treatments-xml/data/03/D7/C1/03D7C10CFF9DF629A8892614FB1B5189.xml

<document id="3B79A59555879586EA7DDA12FE41F88F" ID-CLB-Dataset="9158" ID-DOI="10.11646/zootaxa.4838.1.4" ID-GBIF-Dataset="dfdc4db4-0057-411a-8ea7-a1c437c44b56" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403631" ID-ZooBank="5F56E910-9BCA-418A-A331-A5BF56C2F67C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609351495522" checkinUser="plazi" docAuthor="Rivera-Correa, Mauricio &amp; Daza, Juan M." docDate="2020" docId="03D7C10CFF9DF629A8892614FB1B5189" docLanguage="en" docName="zootaxa.4838.1.4.pdf" docOrigin="Zootaxa 4838 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pristimantis zorro Rivera-Correa &amp; Daza 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="90" masterDocId="FFEEB974FF9FF621A81E204FFFD45344" masterDocTitle="Out of the blue: A new rain frog species of the genus Pristimantis (Anura Craugastoridae) from the northern Cordillera Central in Colombia" masterLastPageNumber="101" masterPageNumber="83" pageNumber="85" updateTime="1701998628814" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E28373A43E42CC0DE450585CE4DF15BC">Out of the blue: A new rain frog species of the genus Pristimantis (Anura Craugastoridae) from the northern Cordillera Central in Colombia</mods:title>
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<mods:namePart id="D0F69CD9AC87B16D3E7E02F977097355">Rivera-Correa, Mauricio</mods:namePart>
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<mods:affiliation id="2BD98E965D69F8095D3B01AF847C50B3">Grupo Herpetológico de Antioquia, Instituto de Biología, Universidad de Antioquia, Medellín, Colombia. https: // orcid. org / 0000 - 0001 - 5033 - 5480 &amp; Semillero de Investigación en Biodiversidad de Anfibios (BIO), Seccional Oriente, Universidad de Antioquia</mods:affiliation>
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<mods:namePart id="E7DDB10833F6AC4C2EAB5B706FB67291">Daza, Juan M.</mods:namePart>
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<mods:affiliation id="E94CB721FA1920507BCF92C2481B93AA">Grupo Herpetológico de Antioquia, Instituto de Biología, Universidad de Antioquia, Medellín, Colombia. https: // orcid. org / 0000 - 0001 - 5033 - 5480</mods:affiliation>
<mods:nameIdentifier id="206CCF63972117A921733B62B39CF215" type="email">jumadaza@gmail.com</mods:nameIdentifier>
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<taxonomicName id="4C7E0B99FF9DF623A8892614FEA55531" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[151,369,1627,1653]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="85" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF9DF623A8892614FEA55531" bold="true" box="[151,369,1627,1653]" italics="true" pageId="2" pageNumber="85">Pristimantis zorro</emphasis>
</taxonomicName>
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(
<figureCitation id="13456C9FFF9DF623A8812630FEDE55DD" box="[159,266,1663,1689]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,1615,1639]" captionStartId-1="5.[151,250,1792,1816]" captionStartId-2="8.[151,250,1501,1525]" captionTargetBox-0="[267,1320,205,1584]" captionTargetBox-1="[158,1424,1004,1760]" captionTargetBox-2="[156,1429,744,1474]" captionTargetId-0="figure-33@4.[256,1331,181,1592]" captionTargetId-1="figure-405@5.[151,1436,993,1769]" captionTargetId-2="figure-316@8.[151,1437,741,1478]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="8" captionText-0="FIGURE 2. Adult males of Pristimantis zorro sp. nov. in life: (A) MHUA-A 8813, SVL 20.1 mm, holotype; (B) MHUA-A 8814, SVL 20.5 mm, paratype; (C) MHUA-A 8815, SVL 20.3 mm, paratype, adult male; (D) MHUA-A 8816, SVL 19.5 mm, paratype; (E) MHUA-A 8817, SVL 21.5 mm, paratype; (F) MHUA-A 11165, SVL 20.1 mm, paratype; (G) MHUA-A 11166, SVL 21.3 mm, paratype; (H) MHUA-A 11167, SVL 20.8 mm, paratype. Photos by Adriana Restrepo, Carlos Marín and JMD." captionText-1="FIGURE 3. Paratype of Pristimantis zorro sp. nov. in preservative. MHUA-A 11165, SVL 20.1 mm, adult male. Photos by MRC." captionText-2="FIGURE 4. Advertisement call of Pristimantis zorro sp. nov. (MHUA-A 8813, holotype). (A). Spectrogram; (B) Oscillogram; (C) Power spectrum. Photo by JMD." figureDoi-0="http://doi.org/10.5281/zenodo.4403635" figureDoi-1="http://doi.org/10.5281/zenodo.4403639" figureDoi-2="http://doi.org/10.5281/zenodo.4403641" httpUri-0="https://zenodo.org/record/4403635/files/figure.png" httpUri-1="https://zenodo.org/record/4403639/files/figure.png" httpUri-2="https://zenodo.org/record/4403641/files/figure.png" pageId="2" pageNumber="85">Figs. 2–4</figureCitation>
)
</paragraph>
<paragraph id="8BC1701AFF9DF623A8892688FB545441" blockId="2.[151,1437,1735,2013]" pageId="2" pageNumber="85">
<materialsCitation id="3B167A47FF9DF623A8892688FB545441" ID-GBIF-Occurrence="3006039302" collectingDate="2015-05-21" collectionCode="MHUA-A" collectorName="Vereda El Tronco &amp; Juan M. Daza." country="Colombia" elevation="1860" latitude="6.30199" location="Vereda El Tronco" longLatPrecision="1" longitude="-75.14165" municipality="Municipio de Guatape" pageId="2" pageNumber="85" specimenCode="MHUA-A 8813" specimenCount="1" stateProvince="Antioquia" typeStatus="holotype">
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.
</emphasis>
<specimenCode id="DBD8D861FF9DF623A9102688FE1755A5" box="[270,451,1735,1761]" collectionCode="MHUA-A" pageId="2" pageNumber="85">MHUA-A 8813</specimenCode>
, adult male, 
<collectingCountry id="F369308AFF9DF623AA492688FD1C55A5" box="[599,712,1735,1761]" name="Colombia" pageId="2" pageNumber="85">Colombia</collectingCountry>
, 
<collectingRegion id="49BABEF8FF9DF623AAC82688FBC655A5" box="[726,1042,1735,1761]" country="Colombia" name="Antioquia" pageId="2" pageNumber="85">Departamento de Antioquia</collectingRegion>
, 
<collectingMunicipality id="6BA5EA60FF9DF623AC012688FAF455A5" box="[1055,1312,1735,1761]" pageId="2" pageNumber="85">Municipio de Guatapé</collectingMunicipality>
, 
<collectorName id="268B15CCFF9DF623AD332688FF335441" pageId="2" pageNumber="85">
<location id="8EA126C1FF9DF623AD332688FF335441" LSID="urn:lsid:plazi:treatment:03D7C10CFF9DF629A8892614FB1B5189:8EA126C1FF9DF623AD332688FF335441" country="Colombia" latitude="6.30199" longLatPrecision="1" longitude="-75.14165" municipality="Municipio de Guatape" name="Vereda El Tronco" pageId="2" pageNumber="85" stateProvince="Antioquia">Vereda El Tronco</location>
</collectorName>
(
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, 
<geoCoordinate id="EE4A16DDFF9DF623A97E26A5FE025441" box="[352,470,1770,1797]" degrees="75.14165" direction="west" orientation="longitude" pageId="2" pageNumber="85" precision="1" value="-75.14165">-75.14165</geoCoordinate>
; 
<elevation id="00539729FF9DF623A9FF26A4FDA65442" box="[481,626,1771,1798]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="2" pageNumber="85" unit="m" value="1860.0">
<quantity id="4C86DDFFFF9DF623A9FF26A4FDE25441" box="[481,566,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="2" pageNumber="85" unit="m" value="1860.0">1860 m</quantity>
a.s.l.
</elevation>
), collected on 
<date id="FFC056DAFF9DF623AB0626A4FC605441" box="[792,948,1770,1797]" pageId="2" pageNumber="85" value="2015-05-21">
<collectingDate id="EF84AF32FF9DF623AB0626A4FC605441" box="[792,948,1770,1797]" pageId="2" pageNumber="85" value="2015-05-21">May 21, 2015</collectingDate>
</date>
by 
<collectorName id="268B15CCFF9DF623ABC026A4FB545441" box="[990,1152,1771,1797]" pageId="2" pageNumber="85">Juan M. Daza.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BC1701AFF9DF623A8D92740FB675499" blockId="2.[151,1437,1735,2013]" pageId="2" pageNumber="85">
<materialsCitation id="3B167A47FF9DF623A8D92740FBAF546D" ID-GBIF-Occurrence="3006039303" box="[199,1147,1807,1833]" collectionCode="MHUA-A" pageId="2" pageNumber="85" specimenCode="MHUA-A 8814-8817" specimenCount="2" typeStatus="paratype">
<emphasis id="B90AAC08FF9DF623A8D92740FE94546C" box="[199,320,1807,1832]" italics="true" pageId="2" pageNumber="85">
<typeStatus id="54C5CEB8FF9DF623A8D92740FEE8546C" box="[199,316,1807,1832]" pageId="2" pageNumber="85" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DBD8D861FF9DF623A9562740FDEB546D" box="[328,575,1807,1833]" collectionCode="MHUA-A" pageId="2" pageNumber="85">MHUA-A 8814–8817</specimenCode>
adult males; all collected along with the 
<typeStatus id="54C5CEB8FF9DF623AC0B2740FBAF546D" box="[1045,1147,1807,1833]" pageId="2" pageNumber="85" type="holotype">holotype</typeStatus>
</materialsCitation>
. 
<materialsCitation id="3B167A47FF9DF623AC982740FBC25409" ID-GBIF-Occurrence="3006039305" collectingDate="2017-04-09" collectionCode="MHUA-A" location="Carlos Marin" pageId="2" pageNumber="85" specimenCode="MHUA-A 11165-11167" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBD8D861FF9DF623AC982740FA4C546D" box="[1158,1432,1806,1833]" collectionCode="MHUA-A" pageId="2" pageNumber="85">MHUA-A 11165–11167</specimenCode>
, adult males; all collected at the type locality on 
<date id="FFC056DAFF9DF623AAA6277CFC855409" box="[696,849,1843,1869]" pageId="2" pageNumber="85" value="2017-04-09">
<collectingDate id="EF84AF32FF9DF623AAA6277CFC855409" box="[696,849,1843,1869]" pageId="2" pageNumber="85" value="2017-04-09">April 9, 2017</collectingDate>
</date>
by 
<location id="8EA126C1FF9DF623AB63277CFBC25409" LSID="urn:lsid:plazi:treatment:03D7C10CFF9DF629A8892614FB1B5189:8EA126C1FF9DF623AB63277CFBC25409" box="[893,1046,1843,1869]" name="Carlos Marin" pageId="2" pageNumber="85">Carlos Marin</location>
</materialsCitation>
. 
<materialsCitation id="3B167A47FF9DF623AC3C277CFB055435" ID-GBIF-Occurrence="3006039301" collectingDate="2016-10-15" collectionCode="MHUA-A" country="Mauritius" location="Mauricio" pageId="2" pageNumber="85" specimenCode="MHUA-A 12092" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBD8D861FF9DF623AC3C277CFB365409" box="[1058,1250,1843,1869]" collectionCode="MHUA-A" pageId="2" pageNumber="85">MHUA-A 12092</specimenCode>
adult male; collected at the type locality on 
<date id="FFC056DAFF9DF623A9FF2718FD725435" box="[481,678,1878,1905]" pageId="2" pageNumber="85" value="2016-10-15">
<collectingDate id="EF84AF32FF9DF623A9FF2718FD725435" box="[481,678,1878,1905]" pageId="2" pageNumber="85" value="2016-10-15">October 15, 2016</collectingDate>
</date>
by 
<collectingCountry id="F369308AFF9DF623AACC2718FCED5435" box="[722,825,1879,1905]" name="Mauritius" pageId="2" pageNumber="85">Mauricio</collectingCountry>
Rivera-Correa and Daniela Murillo
</materialsCitation>
. 
<materialsCitation id="3B167A47FF9DF623ACC32718FDCF54FD" ID-GBIF-Occurrence="3006039306" collectingDate="2019-11-11" collectionCode="MHUA-A" country="Mauritius" county="Carlos Rivera" location="Rivera-Correa" municipality="Alejandro Ramirez" pageId="2" pageNumber="85" specimenCode="MHUA-A 12093" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBD8D861FF9DF623ACC32718FA485435" box="[1245,1436,1879,1905]" collectionCode="MHUA-A" pageId="2" pageNumber="85">MHUA-A 12093</specimenCode>
adult male; collected at the type locality on 
<date id="FFC056DAFF9DF623AA962734FCB354D1" box="[648,871,1915,1941]" pageId="2" pageNumber="85" value="2019-11-11">
<collectingDate id="EF84AF32FF9DF623AA962734FCB354D1" box="[648,871,1915,1941]" pageId="2" pageNumber="85" value="2019-11-11">November 11, 2019</collectingDate>
</date>
by 
<collectingCountry id="F369308AFF9DF623AB8D2734FC2E54D1" box="[915,1018,1915,1941]" name="Mauritius" pageId="2" pageNumber="85">Mauricio</collectingCountry>
<location id="8EA126C1FF9DF623AC1C2734FB7154D1" LSID="urn:lsid:plazi:treatment:03D7C10CFF9DF629A8892614FB1B5189:8EA126C1FF9DF623AC1C2734FB7154D1" box="[1026,1189,1915,1941]" country="Mauritius" county="Carlos Rivera" municipality="Alejandro Ramirez" name="Rivera-Correa" pageId="2" pageNumber="85">Rivera-Correa</location>
, 
<location id="8EA126C1FF9DF623ACAF2734FAB454D1" LSID="urn:lsid:plazi:treatment:03D7C10CFF9DF629A8892614FB1B5189:8EA126C1FF9DF623ACAF2734FAB454D1" box="[1201,1376,1915,1941]" country="Mauritius" county="Carlos Rivera" municipality="Alejandro Ramirez" name="Jaime Culebras" pageId="2" pageNumber="85">Jaime Culebras</location>
, 
<collectingMunicipality id="6BA5EA60FF9DF623AD742734FE9154FD" pageId="2" pageNumber="85">Alejandro Ramírez</collectingMunicipality>
and 
<collectingCounty id="62A00896FF9DF623A96327D0FDCF54FD" box="[381,539,1951,1977]" pageId="2" pageNumber="85">Carlos Rivera</collectingCounty>
</materialsCitation>
. 
<materialsCitation id="3B167A47FF9DF623AA3827D0FB645499" ID-GBIF-Occurrence="3006039304" collectingDate="2020-02-08" collectionCode="MHUA-A" country="Mauritius" county="Hader Correa" location="Rivera-Correa" municipality="Diego Botero" pageId="2" pageNumber="85" specimenCode="MHUA-A 12094" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBD8D861FF9DF623AA3827D0FD3154FD" box="[550,741,1951,1977]" collectionCode="MHUA-A" pageId="2" pageNumber="85">MHUA-A 12094</specimenCode>
adult male; collected at the type locality on 
<date id="FFC056DAFF9DF623ACC527D0FA4854FD" box="[1243,1436,1951,1977]" pageId="2" pageNumber="85" value="2020-02-08">
<collectingDate id="EF84AF32FF9DF623ACC527D0FA4854FD" box="[1243,1436,1951,1977]" pageId="2" pageNumber="85" value="2020-02-08">February 8, 2020</collectingDate>
</date>
by 
<collectingCountry id="F369308AFF9DF623A8A4278CFEF55499" box="[186,289,1987,2013]" name="Mauritius" pageId="2" pageNumber="85">Mauricio</collectingCountry>
<location id="8EA126C1FF9DF623A936278CFE1F5499" LSID="urn:lsid:plazi:treatment:03D7C10CFF9DF629A8892614FB1B5189:8EA126C1FF9DF623A936278CFE1F5499" box="[296,459,1987,2013]" country="Mauritius" county="Hader Correa" municipality="Diego Botero" name="Rivera-Correa" pageId="2" pageNumber="85">Rivera-Correa</location>
, 
<location id="8EA126C1FF9DF623A9C8278CFD715499" LSID="urn:lsid:plazi:treatment:03D7C10CFF9DF629A8892614FB1B5189:8EA126C1FF9DF623A9C8278CFD715499" box="[470,677,1987,2013]" country="Mauritius" county="Hader Correa" municipality="Diego Botero" name="Khristian Venegas" pageId="2" pageNumber="85">Khristian Venegas</location>
, 
<location id="8EA126C1FF9DF623AAAE278CFCEE5499" LSID="urn:lsid:plazi:treatment:03D7C10CFF9DF629A8892614FB1B5189:8EA126C1FF9DF623AAAE278CFCEE5499" box="[688,826,1987,2013]" country="Mauritius" county="Hader Correa" municipality="Diego Botero" name="Eliza Patino" pageId="2" pageNumber="85">Eliza Patiño</location>
, 
<collectingMunicipality id="6BA5EA60FF9DF623AB58278CFC095499" box="[838,989,1987,2013]" pageId="2" pageNumber="85">Diego Botero</collectingMunicipality>
and 
<collectingCounty id="62A00896FF9DF623AC0A278CFB645499" box="[1044,1200,1987,2013]" pageId="2" pageNumber="85">Hader Correa</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF012092FF9CF622A889271DFB655492" ID-DOI="http://doi.org/10.5281/zenodo.4403633" ID-Zenodo-Dep="4403633" httpUri="https://zenodo.org/record/4403633/files/figure.png" pageId="3" pageNumber="86" startId="3.[151,250,1874,1898]" targetBox="[234,1356,188,1840]" targetPageId="3">
<paragraph id="8BC1701AFF9CF622A889271DFB655492" blockId="3.[151,1436,1874,2006]" pageId="3" pageNumber="86">
<emphasis id="B90AAC08FF9CF622A889271DFEC1542E" bold="true" box="[151,277,1874,1898]" pageId="3" pageNumber="86">FIGURE 1.</emphasis>
(A) Maximum likelihood tree (-lnL=104073.3) depicting evolutionary relationships within 
<taxonomicName id="4C7E0B99FF9CF622ACD6271CFA90542E" box="[1224,1348,1875,1898]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="86" phylum="Chordata" rank="genus">
<emphasis id="B90AAC08FF9CF622ACD6271CFA90542E" box="[1224,1348,1875,1898]" italics="true" pageId="3" pageNumber="86">Pristimantis</emphasis>
</taxonomicName>
inferred from a partitioned analysis using 4103 sites, four genomic regions, and 188 terminals. The complete tree is provided as appendix 1. (B) Uncorrected genetic distances between species most closely related to 
<taxonomicName id="4C7E0B99FF9CF622ABBD27D4FB8F54F6" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[931,1115,1947,1970]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="86" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF9CF622ABBD27D4FB8F54F6" box="[931,1115,1947,1970]" italics="true" pageId="3" pageNumber="86">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF9CF622AC7C27D5FB6454F5" box="[1122,1200,1946,1969]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
Genetic distance given in percentage and based on 529 aligned sites of 16S. Colored terminals match colors in Figure 5 (Map).
</paragraph>
</caption>
<caption id="DF012092FF9BF625A8892600FA465597" ID-DOI="http://doi.org/10.5281/zenodo.4403635" ID-Zenodo-Dep="4403635" httpUri="https://zenodo.org/record/4403635/files/figure.png" pageId="4" pageNumber="87" startId="4.[151,250,1615,1639]" targetBox="[267,1320,205,1584]" targetPageId="4">
<paragraph id="8BC1701AFF9BF625A8892600FA465597" blockId="4.[151,1437,1615,1748]" pageId="4" pageNumber="87">
<emphasis id="B90AAC08FF9BF625A8892600FEC1552C" bold="true" box="[151,277,1615,1640]" pageId="4" pageNumber="87">FIGURE 2.</emphasis>
Adult males of 
<taxonomicName id="4C7E0B99FF9BF625A9A1261FFDAD5523" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[447,633,1616,1639]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="87" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF9BF625A9A1261FFDAD5523" box="[447,633,1616,1639]" italics="true" pageId="4" pageNumber="87">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF9BF625AA9F261FFD065523" box="[641,722,1616,1639]" pageId="4" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
in life: (A) MHUA-A 8813, SVL 20.1 mm, holotype; (B) MHUA-A 8814, SVL 20.5 mm, paratype; (C) MHUA-A 8815, SVL 20.3 mm, paratype, adult male; (D) MHUA-A 8816, SVL 19.5 mm, paratype; (E) MHUA-A 8817, SVL 21.5 mm, paratype; (F) MHUA-A 11165, SVL 20.1 mm, paratype; (G) MHUA-A 11166, SVL 21.3 mm, paratype; (H) MHUA-A 11167, SVL 20.8 mm, paratype. Photos by Adriana Restrepo, Carlos Marín and JMD.
</paragraph>
</caption>
<subSubSection id="C3642391FF9BF624A8D9274DFBF652E9" lastPageId="5" lastPageNumber="88" pageId="4" pageNumber="87" type="diagnosis">
<paragraph id="8BC1701AFF9BF624A8D9274DFBF652E9" blockId="4.[151,1437,1794,2036]" lastBlockId="5.[151,1437,151,969]" lastPageId="5" lastPageNumber="88" pageId="4" pageNumber="87">
<emphasis id="B90AAC08FF9BF625A8D9274DFE955458" bold="true" box="[199,321,1794,1820]" pageId="4" pageNumber="87">
<emphasis id="B90AAC08FF9BF625A8D9274DFEEE5458" bold="true" box="[199,314,1794,1820]" italics="true" pageId="4" pageNumber="87">Diagnosis</emphasis>
.
</emphasis>
We assigned the new species to the genus 
<taxonomicName id="4C7E0B99FF9BF625AB5A274CFC195458" box="[836,973,1795,1820]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis id="B90AAC08FF9BF625AB5A274CFC195458" box="[836,973,1795,1820]" italics="true" pageId="4" pageNumber="87">Pristimantis</emphasis>
</taxonomicName>
based on our phylogenetic results. The new species (
<figureCitation id="13456C9FFF9BF625A9302769FEAF5404" box="[302,379,1830,1856]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1639]" captionTargetBox="[267,1320,205,1584]" captionTargetId="figure-33@4.[256,1331,181,1592]" captionTargetPageId="4" captionText="FIGURE 2. Adult males of Pristimantis zorro sp. nov. in life: (A) MHUA-A 8813, SVL 20.1 mm, holotype; (B) MHUA-A 8814, SVL 20.5 mm, paratype; (C) MHUA-A 8815, SVL 20.3 mm, paratype, adult male; (D) MHUA-A 8816, SVL 19.5 mm, paratype; (E) MHUA-A 8817, SVL 21.5 mm, paratype; (F) MHUA-A 11165, SVL 20.1 mm, paratype; (G) MHUA-A 11166, SVL 21.3 mm, paratype; (H) MHUA-A 11167, SVL 20.8 mm, paratype. Photos by Adriana Restrepo, Carlos Marín and JMD." figureDoi="http://doi.org/10.5281/zenodo.4403635" httpUri="https://zenodo.org/record/4403635/files/figure.png" pageId="4" pageNumber="87">Figs. 2</figureCitation>
A-H) is characterized by a combination of (1) skin texture of dorsum smooth, venter areolate; dorsolateral and discoidal fold absent; (2) tympanic membrane and tympanic annulus evident, supratympanic fold differentiated; horizontal diameter of tympanum 23–33% of eye diameter; (3) snout moderate in length, with small rostral papilla, subacuminate in dorsal view and truncate in profile; (4) tubercles on upper eyelids and cranial crests absent; (5) dentigerous process of the vomer present; (6) males with vocal slits and median subgular vocal sac; nuptial pads absent; (7) Finger I shorter than Finger II; discs broadly expanded and elliptical; (8) fingers with nar- row lateral fringes; (9) antebrachial and ulnar tubercles absent; (10) tubercles on heel very small and outer edge of tarsus absent; inner tarsal fold absent; (11) inner metatarsal tubercle oval, two-to-three times as long as round outer metatarsal tubercle; supernumerary plantar tubercles small and low, at the base of Toes III and IV; (12) toes with narrow lateral fringes; webbing absent; fifth toe longer than third; (13) in life, dorsum light green to green-yellow; well-defined canthal stripe; blotch covering the front of the face, including inter-nostrils area forming a facial mask; inter-orbital brown bar and supra-tympanic brown stripe present; blotches on the wrist of the hand, forming a bracelet (
<figureCitation id="13456C9FFF9AF624A8DC2120FED252CD" box="[194,262,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1639]" captionTargetBox="[267,1320,205,1584]" captionTargetId="figure-33@4.[256,1331,181,1592]" captionTargetPageId="4" captionText="FIGURE 2. Adult males of Pristimantis zorro sp. nov. in life: (A) MHUA-A 8813, SVL 20.1 mm, holotype; (B) MHUA-A 8814, SVL 20.5 mm, paratype; (C) MHUA-A 8815, SVL 20.3 mm, paratype, adult male; (D) MHUA-A 8816, SVL 19.5 mm, paratype; (E) MHUA-A 8817, SVL 21.5 mm, paratype; (F) MHUA-A 11165, SVL 20.1 mm, paratype; (G) MHUA-A 11166, SVL 21.3 mm, paratype; (H) MHUA-A 11167, SVL 20.8 mm, paratype. Photos by Adriana Restrepo, Carlos Marín and JMD." figureDoi="http://doi.org/10.5281/zenodo.4403635" httpUri="https://zenodo.org/record/4403635/files/figure.png" pageId="5" pageNumber="88">Fig. 2</figureCitation>
); iris is copper with fine brown reticulation; yellow throat; venter cream; posterior surfaces of thighs and shanks yellow; (14) adults small, SVL in males 19.5–21.5 (20.5 ± 0.495, 
<emphasis id="B90AAC08FF9AF624ABD521DCFC0D52E8" box="[971,985,403,428]" italics="true" pageId="5" pageNumber="88">n</emphasis>
= 11).
</paragraph>
</subSubSection>
<subSubSection id="C3642391FF9AF626A8D921F8FBE355E7" lastPageId="7" lastPageNumber="90" pageId="5" pageNumber="88" type="description">
<paragraph id="8BC1701AFF9AF624A8D921F8FBC451B5" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="88">
<emphasis id="B90AAC08FF9AF624A8D921F8FDDC5295" bold="true" box="[199,520,438,465]" pageId="5" pageNumber="88">
<emphasis id="B90AAC08FF9AF624A8D921F8FDD55295" bold="true" box="[199,513,438,465]" italics="true" pageId="5" pageNumber="88">
Description of the 
<typeStatus id="54C5CEB8FF9AF624A98121F9FDD55295" box="[415,513,438,465]" pageId="5" pageNumber="88" type="holotype">holotype</typeStatus>
</emphasis>
.
</emphasis>
Adult male, head as wide as long; snout relatively short, acuminate in dorsal and truncate in lateral view, but with small conical papilla at tip (snout–eye distance 16% of SVL); 
<emphasis id="B90AAC08FF9AF624ACC02194FA4852B1" box="[1246,1436,475,501]" italics="true" pageId="5" pageNumber="88">
canthus 
<taxonomicName id="4C7E0B99FF9AF624AD202194FA4852B1" baseAuthorityName="Werner" baseAuthorityYear="1896" box="[1342,1436,475,501]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="rostralis">rostralis</taxonomicName>
</emphasis>
indistinct; loreal region slightly concave; nostrils directed anterolaterally, protuberant, inter-nostrils area slightly concave in dorsal view; interorbital area flat, as broad as upper eyelid; cranial crests absent; upper eyelid without tubercles; tympanic membrane and tympanic annulus distinct, round; supratympanic fold differentiated; tympanum diameter 26% of eye diameter; postrictal tubercles low, inconspicuous. Choanae small, nearly rounded, not concealed by palatal shelf of maxillary; dentigerous process of the vomer present, elliptical, low, with three teeth each; tongue longer that wide, posterior one-half free from floor of mouth. Texture of skin of dorsum, flanks and venter smooth, dorsolateral, thoracic and discoidal folds absent; cloacal sheath absent.
</paragraph>
<paragraph id="8BC1701AFF9AF624A8D922B4FBC4508D" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="88">Forearm slender; radio–ulna length 20% of SVL; ulnar tubercles and ulnar fold absent; hand length longer than radio–ulna length (hand length 31% of SVL); fingers with narrow lateral fringes; relative lengths of fingers I &lt;II &lt;IV &lt;III; palmar tubercle bifid, thenar tubercle oval, inconspicuous; subarticular tubercles round, low; supernumerary palmar tubercles present at the base of all fingers, low, inconspicuous; disc cover of Finger I slightly expanded, those of Fingers II–IV extensively expanded; outer discs of fingers as wide as those of toes; all disc covers with elliptical ventral pads defined by circummarginal grooves. Nuptial pads absent.</paragraph>
<caption id="DF012092FF9AF624A889274FFF035479" ID-DOI="http://doi.org/10.5281/zenodo.4403639" ID-Zenodo-Dep="4403639" httpUri="https://zenodo.org/record/4403639/files/figure.png" pageId="5" pageNumber="88" startId="5.[151,250,1792,1816]" targetBox="[158,1424,1004,1760]" targetPageId="5">
<paragraph id="8BC1701AFF9AF624A889274FFF035479" blockId="5.[151,1436,1792,1853]" pageId="5" pageNumber="88">
<emphasis id="B90AAC08FF9AF624A889274FFEC1545C" bold="true" box="[151,277,1792,1816]" pageId="5" pageNumber="88">FIGURE 3.</emphasis>
Paratype of 
<taxonomicName id="4C7E0B99FF9AF624A982274EFD82545C" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[412,598,1793,1816]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="88" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF9AF624A982274EFD82545C" box="[412,598,1793,1816]" italics="true" pageId="5" pageNumber="88">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF9AF624AA40274EFD7B545C" box="[606,687,1793,1816]" pageId="5" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
in preservative. MHUA-A 11165, SVL 20.1 mm, adult male. Photos by MRC.
</paragraph>
</caption>
<paragraph id="8BC1701AFF9AF627A8D92724FBAD5391" blockId="5.[151,1437,1899,2033]" lastBlockId="6.[151,1437,151,717]" lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88">Hind limbs relatively slender; tibia length 56% of SVL; foot length 76% of tibia length; tarsal fold and tarsal tubercles absent; heel (tibia-tarsal articulation) without tubercles; toes with narrow lateral fringes; subarticular tubercles round, low; inner metatarsal tubercle oval, about 2x as long as wide; subconical outer tubercle; supernumerary plantar tubercles inconspicuous; disc covers slightly expanded; toes with defined pads; disc pads nearly ellipti- cal; relative lengths of toes I &lt;II &lt;III &lt;V &lt;IV; tip of Toe V reaching distal border of distal subarticular tubercle of Toe IV; tip of Toe III reaching proximal border of medial subarticular tubercle of Toe IV.</paragraph>
<paragraph id="8BC1701AFF99F627A8D92091FBC15125" blockId="6.[151,1437,151,717]" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8D92091FDD753BD" bold="true" box="[199,515,222,249]" italics="true" pageId="6" pageNumber="89">
Coloration of the 
<typeStatus id="54C5CEB8FF99F627A9842091FE2B53BD" box="[410,511,222,249]" pageId="6" pageNumber="89" type="holotype">holotype</typeStatus>
.
</emphasis>
In life, the 
<typeStatus id="54C5CEB8FF99F627AA902090FD2453BD" box="[654,752,223,249]" pageId="6" pageNumber="89" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C7E0B99FF99F627AB072090FC3353BC" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[793,999,223,248]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="89" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF99F627AB072090FC3353BC" box="[793,999,223,248]" italics="true" pageId="6" pageNumber="89">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF99F627ABEE2090FB9753BD" box="[1008,1091,223,249]" pageId="6" pageNumber="89" rank="species">sp. nov</taxonomicNameLabel>
is yellow dorsally, with some dark brown blotches and spots and yellow-green in the supra-ocular region. The 
<typeStatus id="54C5CEB8FF99F627AC0C214CFBA05259" box="[1042,1140,259,285]" pageId="6" pageNumber="89" type="holotype">holotype</typeStatus>
has a well-defined canthal brown stripe covering the front of the face, extending to the inter-nostrils region and forming a facial mask. The inter-orbital brown bar and supra-tympanic brown stripe is present, extending towards the flanks to the middle of the body. The wrist of the hands is intensely pigmented in the dorsal region; the flanks and limbs have scattered brown spots; throat, axillae, undersides and posterior surfaces of thighs immaculate with light yellow coloration; venter is white; palmar and plantar side light yellow; ventral side of limbs and thighs light yellow without spots or marks. The iris is cooper with fine brown reticulation (
<figureCitation id="13456C9FFF99F627AAE22194FC8252B1" box="[764,854,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1639]" captionTargetBox="[267,1320,205,1584]" captionTargetId="figure-33@4.[256,1331,181,1592]" captionTargetPageId="4" captionText="FIGURE 2. Adult males of Pristimantis zorro sp. nov. in life: (A) MHUA-A 8813, SVL 20.1 mm, holotype; (B) MHUA-A 8814, SVL 20.5 mm, paratype; (C) MHUA-A 8815, SVL 20.3 mm, paratype, adult male; (D) MHUA-A 8816, SVL 19.5 mm, paratype; (E) MHUA-A 8817, SVL 21.5 mm, paratype; (F) MHUA-A 11165, SVL 20.1 mm, paratype; (G) MHUA-A 11166, SVL 21.3 mm, paratype; (H) MHUA-A 11167, SVL 20.8 mm, paratype. Photos by Adriana Restrepo, Carlos Marín and JMD." figureDoi="http://doi.org/10.5281/zenodo.4403635" httpUri="https://zenodo.org/record/4403635/files/figure.png" pageId="6" pageNumber="89">Fig. 2A</figureCitation>
). In preservative, the yellow of the body turned into pale cream, both on the dorsum and on the flanks; the brown botches o and marks became darker almost black and dorsum show traces of very small black dots throughout the body, only perceptible to stereoscope. In addition, arm and legs cream with dark brown spots and blotches, cream belly without marks.
</paragraph>
<paragraph id="8BC1701AFF99F627A8D92224FED15189" blockId="6.[151,1437,151,717]" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8D92224FD3C51C1" bold="true" box="[199,744,618,645]" italics="true" pageId="6" pageNumber="89">
Measurements of the 
<typeStatus id="54C5CEB8FF99F627A9DA2225FDF251C1" box="[452,550,618,645]" pageId="6" pageNumber="89" type="holotype">holotype</typeStatus>
(in millimeters).
</emphasis>
SVL: 20.1; HL: 7.5 HW: 7.7; ED: 3.0; END: 2.4; NSD: 0.9; IND: 1.7; AMD: 4.4; TD: 0.8; FAL: 4.0; FAB: 1.2; HAL: 62; THL: 10.7; TL: 11.3; TAL: 5.6; FL: 8.6; TFD: 1.0; FTD: 1.9.
</paragraph>
<caption id="DF012092FF99F627A88922B3FC08507C" pageId="6" pageNumber="89">
<paragraph id="8BC1701AFF99F627A88922B3FC08507C" blockId="6.[151,1437,764,825]" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A88922B3FEDD5050" bold="true" box="[151,265,764,788]" pageId="6" pageNumber="89">TABLE 1.</emphasis>
Morphological variation (in mm) of the type series of 
<taxonomicName id="4C7E0B99FF99F627AB5822B2FBD35050" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[838,1031,765,788]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="89" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF99F627AB5822B2FBD35050" box="[838,1031,765,788]" italics="true" pageId="6" pageNumber="89">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF99F627AC1322B2FB8A5050" box="[1037,1118,765,788]" pageId="6" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
Min. = minimum value; Max. = maximum value; Mean = arithmetic mean value; S.D. = standard deviation).
</paragraph>
</caption>
<paragraph id="8BC1701AFF99F627A9642313FAEB5448" pageId="6" pageNumber="89">
<table id="F97E82BAFF9909DEA8BD2307FAEB5448" box="[163,1343,840,1804]" gridcols="5" gridrows="19" pageId="6" pageNumber="89">
<tr id="354E7258FF9909DEA8BD2307FAEB50CD" box="[163,1343,840,905]" gridrow="0" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD2307FD3250CD" box="[163,742,840,905]" gridcol="0" gridrow="0" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A9642313FD605031" bold="true" box="[378,692,860,885]" pageId="6" pageNumber="89">Morphometric Measurement</emphasis>
</th>
<th id="769F1B24FF9909DEABB52307FC0850CD" box="[939,988,840,905]" gridcol="1" gridrow="0" pageId="6" pageNumber="89">Min.</th>
<th id="769F1B24FF9909DEAC1A2307FB3250CD" box="[1028,1254,840,905]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627AC532307FB325024" bold="true" box="[1101,1254,840,864]" pageId="6" pageNumber="89">Males (n = 11)</emphasis>
Max. Mean
</th>
<th id="769F1B24FF9909DEAD182307FAEB50CD" box="[1286,1343,840,905]" gridcol="4" gridrow="0" pageId="6" pageNumber="89">S.D.</th>
</tr>
<tr id="354E7258FF9909DEA8BD23D2FAEB50F2" box="[163,1343,925,950]" gridrow="1" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD23D2FD3250F2" box="[163,742,925,950]" gridcol="0" gridrow="1" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD23D2FE7E50F2" bold="true" box="[163,426,925,950]" pageId="6" pageNumber="89">SVL = snout-vent length</emphasis>
</th>
<td id="769F1B24FF9909DEABB523D2FC0850F2" box="[939,988,925,950]" gridcol="1" gridrow="1" pageId="6" pageNumber="89">19.5</td>
<td id="769F1B24FF9909DEAC1A23D2FBE150F2" box="[1028,1077,925,950]" gridcol="2" gridrow="1" pageId="6" pageNumber="89">21.5</td>
<td id="769F1B24FF9909DEAC7B23D2FB3250F2" box="[1125,1254,925,950]" gridcol="3" gridrow="1" pageId="6" pageNumber="89">20.5</td>
<td id="769F1B24FF9909DEAD1823D2FAEB50F2" box="[1286,1343,925,950]" gridcol="4" gridrow="1" pageId="6" pageNumber="89">0.495</td>
</tr>
<tr id="354E7258FF9909DEA8BD239FFAEB50AC" box="[163,1343,976,1000]" gridrow="2" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD239FFD3250AC" box="[163,742,976,1000]" gridcol="0" gridrow="2" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD239FFE8B50AC" bold="true" box="[163,351,976,1000]" pageId="6" pageNumber="89">HL = head lenght</emphasis>
</th>
<td id="769F1B24FF9909DEABB5239FFC0850AC" box="[939,988,976,1000]" gridcol="1" gridrow="2" pageId="6" pageNumber="89">7.0</td>
<td id="769F1B24FF9909DEAC1A239FFBE150AC" box="[1028,1077,976,1000]" gridcol="2" gridrow="2" pageId="6" pageNumber="89">7.8</td>
<td id="769F1B24FF9909DEAC7B239FFB3250AC" box="[1125,1254,976,1000]" gridcol="3" gridrow="2" pageId="6" pageNumber="89">7.4</td>
<td id="769F1B24FF9909DEAD18239FFAEB50AC" box="[1286,1343,976,1000]" gridcol="4" gridrow="2" pageId="6" pageNumber="89">0.305</td>
</tr>
<tr id="354E7258FF9909DEA8BD244DFAEB575E" box="[163,1343,1026,1050]" gridrow="3" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD244DFD32575E" box="[163,742,1026,1050]" gridcol="0" gridrow="3" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD244DFEB7575E" bold="true" box="[163,355,1026,1050]" pageId="6" pageNumber="89">HW = head width</emphasis>
</th>
<td id="769F1B24FF9909DEABB5244DFC08575E" box="[939,988,1026,1050]" gridcol="1" gridrow="3" pageId="6" pageNumber="89">7.4</td>
<td id="769F1B24FF9909DEAC1A244DFBE1575E" box="[1028,1077,1026,1050]" gridcol="2" gridrow="3" pageId="6" pageNumber="89">8.4</td>
<td id="769F1B24FF9909DEAC7B244DFB32575E" box="[1125,1254,1026,1050]" gridcol="3" gridrow="3" pageId="6" pageNumber="89">8.0</td>
<td id="769F1B24FF9909DEAD18244DFAEB575E" box="[1286,1343,1026,1050]" gridcol="4" gridrow="3" pageId="6" pageNumber="89">0.298</td>
</tr>
<tr id="354E7258FF9909DEA8BD247BFAEB5709" box="[163,1343,1076,1101]" gridrow="4" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD247BFD325709" box="[163,742,1076,1101]" gridcol="0" gridrow="4" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD247BFEBF5708" bold="true" box="[163,363,1076,1100]" pageId="6" pageNumber="89">ED = eye diameter</emphasis>
</th>
<td id="769F1B24FF9909DEABB5247BFC085709" box="[939,988,1076,1101]" gridcol="1" gridrow="4" pageId="6" pageNumber="89">2.7</td>
<td id="769F1B24FF9909DEAC1A247BFBE15709" box="[1028,1077,1076,1101]" gridcol="2" gridrow="4" pageId="6" pageNumber="89">3.1</td>
<td id="769F1B24FF9909DEAC7B247BFB325709" box="[1125,1254,1076,1101]" gridcol="3" gridrow="4" pageId="6" pageNumber="89">2.9</td>
<td id="769F1B24FF9909DEAD18247BFAEB5709" box="[1286,1343,1076,1101]" gridcol="4" gridrow="4" pageId="6" pageNumber="89">0.133</td>
</tr>
<tr id="354E7258FF9909DEA8BD2429FAEB573B" box="[163,1343,1126,1151]" gridrow="5" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD2429FD32573B" box="[163,742,1126,1151]" gridcol="0" gridrow="5" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD2428FE0A573A" bold="true" box="[163,478,1126,1151]" pageId="6" pageNumber="89">END = eye to nostril distance</emphasis>
</th>
<td id="769F1B24FF9909DEABB52429FC08573B" box="[939,988,1126,1151]" gridcol="1" gridrow="5" pageId="6" pageNumber="89">2.2</td>
<td id="769F1B24FF9909DEAC1A2429FBE1573B" box="[1028,1077,1126,1151]" gridcol="2" gridrow="5" pageId="6" pageNumber="89">2.6</td>
<td id="769F1B24FF9909DEAC7B2429FB32573B" box="[1125,1254,1126,1151]" gridcol="3" gridrow="5" pageId="6" pageNumber="89">2.5</td>
<td id="769F1B24FF9909DEAD182429FAEB573B" box="[1286,1343,1126,1151]" gridcol="4" gridrow="5" pageId="6" pageNumber="89">0.118</td>
</tr>
<tr id="354E7258FF9909DEA8BD24D6FAEB57F5" box="[163,1343,1177,1201]" gridrow="6" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD24D6FD3257F5" box="[163,742,1177,1201]" gridcol="0" gridrow="6" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD24D6FDE657F5" bold="true" box="[163,562,1177,1201]" pageId="6" pageNumber="89">NSD = nostril to tip of snout distance</emphasis>
</th>
<td id="769F1B24FF9909DEABB524D6FC0857F5" box="[939,988,1177,1201]" gridcol="1" gridrow="6" pageId="6" pageNumber="89">0.8</td>
<td id="769F1B24FF9909DEAC1A24D6FBE157F5" box="[1028,1077,1177,1201]" gridcol="2" gridrow="6" pageId="6" pageNumber="89">1.0</td>
<td id="769F1B24FF9909DEAC7B24D6FB3257F5" box="[1125,1254,1177,1201]" gridcol="3" gridrow="6" pageId="6" pageNumber="89">0.9</td>
<td id="769F1B24FF9909DEAD1824D6FAEB57F5" box="[1286,1343,1177,1201]" gridcol="4" gridrow="6" pageId="6" pageNumber="89">0.060</td>
</tr>
<tr id="354E7258FF9909DEA8BD2484FAEB57A7" box="[163,1343,1227,1251]" gridrow="7" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD2484FD3257A7" box="[163,742,1227,1251]" gridcol="0" gridrow="7" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD2484FE1457A7" bold="true" box="[163,448,1227,1251]" pageId="6" pageNumber="89">IND = internarial distance</emphasis>
</th>
<td id="769F1B24FF9909DEABB52484FC0857A7" box="[939,988,1227,1251]" gridcol="1" gridrow="7" pageId="6" pageNumber="89">1.6</td>
<td id="769F1B24FF9909DEAC1A2484FBE157A7" box="[1028,1077,1227,1251]" gridcol="2" gridrow="7" pageId="6" pageNumber="89">1.8</td>
<td id="769F1B24FF9909DEAC7B2484FB3257A7" box="[1125,1254,1227,1251]" gridcol="3" gridrow="7" pageId="6" pageNumber="89">1.7</td>
<td id="769F1B24FF9909DEAD182484FAEB57A7" box="[1286,1343,1227,1251]" gridcol="4" gridrow="7" pageId="6" pageNumber="89">0.094</td>
</tr>
<tr id="354E7258FF9909DEA8BD24B2FAEB5652" box="[163,1343,1277,1302]" gridrow="8" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD24B2FD325652" box="[163,742,1277,1302]" gridcol="0" gridrow="8" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD24B2FD325652" bold="true" box="[163,742,1277,1302]" pageId="6" pageNumber="89">AMD = distance between the anterior margins of eyes</emphasis>
</th>
<td id="769F1B24FF9909DEABB524B2FC085652" box="[939,988,1277,1302]" gridcol="1" gridrow="8" pageId="6" pageNumber="89">4.1</td>
<td id="769F1B24FF9909DEAC1A24B2FBE15652" box="[1028,1077,1277,1302]" gridcol="2" gridrow="8" pageId="6" pageNumber="89">4.8</td>
<td id="769F1B24FF9909DEAC7B24B2FB325652" box="[1125,1254,1277,1302]" gridcol="3" gridrow="8" pageId="6" pageNumber="89">4.5</td>
<td id="769F1B24FF9909DEAD1824B2FAEB5652" box="[1286,1343,1277,1302]" gridcol="4" gridrow="8" pageId="6" pageNumber="89">0.218</td>
</tr>
<tr id="354E7258FF9909DEA8BD2560FAEB560C" box="[163,1343,1327,1352]" gridrow="9" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD2560FD32560C" box="[163,742,1327,1352]" gridcol="0" gridrow="9" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD257FFE6A5603" bold="true" box="[163,446,1327,1352]" pageId="6" pageNumber="89">TD = tympanum diameter</emphasis>
</th>
<td id="769F1B24FF9909DEABB52560FC08560C" box="[939,988,1327,1352]" gridcol="1" gridrow="9" pageId="6" pageNumber="89">0.7</td>
<td id="769F1B24FF9909DEAC1A2560FBE1560C" box="[1028,1077,1327,1352]" gridcol="2" gridrow="9" pageId="6" pageNumber="89">1.0</td>
<td id="769F1B24FF9909DEAC7B2560FB32560C" box="[1125,1254,1327,1352]" gridcol="3" gridrow="9" pageId="6" pageNumber="89">0.9</td>
<td id="769F1B24FF9909DEAD182560FAEB560C" box="[1286,1343,1327,1352]" gridcol="4" gridrow="9" pageId="6" pageNumber="89">0.092</td>
</tr>
<tr id="354E7258FF9909DEA8BD252DFAEB563E" box="[163,1343,1378,1402]" gridrow="10" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD252DFD32563E" box="[163,742,1378,1402]" gridcol="0" gridrow="10" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD252DFE44563E" bold="true" box="[163,400,1378,1402]" pageId="6" pageNumber="89">FAL = forearm length</emphasis>
</th>
<td id="769F1B24FF9909DEABB5252DFC08563E" box="[939,988,1378,1402]" gridcol="1" gridrow="10" pageId="6" pageNumber="89">3.8</td>
<td id="769F1B24FF9909DEAC1A252DFBE1563E" box="[1028,1077,1378,1402]" gridcol="2" gridrow="10" pageId="6" pageNumber="89">4.8</td>
<td id="769F1B24FF9909DEAC7B252DFB32563E" box="[1125,1254,1378,1402]" gridcol="3" gridrow="10" pageId="6" pageNumber="89">4.4</td>
<td id="769F1B24FF9909DEAD18252DFAEB563E" box="[1286,1343,1378,1402]" gridcol="4" gridrow="10" pageId="6" pageNumber="89">0.279</td>
</tr>
<tr id="354E7258FF9909DEA8BD25DBFAEB56E9" box="[163,1343,1428,1453]" gridrow="11" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD25DBFD3256E9" box="[163,742,1428,1453]" gridcol="0" gridrow="11" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD25DBFE7656E8" bold="true" box="[163,418,1428,1452]" pageId="6" pageNumber="89">FAB = forearm breadth</emphasis>
</th>
<td id="769F1B24FF9909DEABB525DBFC0856E9" box="[939,988,1428,1453]" gridcol="1" gridrow="11" pageId="6" pageNumber="89">1.2</td>
<td id="769F1B24FF9909DEAC1A25DBFBE156E9" box="[1028,1077,1428,1453]" gridcol="2" gridrow="11" pageId="6" pageNumber="89">1.6</td>
<td id="769F1B24FF9909DEAC7B25DBFB3256E9" box="[1125,1254,1428,1453]" gridcol="3" gridrow="11" pageId="6" pageNumber="89">1.4</td>
<td id="769F1B24FF9909DEAD1825DBFAEB56E9" box="[1286,1343,1428,1453]" gridcol="4" gridrow="11" pageId="6" pageNumber="89">0.117</td>
</tr>
<tr id="354E7258FF9909DEA8BD2589FAEB569B" box="[163,1343,1478,1503]" gridrow="12" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD2589FD32569B" box="[163,742,1478,1503]" gridcol="0" gridrow="12" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD2589FEA0569B" bold="true" box="[163,372,1478,1503]" pageId="6" pageNumber="89">HAL = hand length</emphasis>
</th>
<td id="769F1B24FF9909DEABB52589FC08569B" box="[939,988,1478,1503]" gridcol="1" gridrow="12" pageId="6" pageNumber="89">5.7</td>
<td id="769F1B24FF9909DEAC1A2589FBE1569B" box="[1028,1077,1478,1503]" gridcol="2" gridrow="12" pageId="6" pageNumber="89">6.7</td>
<td id="769F1B24FF9909DEAC7B2589FB32569B" box="[1125,1254,1478,1503]" gridcol="3" gridrow="12" pageId="6" pageNumber="89">6.3</td>
<td id="769F1B24FF9909DEAD182589FAEB569B" box="[1286,1343,1478,1503]" gridcol="4" gridrow="12" pageId="6" pageNumber="89">0.330</td>
</tr>
<tr id="354E7258FF9909DEA8BD25B6FAEB5555" box="[163,1343,1529,1553]" gridrow="13" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD25B6FD325555" box="[163,742,1529,1553]" gridcol="0" gridrow="13" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD25B6FEA05555" bold="true" box="[163,372,1529,1553]" pageId="6" pageNumber="89">THL = thigh length</emphasis>
</th>
<td id="769F1B24FF9909DEABB525B6FC085555" box="[939,988,1529,1553]" gridcol="1" gridrow="13" pageId="6" pageNumber="89">9.6</td>
<td id="769F1B24FF9909DEAC1A25B6FBE15555" box="[1028,1077,1529,1553]" gridcol="2" gridrow="13" pageId="6" pageNumber="89">11.4</td>
<td id="769F1B24FF9909DEAC7B25B6FB325555" box="[1125,1254,1529,1553]" gridcol="3" gridrow="13" pageId="6" pageNumber="89">10.6</td>
<td id="769F1B24FF9909DEAD1825B6FAEB5555" box="[1286,1343,1529,1553]" gridcol="4" gridrow="13" pageId="6" pageNumber="89">0.576</td>
</tr>
<tr id="354E7258FF9909DEA8BD2664FAEB5507" box="[163,1343,1579,1603]" gridrow="14" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD2664FD325507" box="[163,742,1579,1603]" gridcol="0" gridrow="14" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD2664FE8E5507" bold="true" box="[163,346,1579,1603]" pageId="6" pageNumber="89">TL = tibia length</emphasis>
</th>
<td id="769F1B24FF9909DEABB52664FC085507" box="[939,988,1579,1603]" gridcol="1" gridrow="14" pageId="6" pageNumber="89">10.9</td>
<td id="769F1B24FF9909DEAC1A2664FBE15507" box="[1028,1077,1579,1603]" gridcol="2" gridrow="14" pageId="6" pageNumber="89">12.2</td>
<td id="769F1B24FF9909DEAC7B2664FB325507" box="[1125,1254,1579,1603]" gridcol="3" gridrow="14" pageId="6" pageNumber="89">11.6</td>
<td id="769F1B24FF9909DEAD182664FAEB5507" box="[1286,1343,1579,1603]" gridcol="4" gridrow="14" pageId="6" pageNumber="89">0.463</td>
</tr>
<tr id="354E7258FF9909DEA8BD2612FAEB5532" box="[163,1343,1629,1654]" gridrow="15" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD2612FD325532" box="[163,742,1629,1654]" gridcol="0" gridrow="15" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD2612FEA35531" bold="true" box="[163,375,1629,1653]" pageId="6" pageNumber="89">TAL = tarsal length</emphasis>
</th>
<td id="769F1B24FF9909DEABB52612FC085532" box="[939,988,1629,1654]" gridcol="1" gridrow="15" pageId="6" pageNumber="89">5.3</td>
<td id="769F1B24FF9909DEAC1A2612FBE15532" box="[1028,1077,1629,1654]" gridcol="2" gridrow="15" pageId="6" pageNumber="89">6.4</td>
<td id="769F1B24FF9909DEAC7B2612FB325532" box="[1125,1254,1629,1654]" gridcol="3" gridrow="15" pageId="6" pageNumber="89">6.0</td>
<td id="769F1B24FF9909DEAD182612FAEB5532" box="[1286,1343,1629,1654]" gridcol="4" gridrow="15" pageId="6" pageNumber="89">0.304</td>
</tr>
<tr id="354E7258FF9909DEA8BD26C0FAEB55EC" box="[163,1343,1679,1704]" gridrow="16" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD26C0FD3255EC" box="[163,742,1679,1704]" gridcol="0" gridrow="16" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD26DFFE8555EC" bold="true" box="[163,337,1679,1704]" pageId="6" pageNumber="89">FL = foot lenght</emphasis>
</th>
<td id="769F1B24FF9909DEABB526C0FC0855EC" box="[939,988,1679,1704]" gridcol="1" gridrow="16" pageId="6" pageNumber="89">8.4</td>
<td id="769F1B24FF9909DEAC1A26C0FBE155EC" box="[1028,1077,1679,1704]" gridcol="2" gridrow="16" pageId="6" pageNumber="89">10.2</td>
<td id="769F1B24FF9909DEAC7B26C0FB3255EC" box="[1125,1254,1679,1704]" gridcol="3" gridrow="16" pageId="6" pageNumber="89">9.3</td>
<td id="769F1B24FF9909DEAD1826C0FAEB55EC" box="[1286,1343,1679,1704]" gridcol="4" gridrow="16" pageId="6" pageNumber="89">0.502</td>
</tr>
<tr id="354E7258FF9909DEA8BD268DFAEB559E" box="[163,1343,1730,1754]" gridrow="17" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD268DFD32559E" box="[163,742,1730,1754]" gridcol="0" gridrow="17" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD268DFDDD559E" bold="true" box="[163,521,1730,1754]" pageId="6" pageNumber="89">TFD = third finger disk diameter</emphasis>
</th>
<td id="769F1B24FF9909DEABB5268DFC08559E" box="[939,988,1730,1754]" gridcol="1" gridrow="17" pageId="6" pageNumber="89">0.9</td>
<td id="769F1B24FF9909DEAC1A268DFBE1559E" box="[1028,1077,1730,1754]" gridcol="2" gridrow="17" pageId="6" pageNumber="89">1.2</td>
<td id="769F1B24FF9909DEAC7B268DFB32559E" box="[1125,1254,1730,1754]" gridcol="3" gridrow="17" pageId="6" pageNumber="89">1.0</td>
<td id="769F1B24FF9909DEAD18268DFAEB559E" box="[1286,1343,1730,1754]" gridcol="4" gridrow="17" pageId="6" pageNumber="89">0.087</td>
</tr>
<tr id="354E7258FF9909DEA8BD26BBFAEB5448" box="[163,1343,1780,1804]" gridrow="18" pageId="6" pageNumber="89">
<th id="769F1B24FF9909DEA8BD26BBFD325448" box="[163,742,1780,1804]" gridcol="0" gridrow="18" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8BD26BBFE215448" bold="true" box="[163,501,1780,1804]" pageId="6" pageNumber="89">FFD = fourth toe disk diameter</emphasis>
</th>
<td id="769F1B24FF9909DEABB526BBFC085448" box="[939,988,1780,1804]" gridcol="1" gridrow="18" pageId="6" pageNumber="89">0.8</td>
<td id="769F1B24FF9909DEAC1A26BBFBE15448" box="[1028,1077,1780,1804]" gridcol="2" gridrow="18" pageId="6" pageNumber="89">1.1</td>
<td id="769F1B24FF9909DEAC7B26BBFB325448" box="[1125,1254,1780,1804]" gridcol="3" gridrow="18" pageId="6" pageNumber="89">0.9</td>
<td id="769F1B24FF9909DEAD1826BBFAEB5448" box="[1286,1343,1780,1804]" gridcol="4" gridrow="18" pageId="6" pageNumber="89">0.083</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC1701AFF99F626A8D9270EFBB15259" blockId="6.[151,1436,1856,2027]" lastBlockId="7.[151,1437,150,501]" lastPageId="7" lastPageNumber="90" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8D9270EFD6E541F" box="[199,698,1856,1883]" italics="true" pageId="6" pageNumber="89">
<emphasis id="B90AAC08FF99F627A8D9270EFE34541F" bold="true" box="[199,480,1856,1883]" italics="true" pageId="6" pageNumber="89">
Variation of 
<typeStatus id="54C5CEB8FF99F627A941270EFE5A541F" box="[351,398,1857,1883]" pageId="6" pageNumber="89">type</typeStatus>
series.
</emphasis>
<taxonomicName id="4C7E0B99FF99F627A9F5270DFD6E541F" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[491,698,1858,1883]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="89" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">Pristimantis zorro</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A2391173FF99F627AADA270EFCCB541F" box="[708,799,1857,1883]" pageId="6" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
has dorsum light yellow in (MHUA-A 8815, 8816) to green-yellow (MHUA-A 8814, 8817) and presenting conspicuous brown spots, blotches and marks with variably extension (
<figureCitation id="13456C9FFF99F627A90F27C6FE8254E7" box="[273,342,1929,1955]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1639]" captionTargetBox="[267,1320,205,1584]" captionTargetId="figure-33@4.[256,1331,181,1592]" captionTargetPageId="4" captionText="FIGURE 2. Adult males of Pristimantis zorro sp. nov. in life: (A) MHUA-A 8813, SVL 20.1 mm, holotype; (B) MHUA-A 8814, SVL 20.5 mm, paratype; (C) MHUA-A 8815, SVL 20.3 mm, paratype, adult male; (D) MHUA-A 8816, SVL 19.5 mm, paratype; (E) MHUA-A 8817, SVL 21.5 mm, paratype; (F) MHUA-A 11165, SVL 20.1 mm, paratype; (G) MHUA-A 11166, SVL 21.3 mm, paratype; (H) MHUA-A 11167, SVL 20.8 mm, paratype. Photos by Adriana Restrepo, Carlos Marín and JMD." figureDoi="http://doi.org/10.5281/zenodo.4403635" httpUri="https://zenodo.org/record/4403635/files/figure.png" pageId="6" pageNumber="89">Fig. 2</figureCitation>
). Individuals have a well-defined canthal brown stripe and blotch covering the front of the face; in addition to a strong pigmentation in the inter-nostrils region forming a mask-like pattern. The extension and shape of the inter-orbital brown bar and supra-tympanic brown stripe vary between specimens, and is weakly stained in the inter-orbital region in MHUA-A 11165 (
<figureCitation id="13456C9FFF98F626AA9E20D8FD1553F5" box="[640,705,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[158,1424,1004,1760]" captionTargetId="figure-405@5.[151,1436,993,1769]" captionTargetPageId="5" captionText="FIGURE 3. Paratype of Pristimantis zorro sp. nov. in preservative. MHUA-A 11165, SVL 20.1 mm, adult male. Photos by MRC." figureDoi="http://doi.org/10.5281/zenodo.4403639" httpUri="https://zenodo.org/record/4403639/files/figure.png" pageId="7" pageNumber="90">Fig. 3</figureCitation>
) and in the supratimpanic region in MHUA-A 8816, and partially interrupted in MHUA-A 11165, MHUA-A 11167. The brown coloration of the marks on the wrist is less intense in MHUA-A 8814 and MHUA-A 8816. Additionally, in MHUA-A 8813, 8814 there are two spots in both the scapular and the urostil region. Variations in morphometric measurements are shown in 
<tableCitation id="C6FC45A1FF98F626AC15214CFB8B5259" box="[1035,1119,259,285]" captionStart="TABLE 1" captionStartId="6.[151,239,764,788]" captionText="TABLE 1. Morphological variation (in mm) of the type series of Pristimantis zorro sp. nov. Min. = minimum value; Max. = maximum value; Mean = arithmetic mean value; S.D. = standard deviation)." pageId="7" pageNumber="90">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BC1701AFF98F626A8D92169FB1552B1" blockId="7.[151,1437,150,501]" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A8D92169FE885205" bold="true" box="[199,348,294,321]" italics="true" pageId="7" pageNumber="90">Vocalization.</emphasis>
The advertisement call of 
<taxonomicName id="4C7E0B99FF98F626AA982168FC855204" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[646,849,295,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF98F626AA982168FC855204" box="[646,849,295,320]" italics="true" pageId="7" pageNumber="90">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF98F626AB492168FC7A5205" box="[855,942,295,321]" pageId="7" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
consists of single-pulsed, short note lacking frequency modulation, with a duration of 0.109 ± 0.015 (
<geoCoordinate id="EE4A16DDFF98F626AB092105FC8C5221" box="[791,856,330,357]" degrees="0.062" direction="north" orientation="latitude" pageId="7" pageNumber="90" precision="55" value="0.062">0.062</geoCoordinate>
<geoCoordinate id="EE4A16DDFF98F626AB462105FC735221" box="[856,935,330,357]" degrees="0.155" direction="west" orientation="longitude" pageId="7" pageNumber="90" precision="55" value="-0.155">–0.155</geoCoordinate>
) seconds (
<figureCitation id="13456C9FFF98F626AC002104FBAD5221" box="[1054,1145,331,357]" captionStart="FIGURE 4" captionStartId="8.[151,250,1501,1525]" captionTargetBox="[156,1429,744,1474]" captionTargetId="figure-316@8.[151,1437,741,1478]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement call of Pristimantis zorro sp. nov. (MHUA-A 8813, holotype). (A). Spectrogram; (B) Oscillogram; (C) Power spectrum. Photo by JMD." figureDoi="http://doi.org/10.5281/zenodo.4403641" httpUri="https://zenodo.org/record/4403641/files/figure.png" pageId="7" pageNumber="90">Fig. 4A</figureCitation>
). We observed up to three harmonics per note. The interval between notes is 1.258 ± 0.547 (
<geoCoordinate id="EE4A16DDFF98F626AB672120FC6E52CE" box="[889,954,367,394]" degrees="0.179" direction="north" orientation="latitude" pageId="7" pageNumber="90" precision="55" value="0.179">0.179</geoCoordinate>
<geoCoordinate id="EE4A16DDFF98F626ABA42120FBDD52CE" box="[954,1033,367,394]" degrees="3.763" direction="west" orientation="longitude" pageId="7" pageNumber="90" precision="55" value="-3.763">–3.763</geoCoordinate>
) seconds (Table 3). Call rate ranges from 34 to 68 calls per minute. The dominant frequency is 2.954 ± 94.434 (
<geoCoordinate id="EE4A16DDFF98F626ABC121DCFBCB52E9" box="[991,1055,403,429]" degrees="2.842" direction="north" orientation="latitude" pageId="7" pageNumber="90" precision="55" value="2.842">2.842</geoCoordinate>
<geoCoordinate id="EE4A16DDFF98F626AC0121DCFBB852E9" box="[1055,1132,403,429]" degrees="3.186" direction="west" orientation="longitude" pageId="7" pageNumber="90" precision="55" value="-3.186">–3.186</geoCoordinate>
kHz), minimum frequency 2.855 ± 0.091 (
<geoCoordinate id="EE4A16DDFF98F626A95E21F8FE555296" box="[320,385,439,466]" degrees="2.670" direction="north" orientation="latitude" pageId="7" pageNumber="90" precision="55" value="2.67">2.670</geoCoordinate>
<geoCoordinate id="EE4A16DDFF98F626A99F21F8FE1B5296" box="[385,463,439,466]" degrees="3.186" direction="west" orientation="longitude" pageId="7" pageNumber="90" precision="55" value="-3.186">–3.186</geoCoordinate>
) kHz, and maximum frequency 3.092 ± 56.584 (
<geoCoordinate id="EE4A16DDFF98F626ABF121F8FBE55295" box="[1007,1073,439,465]" degrees="2.928" direction="north" orientation="latitude" pageId="7" pageNumber="90" precision="55" value="2.928">2.928</geoCoordinate>
<geoCoordinate id="EE4A16DDFF98F626AC2F21F8FB555295" box="[1073,1153,439,465]" degrees="3.273" direction="west" orientation="longitude" pageId="7" pageNumber="90" precision="55" value="-3.273">–3.273</geoCoordinate>
). In qualitative terms, the amplitude of note suddenly increases and it quickly falls into a funnel at the end of it (
<figureCitation id="13456C9FFF98F626AC452194FB6052B1" box="[1115,1204,475,501]" captionStart="FIGURE 4" captionStartId="8.[151,250,1501,1525]" captionTargetBox="[156,1429,744,1474]" captionTargetId="figure-316@8.[151,1437,741,1478]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement call of Pristimantis zorro sp. nov. (MHUA-A 8813, holotype). (A). Spectrogram; (B) Oscillogram; (C) Power spectrum. Photo by JMD." figureDoi="http://doi.org/10.5281/zenodo.4403641" httpUri="https://zenodo.org/record/4403641/files/figure.png" pageId="7" pageNumber="90">Fig. 4B</figureCitation>
).
</paragraph>
<caption id="DF012092FF98F626A889226BFEA051C0" pageId="7" pageNumber="90">
<paragraph id="8BC1701AFF98F626A889226BFEA051C0" blockId="7.[151,1436,548,645]" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A889226BFEDF5178" bold="true" box="[151,267,548,572]" pageId="7" pageNumber="90">TABLE 2.</emphasis>
Advertisement call features of 
<taxonomicName id="4C7E0B99FF98F626AA7F226AFCF05178" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[609,804,549,572]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF98F626AA7F226AFCF05178" box="[609,804,549,572]" italics="true" pageId="7" pageNumber="90">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF98F626AB32226AFC545178" box="[812,896,549,572]" pageId="7" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
recorded at type locality, Guatapé Antioquia, Colombia. Values are reported as mean ± standard deviation (minimum-maximum). Body size (snout-vent length, SVL), air temperature (T. Air).
</paragraph>
</caption>
<paragraph id="8BC1701AFF98F626A8DD22DBFAA256C5" pageId="7" pageNumber="90">
<table id="F97E82BAFF9809DEA8BD22DBFA5B56C5" box="[163,1423,660,1409]" gridcols="7" gridrows="11" pageId="7" pageNumber="90">
<tr id="354E7258FF9809DEA8BD22DBFA5B518E" box="[163,1423,660,714]" gridrow="0" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD22DBFEEB518E" box="[163,319,660,714]" gridcol="0" gridrow="0" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A8DD22DBFEEB518E" bold="true" pageId="7" pageNumber="90">Acoustic measurements</emphasis>
</th>
<th id="769F1B24FF9809DEA94722DBFE3B518E" box="[345,495,660,714]" gridcol="1" gridrow="0" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A94722DBFE3B51E8" bold="true" box="[345,495,660,684]" pageId="7" pageNumber="90">MHUAA8813</emphasis>
(N=58)
</th>
<th id="769F1B24FF9809DEAA1322DBFD70518E" box="[525,676,660,714]" gridcol="2" gridrow="0" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626AA1322DBFD7751E8" bold="true" box="[525,675,660,684]" pageId="7" pageNumber="90">MHUAA8815</emphasis>
(N=78)
</th>
<th id="769F1B24FF9809DEAADF22DBFCB6518E" box="[705,866,660,714]" gridcol="3" gridrow="0" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626AADF22DBFCB651E8" bold="true" box="[705,866,660,684]" pageId="7" pageNumber="90">MHUAA12092</emphasis>
(N=38)
</th>
<th id="769F1B24FF9809DEAB6022DBFBF4518E" box="[894,1056,660,714]" gridcol="4" gridrow="0" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626AB6022DBFBF451E8" bold="true" box="[894,1056,660,684]" pageId="7" pageNumber="90">MHUAA12093</emphasis>
(N=70)
</th>
<th id="769F1B24FF9809DEAC2522DBFB08518E" box="[1083,1244,660,714]" gridcol="5" gridrow="0" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626AC2522DBFB0851E8" bold="true" box="[1083,1244,660,684]" pageId="7" pageNumber="90">MHUAA12094</emphasis>
(N=68)
</th>
<th id="769F1B24FF9809DEACE622DBFA5B518E" box="[1272,1423,660,714]" gridcol="6" gridrow="0" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626AD1022DBFAAD51E8" bold="true" box="[1294,1401,660,684]" pageId="7" pageNumber="90">Summary</emphasis>
(N=312)
</th>
</tr>
<tr id="354E7258FF9809DEA8BD2295FA5B504B" box="[163,1423,730,783]" gridrow="1" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD2295FEEB504B" box="[163,319,730,783]" gridcol="0" gridrow="1" pageId="7" pageNumber="90">Call duration (s)</th>
<td id="769F1B24FF9809DEA9472295FE3B504B" box="[345,495,730,783]" gridcol="1" gridrow="1" pageId="7" pageNumber="90">0.101±0.749 (0.087–0.126)</td>
<td id="769F1B24FF9809DEAA132295FD70504B" box="[525,676,730,783]" gridcol="2" gridrow="1" pageId="7" pageNumber="90">0.108±0.017 (0.062–0.155)</td>
<td id="769F1B24FF9809DEAADF2295FCB6504B" box="[705,866,730,783]" gridcol="3" gridrow="1" pageId="7" pageNumber="90">0.093±0.003 (0.087–0.099)</td>
<td id="769F1B24FF9809DEAB602295FBF4504B" box="[894,1056,730,783]" gridcol="4" gridrow="1" pageId="7" pageNumber="90">0.120±0.009 (0.094–0.134)</td>
<td id="769F1B24FF9809DEAC252295FB08504B" box="[1083,1244,730,783]" gridcol="5" gridrow="1" pageId="7" pageNumber="90">0.084±0.009 (0.068–0.114)</td>
<td id="769F1B24FF9809DEACE62295FA5B504B" box="[1272,1423,730,783]" gridcol="6" gridrow="1" pageId="7" pageNumber="90">0.109±0.015 (0.062–0.155)</td>
</tr>
<tr id="354E7258FF9809DEA8BD2350FA5B5036" box="[163,1423,799,882]" gridrow="2" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD2350FEEB5036" box="[163,319,799,882]" gridcol="0" gridrow="2" pageId="7" pageNumber="90">Intercall interval duration (s)</th>
<td id="769F1B24FF9809DEA9472350FE3B5036" box="[345,495,799,882]" gridcol="1" gridrow="2" pageId="7" pageNumber="90">1.490±0.514 (0.975–3.270)</td>
<td id="769F1B24FF9809DEAA132350FD705036" box="[525,676,799,882]" gridcol="2" gridrow="2" pageId="7" pageNumber="90">0.827±0.464 (0.179–3.115)</td>
<td id="769F1B24FF9809DEAADF2350FCB65036" box="[705,866,799,882]" gridcol="3" gridrow="2" pageId="7" pageNumber="90">1.450±0.514 (0.975–3.270)</td>
<td id="769F1B24FF9809DEAB602350FBF45036" box="[894,1056,799,882]" gridcol="4" gridrow="2" pageId="7" pageNumber="90">1.185±0.332 (0.713–2.873)</td>
<td id="769F1B24FF9809DEAC252350FB085036" box="[1083,1244,799,882]" gridcol="5" gridrow="2" pageId="7" pageNumber="90">1.142±0.747 (0.805–4.257)</td>
<td id="769F1B24FF9809DEACE62350FA5B5036" box="[1272,1423,799,882]" gridcol="6" gridrow="2" pageId="7" pageNumber="90">1.258±0.547 (0.179–3.270)</td>
</tr>
<tr id="354E7258FF9809DEA8BD23CDFA5B5091" box="[163,1423,898,981]" gridrow="3" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD23CDFEEB5091" box="[163,319,898,981]" gridcol="0" gridrow="3" pageId="7" pageNumber="90">Calls rate (No. Calls/ min)</th>
<td id="769F1B24FF9809DEA94723CDFE3B5091" box="[345,495,898,981]" gridcol="1" gridrow="3" pageId="7" pageNumber="90">38</td>
<td id="769F1B24FF9809DEAA1323CDFD705091" box="[525,676,898,981]" gridcol="2" gridrow="3" pageId="7" pageNumber="90">68</td>
<td id="769F1B24FF9809DEAADF23CDFCB65091" box="[705,866,898,981]" gridcol="3" gridrow="3" pageId="7" pageNumber="90">38</td>
<td id="769F1B24FF9809DEAB6023CDFBF45091" box="[894,1056,898,981]" gridcol="4" gridrow="3" pageId="7" pageNumber="90">40</td>
<td id="769F1B24FF9809DEAC2523CDFB085091" box="[1083,1244,898,981]" gridcol="5" gridrow="3" pageId="7" pageNumber="90">34</td>
<td id="769F1B24FF9809DEACE623CDFA5B5091" box="[1272,1423,898,981]" gridcol="6" gridrow="3" pageId="7" pageNumber="90">46</td>
</tr>
<tr id="354E7258FF9809DEA8BD23AAFA5B575F" box="[163,1423,997,1051]" gridrow="4" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD23AAFEEB575F" box="[163,319,997,1051]" gridcol="0" gridrow="4" pageId="7" pageNumber="90">Low frequency (kHz)</th>
<td id="769F1B24FF9809DEA94723AAFE3B575F" box="[345,495,997,1051]" gridcol="1" gridrow="4" pageId="7" pageNumber="90">2,928</td>
<td id="769F1B24FF9809DEAA1323AAFD70575F" box="[525,676,997,1051]" gridcol="2" gridrow="4" pageId="7" pageNumber="90">2,922±0.077 (2,670–3,186)</td>
<td id="769F1B24FF9809DEAADF23AAFCB6575F" box="[705,866,997,1051]" gridcol="3" gridrow="4" pageId="7" pageNumber="90">2,928</td>
<td id="769F1B24FF9809DEAB6023AAFBF4575F" box="[894,1056,997,1051]" gridcol="4" gridrow="4" pageId="7" pageNumber="90">2,756</td>
<td id="769F1B24FF9809DEAC2523AAFB08575F" box="[1083,1244,997,1051]" gridcol="5" gridrow="4" pageId="7" pageNumber="90">3,010±0.3353 (2,958–3,100)</td>
<td id="769F1B24FF9809DEACE623AAFA5B575F" box="[1272,1423,997,1051]" gridcol="6" gridrow="4" pageId="7" pageNumber="90">2,855±0.091 (2,670–3,186)</td>
</tr>
<tr id="354E7258FF9809DEA8BD2464FA5B5724" box="[163,1423,1067,1120]" gridrow="5" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD2464FEEB5724" box="[163,319,1067,1120]" gridcol="0" gridrow="5" pageId="7" pageNumber="90">Dominant frequency (Hz)</th>
<td id="769F1B24FF9809DEA9472464FE3B5724" box="[345,495,1067,1120]" gridcol="1" gridrow="5" pageId="7" pageNumber="90">3,010±0.1939 (2,928–3,100)</td>
<td id="769F1B24FF9809DEAA132464FD705724" box="[525,676,1067,1120]" gridcol="2" gridrow="5" pageId="7" pageNumber="90">3,008±0.7874 (2,842–3.186)</td>
<td id="769F1B24FF9809DEAADF2464FCB65724" box="[705,866,1067,1120]" gridcol="3" gridrow="5" pageId="7" pageNumber="90">3,014</td>
<td id="769F1B24FF9809DEAB602464FBF45724" box="[894,1056,1067,1120]" gridcol="4" gridrow="5" pageId="7" pageNumber="90">3,014</td>
<td id="769F1B24FF9809DEAC252464FB085724" box="[1083,1244,1067,1120]" gridcol="5" gridrow="5" pageId="7" pageNumber="90">3,093±0.1387 (2,842–3,186)</td>
<td id="769F1B24FF9809DEACE62464FA5B5724" box="[1272,1423,1067,1120]" gridcol="6" gridrow="5" pageId="7" pageNumber="90">2.954±94.434 (2,842–3,186)</td>
</tr>
<tr id="354E7258FF9809DEA8BD243FFA5B57E2" box="[163,1423,1136,1190]" gridrow="6" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD243FFEEB57E2" box="[163,319,1136,1190]" gridcol="0" gridrow="6" pageId="7" pageNumber="90">High frequency (Hz)</th>
<td id="769F1B24FF9809DEA947243FFE3B57E2" box="[345,495,1136,1190]" gridcol="1" gridrow="6" pageId="7" pageNumber="90">3,100</td>
<td id="769F1B24FF9809DEAA13243FFD7057E2" box="[525,676,1136,1190]" gridcol="2" gridrow="6" pageId="7" pageNumber="90">3,092±0.5658 (2,928–3,273)</td>
<td id="769F1B24FF9809DEAADF243FFCB657E2" box="[705,866,1136,1190]" gridcol="3" gridrow="6" pageId="7" pageNumber="90">3,100</td>
<td id="769F1B24FF9809DEAB60243FFBF457E2" box="[894,1056,1136,1190]" gridcol="4" gridrow="6" pageId="7" pageNumber="90">2,842</td>
<td id="769F1B24FF9809DEAC25243FFB0857E2" box="[1083,1244,1136,1190]" gridcol="5" gridrow="6" pageId="7" pageNumber="90">3,184±0.2949 (3,100–3,273)</td>
<td id="769F1B24FF9809DEACE6243FFA5B57E2" box="[1272,1423,1136,1190]" gridcol="6" gridrow="6" pageId="7" pageNumber="90">3.092±56.584 (2,928–3.273)</td>
</tr>
<tr id="354E7258FF9809DEA8BD24F9FA5B578A" box="[163,1423,1206,1230]" gridrow="7" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD24F9FEEB578A" box="[163,319,1206,1230]" gridcol="0" gridrow="7" pageId="7" pageNumber="90">No. Harmonics</th>
<td id="769F1B24FF9809DEA94724F9FE3B578A" box="[345,495,1206,1230]" gridcol="1" gridrow="7" pageId="7" pageNumber="90">1</td>
<td id="769F1B24FF9809DEAA1324F9FD70578A" box="[525,676,1206,1230]" gridcol="2" gridrow="7" pageId="7" pageNumber="90">3</td>
<td id="769F1B24FF9809DEAADF24F9FCB6578A" box="[705,866,1206,1230]" gridcol="3" gridrow="7" pageId="7" pageNumber="90">1</td>
<td id="769F1B24FF9809DEAB6024F9FBF4578A" box="[894,1056,1206,1230]" gridcol="4" gridrow="7" pageId="7" pageNumber="90">1</td>
<td id="769F1B24FF9809DEAC2524F9FB08578A" box="[1083,1244,1206,1230]" gridcol="5" gridrow="7" pageId="7" pageNumber="90">0</td>
<td id="769F1B24FF9809DEACE624F9FA5B578A" box="[1272,1423,1206,1230]" gridcol="6" gridrow="7" pageId="7" pageNumber="90">0–3</td>
</tr>
<tr id="354E7258FF9809DEA8BD2491FA5B5675" box="[163,1423,1246,1329]" gridrow="8" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD2491FEEB5675" box="[163,319,1246,1329]" gridcol="0" gridrow="8" pageId="7" pageNumber="90">Harmonic frequency (Hz)</th>
<td id="769F1B24FF9809DEA9472491FE3B5675" box="[345,495,1246,1329]" gridcol="1" gridrow="8" pageId="7" pageNumber="90">5,598–6,029</td>
<td id="769F1B24FF9809DEAA132491FD705675" box="[525,676,1246,1329]" gridcol="2" gridrow="8" pageId="7" pageNumber="90">5,684–5,943 8,441–8,957 14,211–14,901</td>
<td id="769F1B24FF9809DEAADF2491FCB65675" box="[705,866,1246,1329]" gridcol="3" gridrow="8" pageId="7" pageNumber="90">5,943</td>
<td id="769F1B24FF9809DEAB602491FBF45675" box="[894,1056,1246,1329]" gridcol="4" gridrow="8" pageId="7" pageNumber="90">6,048</td>
<td id="769F1B24FF9809DEAC252491FB085675" box="[1083,1244,1246,1329]" gridcol="5" gridrow="8" pageId="7" pageNumber="90">–</td>
<td id="769F1B24FF9809DEACE62491FA5B5675" box="[1272,1423,1246,1329]" gridcol="6" gridrow="8" pageId="7" pageNumber="90">5,598–6,048 8,441–8,957 14,211–14,901</td>
</tr>
<tr id="354E7258FF9809DEA8BD250EFA5B561D" box="[163,1423,1345,1369]" gridrow="9" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD250EFEEB561D" box="[163,319,1345,1369]" gridcol="0" gridrow="9" pageId="7" pageNumber="90">SVL (mm)</th>
<td id="769F1B24FF9809DEA947250EFE3B561D" box="[345,495,1345,1369]" gridcol="1" gridrow="9" pageId="7" pageNumber="90">20.1</td>
<td id="769F1B24FF9809DEAA13250EFD70561D" box="[525,676,1345,1369]" gridcol="2" gridrow="9" pageId="7" pageNumber="90">20.5</td>
<td id="769F1B24FF9809DEAADF250EFCB6561D" box="[705,866,1345,1369]" gridcol="3" gridrow="9" pageId="7" pageNumber="90">20.5</td>
<td id="769F1B24FF9809DEAB60250EFBF4561D" box="[894,1056,1345,1369]" gridcol="4" gridrow="9" pageId="7" pageNumber="90">20.7</td>
<td id="769F1B24FF9809DEAC25250EFB08561D" box="[1083,1244,1345,1369]" gridcol="5" gridrow="9" pageId="7" pageNumber="90">20.6</td>
<td id="769F1B24FF9809DEACE6250EFA5B561D" box="[1272,1423,1345,1369]" gridcol="6" gridrow="9" pageId="7" pageNumber="90">20.1–20.7</td>
</tr>
<tr id="354E7258FF9809DEA8BD2526FA5B56C5" box="[163,1423,1385,1409]" gridrow="10" pageId="7" pageNumber="90">
<th id="769F1B24FF9809DEA8BD2526FEEB56C5" box="[163,319,1385,1409]" gridcol="0" gridrow="10" pageId="7" pageNumber="90">T. Air (°C)</th>
<td id="769F1B24FF9809DEA9472526FE3B56C5" box="[345,495,1385,1409]" gridcol="1" gridrow="10" pageId="7" pageNumber="90">16.0</td>
<td id="769F1B24FF9809DEAA132526FD7056C5" box="[525,676,1385,1409]" gridcol="2" gridrow="10" pageId="7" pageNumber="90">17.0</td>
<td id="769F1B24FF9809DEAADF2526FCB656C5" box="[705,866,1385,1409]" gridcol="3" gridrow="10" pageId="7" pageNumber="90">16.7</td>
<td id="769F1B24FF9809DEAB602526FBF456C5" box="[894,1056,1385,1409]" gridcol="4" gridrow="10" pageId="7" pageNumber="90">20.0</td>
<td id="769F1B24FF9809DEAC252526FB0856C5" box="[1083,1244,1385,1409]" gridcol="5" gridrow="10" pageId="7" pageNumber="90">21.8</td>
<td id="769F1B24FF9809DEACE62526FA5B56C5" box="[1272,1423,1385,1409]" gridcol="6" gridrow="10" pageId="7" pageNumber="90">16.0–21.8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC1701AFF98F626A8D925FFFBE355E7" blockId="7.[151,1437,1456,2024]" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A8D925FFFC46568F" box="[199,914,1456,1483]" italics="true" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A8D925FFFD16568F" bold="true" box="[199,706,1456,1483]" italics="true" pageId="7" pageNumber="90">Geographic distribution and natural history.</emphasis>
<taxonomicName id="4C7E0B99FF98F626AAD625FDFC46568F" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[712,914,1458,1483]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">Pristimantis zorro</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A2391173FF98F626AB8625FEFC39568F" box="[920,1005,1457,1483]" pageId="7" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
is known only from the 
<typeStatus id="54C5CEB8FF98F626ACE625FEFAFC568F" box="[1272,1320,1457,1483]" pageId="7" pageNumber="90">type</typeStatus>
locality at elevations ca. 
<quantity id="4C86DDFFFF98F626A928259AFE5F56AB" box="[310,395,1493,1519]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="7" pageNumber="90" unit="m" value="1860.0">1860 m</quantity>
a.s.l., near the Peñol-Guatape hydroelectric reservoir, on the northeastern Cordillera Central, in 
<collectingRegion id="49BABEF8FF98F626A8AC25B6FEF15557" box="[178,293,1529,1555]" country="Colombia" name="Antioquia" pageId="7" pageNumber="90">Antioquia</collectingRegion>
, 
<collectingCountry id="F369308AFF98F626A92E25B6FE4A5557" box="[304,414,1529,1555]" name="Colombia" pageId="7" pageNumber="90">Colombia</collectingCountry>
(
<figureCitation id="13456C9FFF98F626A9B325B6FE255550" box="[429,497,1529,1556]" captionStart="FIGURE 5" captionStartId="9.[151,250,1815,1839]" captionTargetBox="[160,1429,317,1785]" captionTargetId="figure-67@9.[151,1435,309,1792]" captionTargetPageId="9" captionText="FIGURE 5. Map showing the type locality of Pristimantis zorro sp. nov. and relatives. Pristimantis zorro sp. nov. (green dot) Antioquia, Colombia; P. pulchridormientes (blue dot) Huánuco, Perú; P. pluvialis (red dot) Cusco, Perú; Pristimantis olivaceus (CORBIDI 17473, orange dot) Cusco, Perú and Pristimantis sp (ROM 43978, purple dot) Potaro-Siparuni, Guyana." figureDoi="http://doi.org/10.5281/zenodo.4403643" httpUri="https://zenodo.org/record/4403643/files/figure.png" pageId="7" pageNumber="90">Fig. 5</figureCitation>
). We found individuals on shrubs at 1.0 to 4.0 m from the ground, in open areas and on the side of the road next to mature forest. We observed and collected active frogs between 1830–2100 h. We heard calls at night, during both clear and full moon nights, and overcast and rainy nights. Reproductive activity of females, and other natural history characteristics remain unknown. Sympatric congeneric species include 
<emphasis id="B90AAC08FF98F626AD292629FD9C55E7" italics="true" pageId="7" pageNumber="90">
<taxonomicName id="4C7E0B99FF98F626AD292629FF6955E7" baseAuthorityName="Lynch &amp; Rueda-Almonacid" baseAuthorityYear="1998" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="factiosus">P. factiosus</taxonomicName>
, 
<taxonomicName id="4C7E0B99FF98F626A8D726C5FE9055E7" baseAuthorityName="Lynch &amp; Rueda-Almonacid" baseAuthorityYear="1998" box="[201,324,1673,1699]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="helvolus">P. helvolus</taxonomicName>
, 
<taxonomicName id="4C7E0B99FF98F626A94E26C5FE7855E7" baseAuthorityName="Lynch &amp; Rueda-Almonacid" baseAuthorityYear="1998" box="[336,428,1673,1699]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="lemur">P. lemur</taxonomicName>
, 
<taxonomicName id="4C7E0B99FF98F626A9A926C5FD9055E7" baseAuthorityName="Lynch, Ruiz-Carranza &amp; Ardila-Robayo" baseAuthorityYear="1994" box="[439,580,1674,1699]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="permixtus">P. permixtus</taxonomicName>
,
</emphasis>
<taxonomicName id="4C7E0B99FF98F626AA5126C5FD0055E7" baseAuthorityName="Lynch &amp; Ardila-Robayo" baseAuthorityYear="1999" box="[591,724,1673,1699]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="paisa">
<emphasis id="B90AAC08FF98F626AA5126C5FDB155E7" box="[591,613,1674,1699]" italics="true" pageId="7" pageNumber="90">P.</emphasis>
aff. 
<emphasis id="B90AAC08FF98F626AA8926C5FD0055E7" box="[663,724,1674,1699]" italics="true" pageId="7" pageNumber="90">paisa</emphasis>
</taxonomicName>
<taxonomicName id="4C7E0B99FF98F626AAC526C5FCB855E7" baseAuthorityName="Lynch &amp; Rueda-Almonacid" baseAuthorityYear="1998" box="[731,876,1673,1699]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="suetus">
<emphasis id="B90AAC08FF98F626AAC526C5FD2455E7" box="[731,752,1674,1699]" italics="true" pageId="7" pageNumber="90">P.</emphasis>
aff. 
<emphasis id="B90AAC08FF98F626AB3D26C5FCB855E7" box="[803,876,1674,1699]" italics="true" pageId="7" pageNumber="90">suetus</emphasis>
</taxonomicName>
<emphasis id="B90AAC08FF98F626AB7226C5FCA455E7" box="[876,880,1674,1699]" italics="true" pageId="7" pageNumber="90">,</emphasis>
and 
<taxonomicName id="4C7E0B99FF98F626ABB926C5FBE455E7" baseAuthorityName="Lynch &amp; Rueda-Almonacid" baseAuthorityYear="1999" box="[935,1072,1673,1699]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="viejas">
<emphasis id="B90AAC08FF98F626ABB926C5FC6855E7" box="[935,956,1674,1699]" italics="true" pageId="7" pageNumber="90">P.</emphasis>
aff. 
<emphasis id="B90AAC08FF98F626ABF126C5FBE455E7" box="[1007,1072,1674,1699]" italics="true" pageId="7" pageNumber="90">viejas</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3642391FF98F629A8D926E3FB1B5189" lastPageId="8" lastPageNumber="91" pageId="7" pageNumber="90" type="etymology">
<paragraph id="8BC1701AFF98F626A8D926E3FCD35477" blockId="7.[151,1437,1456,2024]" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A8D926E3FE9C5583" bold="true" box="[199,328,1708,1735]" italics="true" pageId="7" pageNumber="90">Etymology.</emphasis>
The specific name is a patronym in reference to the character El Zorro (fox in Spanish). El Zorro is the secret identity of Don Diego de la Vega, a fictional hero created in 1919 by pulp writer Johnston McCulley. The character has a distinctive black garb, coat, hat and a mask that covers the top of the head from eye level upwards. The name alludes to the facial mask of the new species.
</paragraph>
<paragraph id="8BC1701AFF98F629A8D92772FB1B5189" blockId="7.[151,1437,1456,2024]" lastBlockId="8.[151,1437,151,717]" lastPageId="8" lastPageNumber="91" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A8D92772FD9E5413" bold="true" box="[199,586,1852,1879]" pageId="7" pageNumber="90">
<emphasis id="B90AAC08FF98F626A8D92772FD975413" bold="true" box="[199,579,1852,1879]" italics="true" pageId="7" pageNumber="90">Comparison with related species</emphasis>
.
</emphasis>
<taxonomicName id="4C7E0B99FF98F626AA4D2771FCF55413" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[595,801,1854,1879]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF98F626AA4D2771FCF55413" box="[595,801,1854,1879]" italics="true" pageId="7" pageNumber="90">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF98F626AB342772FC575413" box="[810,899,1853,1879]" pageId="7" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13456C9FFF98F626AB8A2772FC3B541C" box="[916,1007,1853,1880]" captionStart="FIGURE 6" captionStartId="10.[151,250,1403,1427]" captionTargetBox="[154,1431,492,1375]" captionTargetId="figure-192@10.[151,1435,489,1379]" captionTargetPageId="10" captionText="FIGURE 6. Adult males of Pristimantis zorro sp. nov. and related species: (A) Pristimantis zorro sp. nov, paratype (MHUAA12093); (B) Pristimantis olivaceus ZFMK 67132, SVL 19.7 mm, paratype; (C) Pristimantis pluvialis CORBIDI 11862, SVL 20.5 mm, paratype; (D) Pristimantis pulchridormientes CORBIDI 15565, adult male. Photos: (A) MRC; (B) Jörn Köhler; (C) Alessandro Catenazzi; (D) Germán Chávez." figureDoi="http://doi.org/10.5281/zenodo.4403647" httpUri="https://zenodo.org/record/4403647/files/figure.png" pageId="7" pageNumber="90">Fig. 6A</figureCitation>
) is easily distinguished from most of its congeneric by its color pattern, having a light-green to green-yellow dorsum with well-defined canthal stripe, a blotch covering the front of the face, including inter-nostrils area forming a “mask”, an inter-orbital brown bar or blotches and supra-tympanic brown stripe. Another diagnostic feature that is present in 
<taxonomicName id="4C7E0B99FF98F626AC6827E5FB1B5487" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[1142,1231,1962,1987]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF98F626AC6827E5FB1B5487" box="[1142,1231,1962,1987]" italics="true" pageId="7" pageNumber="90">P. zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF98F626ACC827E6FAFA5487" box="[1238,1326,1961,1987]" pageId="7" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
is having blotches on the wrist of the hand. Only two related species, 
<taxonomicName id="4C7E0B99FF98F626AB2E2781FBFA54AC" authority="(Savage 1965)" baseAuthorityName="Savage" baseAuthorityYear="1965" box="[816,1070,1997,2024]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="moro">
<emphasis id="B90AAC08FF98F626AB2E2781FC5154A3" box="[816,901,1998,2023]" italics="true" pageId="7" pageNumber="90">P. moro</emphasis>
(
<bibRefCitation id="EFEF0DEBFF98F626AB8A2782FBF254AC" author="Savage, J. M." box="[916,1062,1997,2024]" pageId="7" pageNumber="90" pagination="106 - 110" refId="ref9082" refString="Savage, J. M. (1965) A new bromeliad frog of the genus Eleutherodactylus from Costa Rica. Bulletin of the Southern California Academy of Sciences, 64, 106 - 110." type="journal article" year="1965">Savage 1965</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C7E0B99FF98F626AC7A2781FA4854A3" authority="(Duellman 1990)" baseAuthorityName="Duellman" baseAuthorityYear="1990" box="[1124,1436,1997,2023]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="90" phylum="Chordata" rank="species" species="schultei">
<emphasis id="B90AAC08FF98F626AC7A2781FB0254A3" box="[1124,1238,1997,2023]" italics="true" pageId="7" pageNumber="90">P. schultei</emphasis>
(
<bibRefCitation id="EFEF0DEBFF98F626ACFA2782FA4054A3" author="Duellman, W. E." box="[1252,1428,1997,2023]" pageId="7" pageNumber="90" pagination="348 - 350" refId="ref7371" refString="Duellman, W. E. (1990) A new species of Eleutherodactylus from the Andes of northern Peru (Anura: Leptodactylidae). Journal of Herpetology, 24, 348 - 350. https: // doi. org / 10.2307 / 1565048" type="journal article" year="1990">Duellman 1990</bibRefCitation>
)
</taxonomicName>
have a similar pattern on the head, with the “mask” and band to interorbital area in some individuals. However, 
<taxonomicName id="4C7E0B99FF97F629AD9620D8FF055390" baseAuthorityName="Savage" baseAuthorityYear="1965" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="moro">
<emphasis id="B90AAC08FF97F629AD9620D8FF055390" italics="true" pageId="8" pageNumber="91">P. moro</emphasis>
</taxonomicName>
has the upper eyelids and area from halfway between eyes to snout bright orange-red and 
<taxonomicName id="4C7E0B99FF97F629ACCB20F4FA9C5391" baseAuthorityName="Duellman" baseAuthorityYear="1990" box="[1237,1352,187,213]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="schultei">
<emphasis id="B90AAC08FF97F629ACCB20F4FA9C5391" box="[1237,1352,187,213]" italics="true" pageId="8" pageNumber="91">P. schultei</emphasis>
</taxonomicName>
having upper eyelids and flanks with tubercles (
<bibRefCitation id="EFEF0DEBFF97F629AA412090FCC653BD" author="Duellman, W. E." box="[607,786,223,249]" pageId="8" pageNumber="91" pagination="348 - 350" refId="ref7371" refString="Duellman, W. E. (1990) A new species of Eleutherodactylus from the Andes of northern Peru (Anura: Leptodactylidae). Journal of Herpetology, 24, 348 - 350. https: // doi. org / 10.2307 / 1565048" type="journal article" year="1990">Duellman 1990</bibRefCitation>
, 
<bibRefCitation id="EFEF0DEBFF97F629AB032090FC2653BD" author="Beraun, A. M. &amp; Duran, V. &amp; Alvarez, S. C. &amp; Venegas, P. J." box="[797,1010,223,249]" pageId="8" pageNumber="91" pagination="281 - 285" refId="ref7215" refString="Beraun, A. M., Duran, V., Alvarez, S. C. &amp; Venegas, P. J. (2014) Distribution extension and an updated map for Pristimantis corrugatus, P. schultei and P. wagteri in northern Peru (Amphibia, Anura, Craugastoridae). Herpetology Notes, 7, 281 - 285." type="journal article" year="2014">
Beraún 
<emphasis id="B90AAC08FF97F629AB682090FC7353BD" box="[886,935,223,249]" italics="true" pageId="8" pageNumber="91">et al</emphasis>
. 2014
</bibRefCitation>
), absent structures in 
<taxonomicName id="4C7E0B99FF97F629ACF82090FAEA53BC" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[1254,1342,223,248]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF97F629ACF82090FAEA53BC" box="[1254,1342,223,248]" italics="true" pageId="8" pageNumber="91">P. zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF97F629AD582090FA4953BD" box="[1350,1437,223,249]" pageId="8" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C7E0B99FF97F629A889214CFC525259" authority="Shepack, von May, Tito and Catenazzi 2016" authorityName="Shepack, von May, Tito and Catenazzi" authorityYear="2016" box="[151,902,259,285]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B90AAC08FF97F629A889214CFE5C5259" box="[151,392,259,285]" italics="true" pageId="8" pageNumber="91">Pristimantis pluvialis</emphasis>
<bibRefCitation id="EFEF0DEBFF97F629A98E214CFC525259" author="Shepack, A. &amp; von May, R. &amp; Tito, A. &amp; Catenazzi, A." box="[400,902,259,285]" pageId="8" pageNumber="91" pagination="143 - 164" refId="ref9118" refString="Shepack, A., von May, R., Tito, A. &amp; Catenazzi, A. (2016) A new species of Pristimantis (Amphibia, Anura, Craugastoridae) from the foothills of the Andes in Manu National Park, southeastern Peru. ZooKeys, 594, 143 - 164. https: // doi. org / 10.3897 / zookeys. 594.8295" type="journal article" year="2016">Shepack, von May, Tito and Catenazzi 2016</bibRefCitation>
</taxonomicName>
, having body reddish-brown (
<figureCitation id="13456C9FFF97F629ACC7214CFAE5525A" box="[1241,1329,259,286]" captionStart="FIGURE 6" captionStartId="10.[151,250,1403,1427]" captionTargetBox="[154,1431,492,1375]" captionTargetId="figure-192@10.[151,1435,489,1379]" captionTargetPageId="10" captionText="FIGURE 6. Adult males of Pristimantis zorro sp. nov. and related species: (A) Pristimantis zorro sp. nov, paratype (MHUAA12093); (B) Pristimantis olivaceus ZFMK 67132, SVL 19.7 mm, paratype; (C) Pristimantis pluvialis CORBIDI 11862, SVL 20.5 mm, paratype; (D) Pristimantis pulchridormientes CORBIDI 15565, adult male. Photos: (A) MRC; (B) Jörn Köhler; (C) Alessandro Catenazzi; (D) Germán Chávez." figureDoi="http://doi.org/10.5281/zenodo.4403647" httpUri="https://zenodo.org/record/4403647/files/figure.png" pageId="8" pageNumber="91">Fig. 6B</figureCitation>
), without interocular and canthal stripe and snout rounded in profile (dorsum light yellow to green-yellow; well-defined canthal stripe and snout truncate in profile in 
<taxonomicName id="4C7E0B99FF97F629AA782104FD685220" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[614,700,331,356]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="zorro">
<emphasis id="B90AAC08FF97F629AA782104FD685220" box="[614,700,331,356]" italics="true" pageId="8" pageNumber="91">P. zorro</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C7E0B99FF97F629AACC2104FAB95221" authority="Chavez and Catenazzi 2016" authorityName="Chavez and Catenazzi" authorityYear="2016" box="[722,1389,331,357]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="pulchridormientes">
<emphasis id="B90AAC08FF97F629AACC2104FBFA5221" box="[722,1070,331,357]" italics="true" pageId="8" pageNumber="91">Pristimantis pulchridormientes</emphasis>
<bibRefCitation id="EFEF0DEBFF97F629AC2B2104FAB95221" author="Chavez, G. &amp; Catenazzi, A." box="[1077,1389,331,357]" pageId="8" pageNumber="91" pagination="113 - 130" refId="ref7280" refString="Chavez, G. &amp; Catenazzi, A. (2016) A new species of frog of the genus Pristimantis from Tingo Maria National Park, Huanuco Department, central Peru (Anura, Craugastoridae). ZooKeys, 610, 113 - 130. https: // doi. org / 10.3897 / zookeys. 610.8507" type="journal article" year="2016">Chávez and Catenazzi 2016</bibRefCitation>
</taxonomicName>
, has bright red coloration (
<figureCitation id="13456C9FFF97F629A9942120FE3452CE" box="[394,480,367,394]" captionStart="FIGURE 6" captionStartId="10.[151,250,1403,1427]" captionTargetBox="[154,1431,492,1375]" captionTargetId="figure-192@10.[151,1435,489,1379]" captionTargetPageId="10" captionText="FIGURE 6. Adult males of Pristimantis zorro sp. nov. and related species: (A) Pristimantis zorro sp. nov, paratype (MHUAA12093); (B) Pristimantis olivaceus ZFMK 67132, SVL 19.7 mm, paratype; (C) Pristimantis pluvialis CORBIDI 11862, SVL 20.5 mm, paratype; (D) Pristimantis pulchridormientes CORBIDI 15565, adult male. Photos: (A) MRC; (B) Jörn Köhler; (C) Alessandro Catenazzi; (D) Germán Chávez." figureDoi="http://doi.org/10.5281/zenodo.4403647" httpUri="https://zenodo.org/record/4403647/files/figure.png" pageId="8" pageNumber="91">Fig. 6C</figureCitation>
) on groins and on the posterior surfaces of thighs and shanks, and dentigerous process of the vomer absent (surfaces of thighs and shanks are yellow and dentigerous process are present in 
<taxonomicName id="4C7E0B99FF97F629ACF721DCFAEB52E8" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[1257,1343,403,428]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="zorro">
<emphasis id="B90AAC08FF97F629ACF721DCFAEB52E8" box="[1257,1343,403,428]" italics="true" pageId="8" pageNumber="91">P. zorro</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C7E0B99FF97F629AD4B21DCFEF05294" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF97F629AD4B21DCFEF05294" italics="true" pageId="8" pageNumber="91">Pristimantis zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF97F629A93421F8FE545295" box="[298,384,439,465]" pageId="8" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
superficially resembles 
<taxonomicName id="4C7E0B99FF97F629AA9121F8FC305295" authority="(Lynch, 1979)" baseAuthorityName="Lynch" baseAuthorityYear="1979" box="[655,996,439,465]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="bromeliaceus">
<emphasis id="B90AAC08FF97F629AA9121F8FC945295" box="[655,832,439,465]" italics="true" pageId="8" pageNumber="91">P. bromeliaceus</emphasis>
(
<bibRefCitation id="EFEF0DEBFF97F629AB5021F8FC085295" author="Lynch, J. D." box="[846,988,439,465]" pageId="8" pageNumber="91" pagination="1 - 62" refId="ref8180" refString="Lynch, J. D. (1979) Leptodactylid frogs of the genus Eleutherodactylus from the Andes of southern Ecuador. Miscellaneous Publication. Museum of Natural History, University of Kansas, 66, 1 - 62. https: // doi. org / 10.5962 / bhl. title. 16268" type="journal article" year="1979">Lynch, 1979</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C7E0B99FF97F629AC0721F8FA855295" authority="(Duellman 1978)" baseAuthorityName="Duellman" baseAuthorityYear="1978" box="[1049,1361,439,465]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="mendax">
<emphasis id="B90AAC08FF97F629AC0721F8FB5F5295" box="[1049,1163,439,465]" italics="true" pageId="8" pageNumber="91">P. mendax</emphasis>
(
<bibRefCitation id="EFEF0DEBFF97F629AC8721F8FA9D5295" author="Duellman, W. E." box="[1177,1353,439,465]" pageId="8" pageNumber="91" pagination="264 - 270" refId="ref7338" refString="Duellman, W. E. (1978) Three new species of Eleutherodactylus from Amazonian Peru (Amphibia: Anura: Leptodactylidae). Herpetologica, 34, 264 - 270." type="journal article" year="1978">Duellman 1978</bibRefCitation>
)
</taxonomicName>
which differs in having prominent tubercles on the eyelid, dorsum and flanks (absent in 
<taxonomicName id="4C7E0B99FF97F629AC012194FBA252B0" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[1055,1142,475,500]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF97F629AC012194FBA252B0" box="[1055,1142,475,500]" italics="true" pageId="8" pageNumber="91">P. zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF97F629AC622194FB0252B1" box="[1148,1238,475,501]" pageId="8" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
), and 
<taxonomicName id="4C7E0B99FF97F629AD062194FCDE515D" authority="(Kohler, Morales, Lotters, Reichle, and Aparicio, 1998)" baseAuthorityName="Kohler, Morales, Lotters, Reichle, and Aparicio" baseAuthorityYear="1998" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AAC08FF97F629AD062194FA4852B1" box="[1304,1436,475,501]" italics="true" pageId="8" pageNumber="91">P. olivaceus</emphasis>
(
<bibRefCitation id="EFEF0DEBFF97F629A88121B0FCD6515D" author="Kohler, J. &amp; Morales, V. R. &amp; Lotters, S. &amp; Reichle, S. &amp; Aparicio, J." box="[159,770,511,537]" pageId="8" pageNumber="91" pagination="93 - 99" refId="ref8051" refString="Kohler, J., Morales, V. R., Lotters, S., Reichle, S. &amp; Aparicio, J. (1998) A new green species of frog, genus Eleutherodactylus, from Bolivia and Peru (Amphibia, Anura, Leptodactylidae). Studies on Neotropical Fauna and Environment, 33, 93 - 99. https: // doi. org / 10.1076 / snfe. 33.2.93.2158" type="journal article" year="1998">Köhler, Morales, Lötters, Reichle, and Aparicio, 1998</bibRefCitation>
)
</taxonomicName>
which has acuminate and protruding snout (subacuminate in dorsal view and truncate in profile in 
<taxonomicName id="4C7E0B99FF97F629AA43226CFD615178" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[605,693,547,572]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF97F629AA43226CFD615178" box="[605,693,547,572]" italics="true" pageId="8" pageNumber="91">P. zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF97F629AAA2226CFCC35179" box="[700,791,547,573]" pageId="8" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
). The “green” Amazonian clade composed by 
<taxonomicName id="4C7E0B99FF97F629AD3F226CFECC5124" class="Amphibia" family="Craugastoridae" genus="Pristimatis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B90AAC08FF97F629AD3F226CFECC5124" italics="true" pageId="8" pageNumber="91">Pristimatis acuminatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E0B99FF97F629A9392208FE3B5125" box="[295,495,583,609]" class="Amphibia" family="Craugastoridae" genus="Pristimatis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="limoncochensis">
<emphasis id="B90AAC08FF97F629A9392208FE3B5125" box="[295,495,583,609]" italics="true" pageId="8" pageNumber="91">P. limoncochensis</emphasis>
</taxonomicName>
, 
<emphasis id="B90AAC08FF97F629A9E32208FDBB5125" box="[509,623,583,609]" italics="true" pageId="8" pageNumber="91">
<taxonomicName id="4C7E0B99FF97F629A9E32208FDBF5125" box="[509,619,583,609]" class="Amphibia" family="Craugastoridae" genus="Pristimatis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="padiali">P. padiali</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C7E0B99FF97F629AAB82208FCEC5125" box="[678,824,583,609]" class="Amphibia" family="Craugastoridae" genus="Pristimatis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="omeviridis">
<emphasis id="B90AAC08FF97F629AAB82208FCEC5125" box="[678,824,583,609]" italics="true" pageId="8" pageNumber="91">P. omeviridis</emphasis>
</taxonomicName>
have well-defined canthal stripe and some individuals also having supratympanic stripe, but lacking a differentiated tympanic annulus and tympanic membrane, which are covered by muscle (
<bibRefCitation id="EFEF0DEBFF97F629A96322C0FD6851ED" author="Ortega-Andrade, H. M. &amp; Rojas-Soto, O. R. &amp; Valencia J. H. &amp; Monteros, A. &amp; Morrone, J. J. &amp; Ron, S. R. &amp; Cannatella D. C." box="[381,700,654,681]" pageId="8" pageNumber="91" pagination="0143392" refId="ref8363" refString="Ortega-Andrade, H. M., Rojas-Soto, O. R., Valencia J. H., Espinosa de los Monteros, A. Morrone, J. J., Ron, S. R. &amp; Cannatella D. C. (2015) Insights from integrative systematics reveal cryptic diversity in Pristimantis frogs (Anura: Craugastoridae) from the Upper Amazon Basin. PLoS One, 10 (11), e 0143392, 1 - 43. https: // doi. org / 10.1371 / journal. pone. 0143392" type="journal article" year="2015">
Ortega-Andrade 
<emphasis id="B90AAC08FF97F629AA2322C0FDA351ED" box="[573,631,655,681]" italics="true" pageId="8" pageNumber="91">et al.</emphasis>
2015
</bibRefCitation>
). 
<taxonomicName id="4C7E0B99FF97F629AACE22C0FC3D51EC" box="[720,1001,655,680]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="enigmaticus">
<emphasis id="B90AAC08FF97F629AACE22C0FC3D51EC" box="[720,1001,655,680]" italics="true" pageId="8" pageNumber="91">Pristimantis enigmaticus</emphasis>
</taxonomicName>
has a differentiated tympanic annulus but lacks vocal slits and vocal sac 
<bibRefCitation id="EFEF0DEBFF97F629AA0622FCFCB55189" author="Ortega-Andrade, H. M. &amp; Rojas-Soto, O. R. &amp; Valencia J. H. &amp; Monteros, A. &amp; Morrone, J. J. &amp; Ron, S. R. &amp; Cannatella D. C." box="[536,865,690,717]" pageId="8" pageNumber="91" pagination="0143392" refId="ref8363" refString="Ortega-Andrade, H. M., Rojas-Soto, O. R., Valencia J. H., Espinosa de los Monteros, A. Morrone, J. J., Ron, S. R. &amp; Cannatella D. C. (2015) Insights from integrative systematics reveal cryptic diversity in Pristimantis frogs (Anura: Craugastoridae) from the Upper Amazon Basin. PLoS One, 10 (11), e 0143392, 1 - 43. https: // doi. org / 10.1371 / journal. pone. 0143392" type="journal article" year="2015">
Ortega-Andrade 
<emphasis id="B90AAC08FF97F629AACB22FCFCD95189" box="[725,781,691,717]" italics="true" pageId="8" pageNumber="91">et al.</emphasis>
(2015)
</bibRefCitation>
, both present in 
<taxonomicName id="4C7E0B99FF97F629AC0422FCFBA65188" authority="Rivera-Correa &amp; Daza, 2020" authorityName="Rivera-Correa &amp; Daza" authorityYear="2020" box="[1050,1138,691,716]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="91" phylum="Chordata" rank="species" species="zorro" status="sp. nov.">
<emphasis id="B90AAC08FF97F629AC0422FCFBA65188" box="[1050,1138,691,716]" italics="true" pageId="8" pageNumber="91">P. zorro</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2391173FF97F629AC6622FCFB1B5189" box="[1144,1231,691,717]" pageId="8" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>