.</emphasis>
♀ terminalia.
<emphasis bold="true" box="[1066,1110,1838,1861]" pageId="65" pageNumber="66">172.</emphasis>
ventral view.
<emphasis bold="true" box="[1258,1303,1838,1861]" pageId="65" pageNumber="66">173.</emphasis>
lateral view. S8= sternite 8, S10 = sternite 10, T8 = tergite 8, T10 = tergite 10.
</paragraph>
</caption>
<subSubSection lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="discussion">
<paragraph blockId="65.[151,1436,1930,2027]" lastBlockId="66.[151,1438,151,428]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<emphasis bold="true" box="[199,324,1930,1955]" pageId="65" pageNumber="66">Comment.</emphasis>
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[331,541,1931,1955]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="steatopygus" status="sp. nov.">
<emphasis box="[331,541,1931,1955]" italics="true" pageId="65" pageNumber="66">Hybos steatopygus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[548,638,1931,1955]" pageId="65" pageNumber="66">
<taxonomicNameLabel box="[548,638,1931,1955]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from Doi Phahompok, in the Daen Lao Range along the border with
<collectingCountry box="[291,367,1967,1990]" name="Myanmar" pageId="65" pageNumber="66">Burma</collectingCountry>
in the far north of
<collectingCountry box="[588,687,1966,1990]" name="Thailand" pageId="65" pageNumber="66">Thailand</collectingCountry>
(
<figureCitation box="[704,801,1966,1990]" captionStart="FIGURES 269–277" captionStartId="89.[151,264,1945,1968]" captionTargetBox="[244,1325,201,1898]" captionTargetId="figure-80@89.[219,1367,165,1924]" captionTargetPageId="89" captionText="FIGURES 269–277. Distribution of Hybos spp. in Thailand plotted at a resolution of 1° of latitude and longitude. 269. H. serratus Yang &amp; Yang. 270. H. shamshevi sp. nov. 271. H. sinclairi sp. nov. 272. H. songbai sp. nov. 273. H. steatopygus sp. nov. 274. H. stigmaticus sp. nov. 275. H. subapicalis sp. nov. 276. H. tetricus sp. nov. 277. H. thaosaeo sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6329911" httpUri="https://zenodo.org/record/6329911/files/figure.png" pageId="65" pageNumber="66">Fig. 273</figureCitation>
) and 48 (79 %) of 61 known individuals were captured above
<quantity box="[224,316,2002,2026]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="65" pageNumber="66" unit="m" value="2000.0">2,000 m</quantity>
in moist hill evergreen forest. Despite extensive trapping efforts on Doi Phahompok,
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[1270,1435,2003,2026]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="steatopygus" status="sp. nov.">
<emphasis box="[1270,1435,2003,2026]" italics="true" pageId="65" pageNumber="66">H. steatopygus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[151,243,152,176]" pageId="66" pageNumber="67">
<taxonomicNameLabel box="[151,243,152,176]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has only been caught in a brief period between late May and early June (
<figureCitation box="[1095,1196,151,175]" captionStart="FIGURES 299–306" captionStartId="94.[151,263,1907,1930]" captionTargetBox="[218,1370,188,1882]" captionTargetId="figure-81@94.[177,1410,165,1918]" captionTargetPageId="94" captionText="FIGURES 299–306. Phenology of Hybos spp. monthly abundance (as % of total number of individuals captured) is plotted against month of capture. 299. H. steatopygus sp. nov. 300. H. stigmaticus sp. nov. 301. H. subapicalis sp. nov. 302. H. thaosaeo sp. nov. 303. H. thepkaisoni sp. nov. 304. H. tilokarati sp. nov. 305. H. xishuangbannaensis Yang &amp; Yang. 306. H. zhejiangensis Yang &amp; Yang." figureDoi="http://doi.org/10.5281/zenodo.6329919" httpUri="https://zenodo.org/record/6329919/files/figure.png" pageId="66" pageNumber="67">Fig. 299</figureCitation>
), coincident with the early part of the wet season.
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[486,699,188,212]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="steatopygus" status="sp. nov.">
<emphasis box="[486,699,188,212]" italics="true" pageId="66" pageNumber="67">Hybos steatopygus</emphasis>
</taxonomicName>
<emphasis bold="true" box="[709,801,188,212]" pageId="66" pageNumber="67">
<taxonomicNameLabel box="[709,801,188,212]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can easily be separated from other
<collectingCountry box="[1215,1314,188,212]" name="Thailand" pageId="66" pageNumber="67">Thailand</collectingCountry>
species in which the hind femur is black with a yellow apex (
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[733,815,225,248]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="men" status="sp. nov.">
<emphasis box="[733,815,225,248]" italics="true" pageId="66" pageNumber="67">H. men</emphasis>
</taxonomicName>
<emphasis bold="true" box="[823,914,224,248]" pageId="66" pageNumber="67">
<taxonomicNameLabel box="[823,914,224,248]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName box="[930,1098,224,248]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="particularis">
<emphasis box="[930,1098,224,248]" italics="true" pageId="66" pageNumber="67">H. particularis</emphasis>
</taxonomicName>
;
<taxonomicName box="[1114,1285,224,248]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="ancistroides">
<emphasis box="[1114,1285,224,248]" italics="true" pageId="66" pageNumber="67">H. ancistroides</emphasis>
</taxonomicName>
) by having a yellow mid femur which is darkened dorsally along its length. Based on morphology of the hypandrium, the species is probably quite closely related to
<taxonomicName authority="Yang &amp; Grootaert, 2006" authorityName="Yang &amp; Grootaert" authorityYear="2006" box="[657,1099,295,320]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bifurcatus">
<emphasis box="[657,807,296,320]" italics="true" pageId="66" pageNumber="67">H. bifurcatus</emphasis>
<bibRefCitation author="Yang, D. &amp; Grootaert, P." box="[818,1099,295,320]" pageId="66" pageNumber="67" pagination="161 - 163" refId="ref81619" refString="Yang, D. &amp; Grootaert, P. (2006) Two new species of Hybos (Diptera, Empidoidea, Hybotidae) from China. Biologia, Bratislava, 61, 161 - 163. http: // dx. doi. org / 10.3157 / 0013 - 872 x (2006) 117 [219: tnsoed] 2.0. co; 2" type="journal article" year="2006">Yang &amp; Grootaert, 2006</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Yang &amp; Grootaert, 2006" authorityName="Yang &amp; Grootaert" authorityYear="2006" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mengqingae">
<emphasis box="[1161,1336,296,320]" italics="true" pageId="66" pageNumber="67">H. mengqingae</emphasis>
<bibRefCitation author="Yang, D. &amp; Grootaert, P." pageId="66" pageNumber="67" pagination="161 - 163" refId="ref81619" refString="Yang, D. &amp; Grootaert, P. (2006) Two new species of Hybos (Diptera, Empidoidea, Hybotidae) from China. Biologia, Bratislava, 61, 161 - 163. http: // dx. doi. org / 10.3157 / 0013 - 872 x (2006) 117 [219: tnsoed] 2.0. co; 2" type="journal article" year="2006">Yang &amp; Grootaert, 2006</bibRefCitation>
</taxonomicName>
from the Nanling Mountains in southeastern
<collectingCountry box="[860,927,332,356]" name="China" pageId="66" pageNumber="67">China</collectingCountry>
and perhaps also
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[1139,1270,332,356]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="sinclairi" status="sp. nov.">
<emphasis box="[1139,1270,332,356]" italics="true" pageId="66" pageNumber="67">H. sinclairi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1279,1372,332,356]" pageId="66" pageNumber="67">
<taxonomicNameLabel box="[1279,1372,332,356]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<collectingCountry box="[151,254,367,391]" name="Thailand" pageId="66" pageNumber="67">Thailand</collectingCountry>
. However,
<taxonomicName authorityName="Yang &amp; Grootaert" authorityYear="2006" box="[391,543,368,392]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bifurcatus">
<emphasis box="[391,543,368,392]" italics="true" pageId="66" pageNumber="67">H. bifurcatus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Yang &amp; Grootaert" authorityYear="2006" box="[609,785,368,392]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mengqingae">
<emphasis box="[609,785,368,392]" italics="true" pageId="66" pageNumber="67">H. mengqingae</emphasis>
</taxonomicName>
have entirely black legs and surstyli of very different morphology and
<taxonomicName authority="PLANT, 2013" authorityName="PLANT" authorityYear="2013" box="[342,469,404,428]" class="Insecta" family="Hybotidae" genus="Hybos" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="sinclairi" status="sp. nov.">
<emphasis box="[342,469,404,428]" italics="true" pageId="66" pageNumber="67">H. sinclairi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[476,566,404,428]" pageId="66" pageNumber="67">
<taxonomicNameLabel box="[476,566,404,428]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by the characters given in the key.
</paragraph>
</subSubSection>
</treatment>
</document>
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