188 lines
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188 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="58E40872F7FF76217BD75DE372A60CC6" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus profundipygus Caterino & Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="350" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="347" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>271</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>401</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040955" LSID="urn:lsid:zoobank.org:act:E9D13FFA-0FE2-4C58-8A71-32301FC8B8C7" httpUri="http://treatment.plazi.org/id/58E40872F7FF76217BD75DE372A60CC6" lastPageId="349" lastPageNumber="350" pageId="346" pageNumber="347">
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<subSubSection lastPageId="347" lastPageNumber="348" pageId="346" pageNumber="347" type="nomenclature">
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<paragraph lastPageId="347" lastPageNumber="348" pageId="346" pageNumber="347">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:E9D13FFA-0FE2-4C58-8A71-32301FC8B8C7" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus profundipygus" order="Coleoptera" pageId="346" pageNumber="347" phylum="Arthropoda" rank="species" species="profundipygus">Operclipygus profundipygus</taxonomicName>
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<taxonomicNameLabel pageId="347" pageNumber="348">
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<pageBreakToken pageId="347" pageNumber="348" start="start">sp</pageBreakToken>
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. n.
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</taxonomicNameLabel>
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Figs 95
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<normalizedToken originalValue="D–E96C–D">D-E96C-D</normalizedToken>
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, F, HMap 33
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</paragraph>
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</subSubSection>
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<subSubSection pageId="347" pageNumber="348" type="type locality">
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<paragraph pageId="347" pageNumber="348">Type locality.</paragraph>
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<paragraph pageId="347" pageNumber="348">
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GUYANA: Region 8: 1 km W Kurupukari, Iwokrama Field Station [
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="4.672">4°40.32'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-58.6845">58°41.07'W</geoCoordinate>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="347" pageNumber="348" type="type material">
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<paragraph pageId="347" pageNumber="348">Type material.</paragraph>
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<paragraph pageId="347" pageNumber="348">
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Holotype male: "GUYANA: Region 8, Iwokrama Forest, 1 km W Kurupukari, Iwokrama Forest Field Stn., 60 m,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.671944">4°40'19"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-58.684444">58°41'4"W</geoCoordinate>
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, 20-25 MAY 2001 R. Brooks,Z.Falin GUY1BF01 034 ex: flight intercept trap"/ "SM0566205 KUNHM-ENT" (SEMC). Paratypes (29): BRAZIL:
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<normalizedToken originalValue="Pará">Para</normalizedToken>
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: 4: Abaetetuba,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.7">1°42'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-48.9">48°54'W</geoCoordinate>
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, 26.
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<normalizedToken originalValue="xi–">xi-</normalizedToken>
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7.xii.1993, FIT (CEMT, FMNH); 1:Altamira -
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<normalizedToken originalValue="Marabá">Maraba</normalizedToken>
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: km 18.,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.15">3°09'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.05">52°03'W</geoCoordinate>
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, v.1985, FIT (CHND); 1:
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<normalizedToken originalValue="Tucuruí">Tucurui</normalizedToken>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.75">3°45'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-49.666668">49°40'W</geoCoordinate>
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, 6-17.vi.1986, FIT (CHND). GUYANA: Region 8: 2: Iwokrama Field Stn., Iwokrama Forest, 1km W Kurupukari,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.671944">4°40'19"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-58.684444">58°41'4"W</geoCoordinate>
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, 60m, 21.v.2001,
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<taxonomicName class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acromyrmex" order="Hymenoptera" pageId="347" pageNumber="348" phylum="Arthropoda" rank="genus">Acromyrmex</taxonomicName>
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refuse pile, R. Brooks & Z. Falin (SEMC), 3: 26-29.v.2001, FIT, R. Brooks & Z. Falin (SEMC), 1: 20-25.v.2001, FIT, R. Brooks & Z. Falin (SEMC); 5: Kurupukari,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="4.6666665">4°40'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-58.666668">58°40'W</geoCoordinate>
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, ix-xi.1992, malaise/FIT (BMNH). FRENCH GUIANA: 1: Montjoly,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="4.9166665">4°55'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-52.266666">52°16'W</geoCoordinate>
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, xii.1977, banana trap (CHND). SURINAME: Marowijne: 1: Perica, 70km E. Paramaribo on East-West Road,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="5.674444">5°40'28"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-54.60861">54°36'31"W</geoCoordinate>
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, 5m, 29-31.v.1999, FIT, Z. Falin, B. DeDijn (SEMC), 3: 31.
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<normalizedToken originalValue="v–">v-</normalizedToken>
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5.vi.1999, FIT, Z. Falin, B. DeDijn (SEMC), 2: 5m, 31.v.1999, FIT, Z. Falin (MSCC, AKTC); Sipaliwini: 2: CI-RAP Surv. Camp 1: on Kutari River,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="2.17535">2°10.521'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-56.7874">56°47.244'W</geoCoordinate>
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, 228m, 19-24.viii.2010, FIT, T. Larsen & A.E.Z. Short (SEMC); 2: upper Palumeu, 225m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="2.477">2.47700°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-55.62941">55.62941°W</geoCoordinate>
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, CI-RAP Survey camp 1, FIT, 10-16.iii.2012, A.E.Z. Short (MSCC); 1: Wanaboo (near Nason), Marowijne River,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.726389">4°43'35"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-54.443333">54°26'36"W</geoCoordinate>
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, 40m, 31.
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<normalizedToken originalValue="v–">v-</normalizedToken>
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5.vi.1999, FIT, Z. Falin, B. DeDijn (SEMC); Wanica: 1: Paramaribo, 12km S. on I. Ghandiweg, and 4.5km W on Mijnzorgweg,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="5.7319446">5°43'55"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-55.251945">55°15'7"W</geoCoordinate>
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, 10m, 6-7.vi.1999, FIT, Z. Falin, B. DeDijn (SEMC).
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</paragraph>
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<paragraph pageId="347" pageNumber="348">
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Other material. BOLIVIA: Beni: 1: Chacobo Indian Village on Rio Benicito,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.333333">12°20'S</geoCoordinate>
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, 66°W, 18-27.vii.1960, under tapir dung, B. Malkin (FMNH), 1: 31.vii-2.viii.1960, B. Malkin (FMNH). ECUADOR: Orellana: 1:
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<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
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Res. Stn.,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.67444444">00°40'28"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-76.64722">76°38'50"W</geoCoordinate>
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, 215m, 5-10.ix.1999, FIT, Primary forest, E.G. Riley (LSAM); 1: P.N.
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<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
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, Via Maxus at Puente
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<normalizedToken originalValue="Piraña">Pirana</normalizedToken>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.65833336">0°39.5'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.433334">76°26'W</geoCoordinate>
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, 20-24.vii.2008, FIT, A.K. Tishechkin (AKTC). PERU: Loreto: 3: Iquitos, 90m, 7.v.1992, FIT, J. Danoff-Berg (SEMC); Madre de Dios: 1: Amazonas Lodge, N Atalaya,
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-12.87">12°52.2'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-71.37666">71°22.6'W</geoCoordinate>
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, 480m, 10-13.xi.2007, FIT, D. Brzoska (SEMC); 2: Tambopata, Reserva Cuzco Amazonico, 15km NE Pto. Maldonado,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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, 200m, 16.vii.1989, FIT, J. Ashe & R. Leschen (SEMC), 2: 13.vii.1989, FIT, J. Ashe & R. Leschen (SEMC), 1: 20.vi.1989, FIT, J. Ashe & R. Leschen (SEMC), 1: 22.vi.1989, FIT, J. Ashe & R. Leschen (CHSM), 1: 26.vi.1989, FIT, J. Ashe & R. Leschen (CHSM); 1: Manu National Park, Cocha Salvador,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.003612">12°0'13"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.52667">71°31'36"W</geoCoordinate>
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, 310m, 20-21.x.2000, FIT, R. Brooks (SEMC).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="348" lastPageNumber="349" pageId="347" pageNumber="348" type="description">
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<paragraph pageId="347" pageNumber="348">Diagnostic description.</paragraph>
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<paragraph lastPageId="348" lastPageNumber="349" pageId="347" pageNumber="348">
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Length: 1.75-1.90 mm, width: 1.47-1.59 mm; body rufo-brunneus, elongate oval, widest at humeri, prothorax rather broad, narrowing more
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<pageBreakToken pageId="348" pageNumber="349" start="start">conspicuously</pageBreakToken>
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posterad; frons weakly depressed at middle, often with microsculpture within depression; frontal stria with sides divergent between eyes, broadly outwardly arcuate across front, continuous with complete supraorbital stria; labrum about two-thirds as long as broad, apical margin straight; left mandible untoothed, right with acute basal tooth; pronotum with sides weakly arcuate, convergent; pronotal disk with distinct narrow, sublinear prescutellar impression; pronotal disk with ground punctation fine, with ~15-20 coarser punctures toward sides; lateral marginal pronotal stria complete at side, ending just mesad anterior corner; lateral submarginal stria complete at side, extending forward to anterior margin, replacing marginal for a short distance, ending behind eye, slightly overlapping end of anterior submarginal stria, which is briefly recurved posterad at sides; central portion of anterior pronotal margin weakly projecting; median pronotal gland openings situated laterad ends of anterior submarginal stria; elytron swollen laterad 1st dorsal stria, with one complete epipleural stria, outer subhumeral in posterior half to two-thirds, inner subhumeral stria absent, striae 1-4 complete, 5th stria present in apical half and with a large basal puncture; sutural stria present in apical three-fourths, widened toward base; prosternal keel rounded, produced at base, with carinal striae sinuately convergent to front, united just short of presternal suture; prosternal lobe with marginal stria more or less complete; mesoventrite emarginate anteriorly, with complete marginal stria; mesometaventral stria arched forward just beyond mesoventral midpoint, continued at sides by lateral metaventral stria, which extends toward middle of metacoxa; 1st abdominal ventrite with two complete lateral striae on each side; propygidium with fine, dense ground punctation at sides, with small, deep, coarse punctures densely clustered in middle two-thirds; pygidium with fine, dense ground punctures throughout, with slightly larger punctures generally sparse, but concentrated along basal margin and toward apex of disk; apical marginal sulcus complete, deep and coarse. Male genitalia (Figs 96
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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, F, H): accessory sclerites present; T8 with sides convergent from base to apex, basal emargination rather narrowly angulate, basal membrane attachment line distad by about half basal emargination's depth, apical emargination narrow, ventrolateral apodemes subacute ventrally, nearly meeting at middle; S8 with apical guides well developed, consistent in width throughout length, apices very slightly divergent, ventral halves fused for most of their length; T9 with apices narrow, slightly convergent; T10 with halves separate along midline; S9 rather narrow, weakly widened to rounded base, apex concave, but without median emargination, apical flanges continuous; tegmen short with sides strongly rounded, narrow at base, medioventral process weak,
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<normalizedToken originalValue="‘U’">'U'</normalizedToken>
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-shaped, projecting beneath just basad midpoint; basal piece nearly as long as tegmen; median lobe about half length of tegmen, with basal apodemes widely separate at gonopore, convergent toward tegmen base.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="349" lastPageNumber="350" pageId="348" pageNumber="349" type="remarks">
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<paragraph pageId="348" pageNumber="349">Remarks.</paragraph>
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<paragraph lastPageId="349" lastPageNumber="350" pageId="348" pageNumber="349">
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The pattern of propygidial punctation (Fig. 95E), with dense ground punctation visible at sides, while larger punctures are concentrated in the middle is unusual, although there are a few distantly related species that share it. Of these, this one alone has a distinct prescutellar impression (Fig. 95D). We restrict the type series
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<pageBreakToken pageId="349" pageNumber="350" start="start">to</pageBreakToken>
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those specimens from the Guianas and northeastern Amazonian region, though there is little evident variation throughout the range.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="349" pageNumber="350" type="etymology">
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<paragraph pageId="349" pageNumber="350">Etymology.</paragraph>
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<paragraph pageId="349" pageNumber="350">The name of this species refers to both the deep pygidial sulcus and the deeply impressed median propygidial punctures.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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