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<document id="BFCBB3FE46161C3F149BA23FC6BC711F" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D13924BFDD5F983C9C72F8A" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius aenescens Heller 1910" docType="treatment" docVersion="10" lastPageNumber="17" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="16" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3DE1C654CB06800CAFAEDBF4DFE5A856">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="E69F5B6D0FEA863E7ED84BF14DE42F30">Telnov, Dmitry</mods:namePart>
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<mods:title id="A04DA1A4732E00B8ECE0ABC0067E39FC">European Journal of Taxonomy</mods:title>
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<mods:date id="04BDF2EE4C0A3B3EC4AE3C0C9C5F0DA9">2020</mods:date>
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<mods:number id="186357D98FA347A6FABCFCC1BEA58F9E">2020-10-19</mods:number>
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<mods:classification id="DA74D5B2CD18513B2FBECC170A190E89">journal article</mods:classification>
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<mods:identifier id="817C3D47B1415B90C8564B5187568833" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D13924BFDD5F983C9C72F8A" ID-DOI="http://doi.org/10.5281/zenodo.4344265" ID-GBIF-Taxon="168648130" ID-Zenodo-Dep="4344265" LSID="urn:lsid:plazi:treatment:1A3787F69D13924BFDD5F983C9C72F8A" httpUri="http://treatment.plazi.org/id/1A3787F69D13924BFDD5F983C9C72F8A" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="DA84656B9D139254FDD5F983CBB82E3A" box="[582,1006,1597,1624]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="922136E09D139254FDD5F983CBB82E3A" blockId="15.[581,1006,1597,1657]" box="[582,1006,1597,1624]" pageId="15" pageNumber="16">
<heading id="C969818C9D139254FDD5F983CBB82E3A" box="[582,1006,1597,1624]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<taxonomicName id="559E4D639D139254FDD5F983CBB82E3A" ID-CoL="3XFH6" authority="Heller, 1910" authorityName="Heller" authorityYear="1910" box="[582,1006,1597,1624]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="aenescens">
<emphasis id="A0EAEAF29D139254FDD5F983CB032E3A" bold="true" box="[582,853,1597,1624]" italics="true" pageId="15" pageNumber="16">Maechidius aenescens</emphasis>
<bibRefCitation id="F60F4B119D139254FCCFF980CBB82E3A" author="Heller K. M." box="[860,1006,1598,1624]" pageId="15" pageNumber="16" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D139254FDD6F9E0CBB82E1B" box="[581,1006,1630,1657]" pageId="15" pageNumber="16" type="description">
<paragraph id="922136E09D139254FDD6F9E0CBB82E1B" blockId="15.[581,1006,1597,1657]" box="[581,1006,1630,1657]" pageId="15" pageNumber="16">
<figureCitation id="0AA52A659D139254FDD6F9E0CAC82E1B" box="[581,670,1630,1657]" captionStart="Figs 18–21" captionStartId="17.[188,240,1928,1954]" captionTargetBox="[353,1214,268,1904]" captionTargetId="figure-12@17.[346,1240,250,1910]" captionTargetPageId="17" captionText="Figs 18–21. Papuan Maechidius Macleay, 1819, habitus, dorsal and lateral views. 18. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 19. M. aiyura sp. nov., holotype, ♂ (BMNH). 20–21. M. alesbezdeki sp. nov. ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122130" httpUri="https://zenodo.org/record/4122130/files/figure.png" pageId="15" pageNumber="16">Figs 18</figureCitation>
,
<figureCitation id="0AA52A659D139254FD38F9E1CA9D2E1B" box="[683,715,1631,1657]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="15" pageNumber="16">97</figureCitation>
,
<figureCitation id="0AA52A659D139254FD44F9E1CB532E1B" box="[727,773,1631,1657]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="15" pageNumber="16">197</figureCitation>
,
<figureCitation id="0AA52A659D139254FC81F9E0CB172E1A" box="[786,833,1630,1656]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="15" pageNumber="16">266</figureCitation>
,
<figureCitation id="0AA52A659D139254FCDEF9E0CB2A2E1A" box="[845,892,1630,1656]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="15" pageNumber="16">286</figureCitation>
, 359, 451
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D13924BFF2EF923CB362ACD" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="922136E09D139254FF2EF923C93A2ED5" blockId="15.[189,1399,1693,1829]" box="[189,364,1693,1719]" pageId="15" pageNumber="16">
<heading id="C969818C9D139254FF2EF923C93A2ED5" bold="true" box="[189,364,1693,1719]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="A0EAEAF29D139254FF2EF923C93A2ED5" bold="true" box="[189,364,1693,1719]" pageId="15" pageNumber="16">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D139254FF79F977CD322F46" blockId="15.[189,1399,1693,1829]" pageId="15" pageNumber="16">
<materialsCitation id="22F63CBD9D139254FF79F977CD322F46" ID-GBIF-Occurrence="2898878512" collectionCode="SNSD" collectorName="Kais. Wilhelmsland Paup Dr. Schlaginhaufen" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<emphasis id="A0EAEAF29D139254FF79F977C9302E81" bold="true" box="[234,358,1737,1763]" pageId="15" pageNumber="16">
<typeStatus id="4D2588429D139254FF79F977C9302E81" box="[234,358,1737,1763]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
<collectingCountry id="EA8976709D139254FF2EF954C98A2F66" box="[189,476,1770,1796]" name="Papua New Guinea" pageId="15" pageNumber="16">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D139254FE64F954CA582F66" box="[503,526,1770,1796]" pageId="15" pageNumber="16" type="female">♀</specimenCount>
; “
<collectorName id="3F6B53369D139254FDBBF954CC642F66" box="[552,1074,1770,1796]" pageId="15" pageNumber="16">Kais.Wilhelmsland Paup Dr.Schlaginhaufen</collectorName>
[p, blue label] //1910 [p] 1 [h] [blue label] //
<taxonomicName id="559E4D639D139254FE16F8B5CA1A2F46" authority="Heller" authorityName="Heller" authorityYear="1910" box="[389,588,1802,1828]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="aenescens">
<emphasis id="A0EAEAF29D139254FE16F8B5C9AA2F46" box="[389,508,1803,1828]" italics="true" pageId="15" pageNumber="16">aenescens</emphasis>
Heller
</taxonomicName>
[h, red label] // Staatl. Museum für Tierkunde.Dresden [p]”;
<collectionCode id="F48FAE259D139254FA82F8B4CD362F46" box="[1297,1376,1802,1828]" collectionName="SNSD" pageId="15" pageNumber="16">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D139254FF79F8F7C9F42F01" blockId="15.[189,1399,1865,2025]" box="[234,418,1865,1891]" pageId="15" pageNumber="16">
<emphasis id="A0EAEAF29D139254FF79F8F7C9F42F01" bold="true" box="[234,418,1865,1891]" pageId="15" pageNumber="16">
<typeStatus id="4D2588429D139254FF79F8F7C9F42F01" box="[234,418,1865,1891]" pageId="15" pageNumber="16" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D139254FF2EF8D4C9E12F8A" blockId="15.[189,1399,1865,2025]" pageId="15" pageNumber="16">
<materialsCitation id="22F63CBD9D139254FF2EF8D4CBA52FC6" ID-GBIF-Occurrence="2898878618" collectionCode="SNSD" collectorName="Kais. Wilhelmsland Paup Dr. Schlaginhaufen" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="15" pageNumber="16" specimenCount="3" specimenCount-female="3" typeStatus="paralectotype">
<collectingCountry id="EA8976709D139254FF2EF8D4C98E2FE6" box="[189,472,1898,1924]" name="Papua New Guinea" pageId="15" pageNumber="16">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D139254FE63F8D4CA642FE6" box="[496,562,1898,1924]" pageId="15" pageNumber="16" type="female">3 ♀♀</specimenCount>
; “
<collectorName id="3F6B53369D139254FDD9F8D7CC072FE1" box="[586,1105,1897,1924]" pageId="15" pageNumber="16">Kais.Wilhelmsland Paup Dr.Schlaginhaufen</collectorName>
[p, blue label] // 1910 [p] 1 [h] [blue label]// Staatl. Museum für Tierkunde.Dresden [p]”;
<collectionCode id="F48FAE259D139254FC3BF834CBA52FC6" box="[936,1011,1930,1956]" collectionName="SNSD" pageId="15" pageNumber="16">SNSD</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D139254FB9FF834C9E22F8A" ID-GBIF-Occurrence="2898878449" collectionCode="SNSD" collectorName="Kais. Wilhelmsland Paup Dr. Schlaginhaufen" country="Papua New Guinea" location="Staatl. Museum fur Tierkunde. Dresden" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<specimenCount id="8498FD699D139254FB9FF834CC6F2FC7" box="[1036,1081,1930,1957]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
; “
<collectorName id="3F6B53369D139254FBC2F834C9CE2FA6" pageId="15" pageNumber="16">Kais.Wilhelmsland Paup Dr.Schlaginhaufen</collectorName>
[p, blue label] // 1910 [p] 1 [h] [blue label] //
<specimenCount id="8498FD699D139254FC27F815CB9C2FA7" box="[948,970,1963,1989]" pageId="15" pageNumber="16" type="female">♀</specimenCount>
[h]//
<location id="9741603B9D139254FB9FF814C9492F8A" LSID="urn:lsid:plazi:treatment:1A3787F69D13924BFDD5F983C9C72F8A:9741603B9D139254FB9FF814C9492F8A" country="Papua New Guinea" name="Staatl. Museum fur Tierkunde. Dresden" pageId="15" pageNumber="16">Staatl. Museum für Tierkunde. Dresden</location>
[p]”;
<collectionCode id="F48FAE259D139254FEF6F870C9E22F8A" box="[357,436,1998,2024]" collectionName="SNSD" pageId="15" pageNumber="16">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D0C924BFF2EFEB7C9302940" blockId="16.[189,358,264,291]" box="[189,358,264,291]" pageId="16" pageNumber="17">
<heading id="C969818C9D0C924BFF2EFEB7C9302940" bold="true" box="[189,358,264,291]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="A0EAEAF29D0C924BFF2EFEB7C9302940" bold="true" box="[189,358,264,291]" pageId="16" pageNumber="17">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0C924BFF2EFE85CD4D291F" blockId="16.[189,1399,315,382]" pageId="16" pageNumber="17">
<materialsCitation id="22F63CBD9D0C924BFF2EFE85CD41291F" ID-GBIF-Occurrence="2898878307" collectingDate="2000-12-16" collectionCode="NHMB" collectorName="L. Cizek" country="Papua New Guinea" latitude="-4.7" location="Salemben village" longLatPrecision="1306" longitude="145.4" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" stateProvince="Madang Prov.">
<collectingCountry id="EA8976709D0C924BFF2EFE85C9B22937" box="[189,484,315,342]" name="Papua New Guinea" pageId="16" pageNumber="17">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D0C924BFD9BFE85CA6C2934" box="[520,570,315,342]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
; “Collection Naturhistorisches Museum Basel // Papua New Guinea
<collectingRegion id="505AF8029D0C924BFF2EFEDDC934291F" box="[189,354,355,381]" country="Papua New Guinea" name="Madang" pageId="16" pageNumber="17">Madang Prov.</collectingRegion>
<collectorName id="3F6B53369D0C924BFEFAFEDDC99B291F" box="[361,461,355,381]" pageId="16" pageNumber="17">L. Cizek</collectorName>
lgt. //
<location id="9741603B9D0C924BFD80FEDDCA88291F" LSID="urn:lsid:plazi:treatment:1A3787F69D13924BFDD5F983C9C72F8A:9741603B9D0C924BFD80FEDDCA88291F" box="[531,734,355,381]" country="Papua New Guinea" latitude="-4.7" longLatPrecision="1306" longitude="145.4" name="Salemben village" pageId="16" pageNumber="17" stateProvince="Madang Prov.">Salemben village</location>
<geoCoordinate id="F7AA50279D0C924BFD76FEDDCB00291C" box="[741,854,355,382]" degrees="145" direction="east" minutes="24" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="145.4">145°24’E</geoCoordinate>
<geoCoordinate id="F7AA50279D0C924BFCCEFEDDCBF9291F" box="[861,943,355,381]" degrees="4" direction="south" minutes="42" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="-4.7">4°42’S</geoCoordinate>
<date id="E62010209D0C924BFC25FEDDCC1E291F" box="[950,1096,355,381]" pageId="16" pageNumber="17" value="2000-12-16">
<collectingDate id="F664E9C89D0C924BFC25FEDDCC1E291F" box="[950,1096,355,381]" pageId="16" pageNumber="17" value="2000-12-16">
16.XII.
<quantity id="55669B059D0C924BFB94FEDDCC1E291F" box="[1031,1096,355,381]" metricMagnitude="6" metricUnit="m" metricValue="2.00075" pageId="16" pageNumber="17" unit="m" value="2000750.0">2000</quantity>
</collectingDate>
</date>
, 750 m”;
<collectionCode id="F48FAE259D0C924BFB28FEDDCD41291F" box="[1211,1303,355,381]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="17">NHMB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D0C924BFF2EFE16CB362ACD" blockId="16.[189,1399,424,688]" pageId="16" pageNumber="17">
<bibRefCitation id="F60F4B119D0C924BFF2EFE16C9D429A1" author="Heller K. M." box="[189,386,424,451]" pageId="16" pageNumber="17" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller (1910: 22)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D0C924BFD04FE17CB6729A0" authorityName="Heller" authorityYear="1910" box="[663,817,425,450]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aenescens">
<emphasis id="A0EAEAF29D0C924BFD04FE17CB6729A0" box="[663,817,425,450]" italics="true" pageId="16" pageNumber="17">M.aenescens</emphasis>
</taxonomicName>
on a number of specimens –although not explicitly stated, he gives a range of sizes. The four
<typeStatus id="4D2588429D0C924BFD2FFE6ECB332988" box="[700,869,464,490]" pageId="16" pageNumber="17" type="paralectotype">paralectotypes</typeStatus>
(
<collectionCode id="F48FAE259D0C924BFCE4FE6ECB932988" box="[887,965,464,490]" pageId="16" pageNumber="17">SNSD</collectionCode>
) do not bear a type label. However, the locality label has the same text and handwriting as in the only specimen labelled “
<taxonomicName id="559E4D639D0C924BFB25FE46CD7B2A73" authorityName="Heller" authorityYear="1910" box="[1206,1325,504,529]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aenescens">
<emphasis id="A0EAEAF29D0C924BFB25FE46CD7B2A73" box="[1206,1325,504,529]" italics="true" pageId="16" pageNumber="17">aenescens</emphasis>
</taxonomicName>
”. The
<typeStatus id="4D2588429D0C924BFF2EFDA1C97F2A5B" box="[189,297,543,569]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
here designated is the only
<typeStatus id="4D2588429D0C924BFDFDFDA1CA9F2A5B" box="[622,713,543,569]" pageId="16" pageNumber="17" type="syntype">syntype</typeStatus>
in Heller’s original handwriting on a red label, which may indicate (but cannot be stated with confidence), that this specimen was selected by the original author as the
<typeStatus id="4D2588429D0C924BFF74FDD0C9042AEA" box="[231,338,622,648]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
. The
<typeStatus id="4D2588429D0C924BFE1CFDD0C9AD2AEA" box="[399,507,622,648]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
designation is made in order to enhance the stability of nomenclature and fix all 5 of Heller’s specimens as members of the type series.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0C924BFF2EFD65C9C72F8A" pageId="16" pageNumber="17" type="description">
<paragraph id="922136E09D0C924BFF2EFD65C9182A97" blockId="16.[189,334,731,757]" box="[189,334,731,757]" pageId="16" pageNumber="17">
<heading id="C969818C9D0C924BFF2EFD65C9182A97" bold="true" box="[189,334,731,757]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="A0EAEAF29D0C924BFF2EFD65C9182A97" bold="true" box="[189,334,731,757]" pageId="16" pageNumber="17">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0C924BFF2EFCB0CCDF2B4A" blockId="16.[189,1161,782,809]" box="[189,1161,782,809]" pageId="16" pageNumber="17">
MEASUREMENTS. Total body length 6 (
<typeStatus id="4D2588429D0C924BFDFDFCB0CA8B2B4B" box="[622,733,782,809]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
) to 7.40 (largest
<typeStatus id="4D2588429D0C924BFC37FCB0CC122B4B" box="[932,1092,782,809]" pageId="16" pageNumber="17" type="paralectotype">paralectotype</typeStatus>
) mm.
</paragraph>
<paragraph id="922136E09D0C924BFF2EFCEACA642F13" blockId="16.[189,1399,851,1905]" pageId="16" pageNumber="17">
Dorsum and venter brown, forebody with slight green lustre. Head flattened dorsally, somewhat depressed on either side at anterior angle of labroclypeus, slightly glossy dorsally and ventrally. Female labroclypeus subtruncate anteriorly (
<figureCitation id="0AA52A659D0C924BFDF4FC1CCAEB2BDF" box="[615,701,930,957]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="16" pageNumber="17">Fig. 97</figureCitation>
), its lateral margins sinuous in both dorsal and lateral views, anterolateral angles large, obtuse angulate and raised up at angle of nearly 90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus broadly rounded to obtuse angulate in dorsal view. Punctures of frons annular, large and shallow. Intervening spaces with very delicate microreticulation, generally larger than punctures. Pubescence scale-like, elongate, variably long. Some setae very short, much shorter than their corresponding punctures. Each seta rises from anterior margin of corresponding puncture. Female antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, glossy dorsally and laterally. Anterior margin deeply emarginate with anterolateral angles protruding anteriad. Basal margin sinuous to broadly rounded. Lateral margin broadly rounded, delicately crenulate all along (
<figureCitation id="0AA52A659D0C924BFF55FA93C9782D25" box="[198,302,1325,1351]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="16" pageNumber="17">Fig. 197</figureCitation>
). Moderately long curved and appressed scale-like seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view except in basal fourth. Hypomeron separated from prosternum by low straight carina, with very long setae on its anterolateral margin opposite to compound eye (
<figureCitation id="0AA52A659D0C924BFEEFFA1AC9B32DDC" box="[380,485,1443,1470]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="16" pageNumber="17">Fig. 266</figureCitation>
). Antennal pocket shallow. Disc with shallow annular punctures, intervening spaces glossy and generally larger than those. Setae similar to those on head, appressed, generally longer. Some setae very short, much shorter than their corresponding punctures. Scutellar shield broadly rounded apically. Elytron glossy, with or without tracks of two vague, almost entirely glabrous longitudinal carinae. Sutural carinae not present. Punctures of elytral disc small, ordinary, intervening spaces in part densely wrinkled, in part glossy (
<figureCitation id="0AA52A659D0C924BFDEBF9D7CAB62EE6" box="[632,736,1641,1668]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="16" pageNumber="17">Fig. 286</figureCitation>
). Setae shorter than those on pronotum, appressed; setae arranged in in part irregular paired longitudinal rows. Most setae extending exactly from one puncture to the next. Female pygidium flattened dorsally, with very large dense punctures which are shagreened on background (
<figureCitation id="0AA52A659D0C924BFEE9F961C9892E98" box="[378,479,1759,1786]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="16" pageNumber="17">Fig. 451</figureCitation>
). Intervening spaces of pygidium glossy, much smaller than punctures. Protibia widened distally, in female with three external teeth: two large acute distal and one inconspicuous strongly obtuse basal (Fig. 359). Protibial terminal spur not present (in females only?). Tarsal claws with pulvilli (in females only?).
</paragraph>
<paragraph id="922136E09D0C924BFF2EF825C9E62FD7" blockId="16.[189,432,1947,1973]" box="[189,432,1947,1973]" pageId="16" pageNumber="17">
<heading id="C969818C9D0C924BFF2EF825C9E62FD7" bold="true" box="[189,432,1947,1973]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="A0EAEAF29D0C924BFF2EF825C9E62FD7" bold="true" box="[189,432,1947,1973]" pageId="16" pageNumber="17">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0C924BFF2EF870C9C72F8A" blockId="16.[189,401,1998,2024]" box="[189,401,1998,2024]" pageId="16" pageNumber="17">Male is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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