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<document id="D942D9E253753BEFA92F1DAAAF890004" ID-DOI="10.11646/zootaxa.5293.2.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="7960048" ID-ZooBank="C42E6A04-811B-4E8C-98CE-0E131FDEF330" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1684797505526" checkinUser="plazi" docAuthor="Ahyong, Shane T. &amp; Ng, Peter K. L." docDate="2023" docId="03A2995BFFDFFFF524F7FEC4FA801299" docLanguage="en" docName="zootaxa.5293.2.12.pdf" docOrigin="Zootaxa 5293 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.2.12" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pronotonyx Ward 1936" docType="treatment" docVersion="3" lastPageNumber="394" masterDocId="FF9BE123FFDEFFF72460FFE3FFF5113C" masterDocTitle="On the identity of Pronotonyx laevis (Miers, 1884) (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="400" masterPageNumber="392" pageNumber="393" updateTime="1685035623545" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="4AB17ED0425204AA3C6AE611EC834014">On the identity of Pronotonyx laevis (Miers, 1884) (Crustacea: Brachyura: Pilumnidae)</mods:title>
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<mods:namePart id="C7EF74FF8EDD74FD256B7616FA7E8AB4">Ahyong, Shane T.</mods:namePart>
<mods:affiliation id="20636A6C11143FF815BC39D83B954864">Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney NSW 2010, Australia, and School of Biological, Earth &amp; Environmental Sciences, University of New South Wales, NSW 2052, Australia</mods:affiliation>
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<mods:namePart id="0533AD176A3B41500580274EB4DA8283">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="CD9F99CF5132A881FBB794C5D6CA6CC7">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
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<taxonomicName id="4C0B53CEFFDFFFF624F7FEC4FE43107D" ID-CoL="7PVRS" authority="Ward 1936" authorityName="Ward" authorityYear="1936" box="[151,438,295,321]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF624F7FEC4FE43107D" bold="true" box="[151,438,295,321]" pageId="1" pageNumber="393">
<emphasis id="B97FF45FFFDFFFF624F7FEC4FED0107D" bold="true" box="[151,293,295,321]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
<bibRefCitation id="EF9A55BCFFDFFFF6254CFEC4FE43107D" author="Ward, M." box="[300,438,295,321]" pageId="1" pageNumber="393" pagination="1 - 13" refId="ref5805" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward 1936</bibRefCitation>
</emphasis>
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<treatmentCitation id="0AAA0E5CFFDFFFF624F7FE84FE501042" author="Ward, M." box="[151,421,358,382]" page="2" pageId="1" pageNumber="393" year="1936">
<taxonomicName id="4C0B53CEFFDFFFF624F7FE84FE501042" ID-CoL="7PVRS" authority="Ward, 1936: 2" authorityName="Ward" authorityPageNumber="2" authorityYear="1936" box="[151,421,358,382]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF624F7FE84FEF91042" box="[151,268,359,382]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
<bibRefCitation id="EF9A55BCFFDFFFF62574FE85FE501042" author="Ward, M." box="[276,421,358,382]" pageId="1" pageNumber="393" pagination="1 - 13" refId="ref5805" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936: 2</bibRefCitation>
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.—
<treatmentCitation id="0AAA0E5CFFDFFFF625A4FE85FDB31042" author="Ng, P. K. L." box="[452,582,358,382]" page="98" pageId="1" pageNumber="393" year="1987">
<bibRefCitation id="EF9A55BCFFDFFFF625A4FE85FDB31042" author="Ng, P. K. L." box="[452,582,358,382]" pageId="1" pageNumber="393" pagination="69 - 111" refId="ref4856" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987: 98</bibRefCitation>
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.—
<treatmentCitation id="0AAA0E5CFFDFFFF62604FE85FCE61042" author="Davie, P. J. F." box="[612,787,358,382]" page="426" pageId="1" pageNumber="393" year="2002">
<bibRefCitation id="EF9A55BCFFDFFFF62604FE85FCE61042" author="Davie, P. J. F." box="[612,787,358,382]" pageId="1" pageNumber="393" pagination="1 - 641" refId="ref4228" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca. Eucarida (Part 2). Decapoda - Anomura, Brachyura. In: Wells, A. &amp; Houston, W. W. K. (Eds.), Zoological Catalogue of Australia. Vol. 19.3 B. CSIRO Publishing, Melbourne, pp. i - xiv + 1 - 641." type="book chapter" year="2002">Davie 2002: 426</bibRefCitation>
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.—Ng
<emphasis id="B97FF45FFFDFFFF62738FE84FC781042" box="[856,909,358,382]" italics="true" pageId="1" pageNumber="393">et al.</emphasis>
2008: 144.—
<treatmentCitation id="0AAA0E5CFFDFFFF62077FE85FACF1042" author="Poore, G. C. B. &amp; Ahyong, S. T." box="[1047,1338,358,382]" page="671" pageId="1" pageNumber="393" year="2023">
<bibRefCitation id="EF9A55BCFFDFFFF62077FE85FACF1042" author="Poore, G. C. B. &amp; Ahyong, S. T." box="[1047,1338,358,382]" pageId="1" pageNumber="393" refId="ref5388" refString="Poore, G. C. B. &amp; Ahyong, S. T. (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Clayton South, xii + 916 pp. https: // doi. org / 10.1071 / 9781486311798" type="book" year="2023">Poore &amp; Ahyong 2023: 671</bibRefCitation>
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, 684, fig. 14.112h.
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<emphasis id="B97FF45FFFDFFFF624F7FE47FEEB1087" bold="true" box="[151,286,419,444]" pageId="1" pageNumber="393">
<typeStatus id="54B096EFFFDFFFF624F7FE47FF391080" box="[151,204,420,444]" pageId="1" pageNumber="393">Type</typeStatus>
species
</emphasis>
.
</emphasis>
<taxonomicName id="4C0B53CEFFDFFFF6254AFE47FD951080" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[298,608,420,444]" class="Malacostraca" family="Pilumnidae" genus="Ceratoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B97FF45FFFDFFFF6254AFE47FE281080" box="[298,477,420,444]" italics="true" pageId="1" pageNumber="393">Ceratoplax laevis</emphasis>
<bibRefCitation id="EF9A55BCFFDFFFF62584FE47FD951080" author="Miers, E. J." box="[484,608,420,444]" pageId="1" pageNumber="393" pagination="178 - 322" refId="ref4539" refString="Miers, E. J. (1884) Part I. The collections from Melanesia. Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 2. British Museum (Natural History), London, pp. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
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, by original designation.
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<emphasis id="B97FF45FFFDFFFF624F7FE0AFEFF133F" bold="true" box="[151,266,489,515]" pageId="1" pageNumber="393">Diagnosis</emphasis>
. Frontal margin of carapace and pereopods with scattered setae, not obscuring margins or surface; carapace wide, regions not indicated, dorsal surface appearing smooth, polished; anterolateral margin convex, entire or with anterolateral teeth scarcely demarcated; posterior margin of epistome distinctly sinuous with the lateral parts distinctly concave; endostome with low oblique ridge. Maxilliped 3 merus with anteroexternal angle auriculiform. Chelae with pollex almost as long as or subequal to palm, dactylus inclined at an angle not exceeding 45° to vertical. P2–P5 slender, unarmed, merus shorter than three-fourths carapace length; coxa without denticulate plate. Male thoracic sternum broad, anterior sternum (sternites 1–4) width 1.7 × length; sternopleonal cavity reaching anteriorly to level of midlength of cheliped coxae. Male pleon T-shaped, somite 1 very wide, reaching laterally to P5 coxae; somite 2 distinctly narrower than somites 1 and 3; somite 3 wide, almost reaching condyles of P5 coxae, separated by distinct gap; coxosternal groove for penis partially exposed. G1 strongly sinuous, distal part strongly curved, hooked.
</paragraph>
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<paragraph id="8BB4284DFFDFFFF624A7FC95FEE5175C" blockId="1.[151,1437,489,2028]" pageId="1" pageNumber="393">
<emphasis id="B97FF45FFFDFFFF624A7FC95FEC112AC" bold="true" box="[199,308,886,912]" pageId="1" pageNumber="393">Remarks</emphasis>
. In describing
<taxonomicName id="4C0B53CEFFDFFFF62587FC95FD4512AC" authorityName="Miers" authorityYear="1884" box="[487,688,886,912]" class="Malacostraca" family="Pilumnidae" genus="Ceratoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B97FF45FFFDFFFF62587FC95FD4512AC" box="[487,688,886,912]" italics="true" pageId="1" pageNumber="393">Ceratoplax laevis</emphasis>
</taxonomicName>
from off northern
<collectingCountry id="F31C68DDFFDFFFF627EFFC95FC0C12AC" box="[911,1017,886,912]" name="Australia" pageId="1" pageNumber="393">Australia</collectingCountry>
,
<bibRefCitation id="EF9A55BCFFDFFFF62067FC95FB2F12AD" author="Miers, E. J." box="[1031,1242,886,913]" pageId="1" pageNumber="393" pagination="178 - 322" refId="ref4539" refString="Miers, E. J. (1884) Part I. The collections from Melanesia. Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 2. British Museum (Natural History), London, pp. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers (1884: 245)</bibRefCitation>
commented that it closely resembled
<taxonomicName id="4C0B53CEFFDFFFF625EAFC78FC9F1288" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[394,874,922,948]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B97FF45FFFDFFFF625EAFC78FDCB1288" box="[394,574,922,948]" italics="true" pageId="1" pageNumber="393">Notonyx nitidus</emphasis>
A.
<bibRefCitation id="EF9A55BCFFDFFFF6260EFC79FC9F1288" author="Milne-Edwards, A." box="[622,874,922,948]" pageId="1" pageNumber="393" pagination="155 - 332" refId="ref4606" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
</taxonomicName>
, but differed in “having the carapace, eyes, and chelipedes entirely glabrous, there is apparently no tooth on the upper margin of the arm, and the dactyli of the ambulatory legs are carinated; the carapace is more quadrate, and the merus of the outer maxillipedes longer, not transverse, with the antero-external angle less prominent”.
<bibRefCitation id="EF9A55BCFFDFFFF62751FBE5FC0D151C" author="Tesch, J. J." box="[817,1016,1030,1056]" pageId="1" pageNumber="393" pagination="149 - 295" refId="ref5770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918: 203</bibRefCitation>
, footnote) included the species in his key to
<taxonomicName id="4C0B53CEFFDFFFF62484FBC9FE971578" authorityName="Stimpson" authorityYear="1858" box="[228,354,1066,1092]" class="Malacostraca" family="Pilumnidae" genus="Ceratoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF62484FBC9FE971578" box="[228,354,1066,1092]" italics="true" pageId="1" pageNumber="393">Ceratoplax</emphasis>
</taxonomicName>
but commented that it was “Probably identical with
<taxonomicName id="4C0B53CEFFDFFFF627D3FBC8FAC81578" authority="A. Milne-Edwards" authorityName="A. Milne-Edwards" authorityYear="1873" box="[947,1341,1066,1092]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B97FF45FFFDFFFF627D3FBC8FB961578" box="[947,1123,1066,1092]" italics="true" pageId="1" pageNumber="393">Notonyx nitidus</emphasis>
A. Milne-Edwards
</taxonomicName>
” but did not elaborate. In establishing
<taxonomicName id="4C0B53CEFFDFFFF625BEFBACFD951554" authorityName="Ward" authorityYear="1936" box="[478,608,1103,1128]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF625BEFBACFD951554" box="[478,608,1103,1128]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
</taxonomicName>
based on specimens from
<collectingRegion id="49CFE6AFFFDFFFF627E5FBADFBF91554" box="[901,1036,1102,1128]" country="Australia" name="Queensland" pageId="1" pageNumber="393">Queensland</collectingRegion>
,
<collectingCountry id="F31C68DDFFDFFFF62075FBADFB8A1554" box="[1045,1151,1102,1128]" name="Australia" pageId="1" pageNumber="393">Australia</collectingCountry>
,
<bibRefCitation id="EF9A55BCFFDFFFF620E8FBADFAC71554" author="Ward, M." box="[1160,1330,1102,1128]" pageId="1" pageNumber="393" pagination="1 - 13" refId="ref5805" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward (1936: 2)</bibRefCitation>
appeared to follow Miers’ (1884) arguments and noted that the genus differed in having: the carapace “broader than long, smooth and shining”; the front “more than one-third the width”; “the anterior margin of the buccal frame...entire”; the merus of maxilliped 3 with “the antero-external angle auriculated”; “a strong tooth on the upper border of the merus of the cheliped”; and the first somite of the male pleon that “almost reaches the coxi of the fifth pair” of legs.
<bibRefCitation id="EF9A55BCFFDFFFF624F7FAE1FECE1420" author="Ng, P. K. L." box="[151,315,1282,1308]" pageId="1" pageNumber="393" pagination="69 - 111" refId="ref4856" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng (1987: 98)</bibRefCitation>
, in reviewing the genera, provisionally retained
<taxonomicName id="4C0B53CEFFDFFFF62700FAE0FC171420" authorityName="Ward" authorityYear="1936" box="[864,994,1283,1308]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF62700FAE0FC171420" box="[864,994,1283,1308]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C0B53CEFFDFFFF62052FAE1FB471420" authorityName="Stimpson" authorityYear="1858" box="[1074,1202,1282,1308]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="392" phylum="Arthropoda" rank="subFamily" subFamily="Rhizopinae">Rhizopinae</taxonomicName>
(
<taxonomicName id="4C0B53CEFFDFFFF620A1FAE1FAB21420" authorityName="Samouelle" authorityYear="1819" box="[1217,1351,1282,1308]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="family">Pilumnidae</taxonomicName>
), as the condition of the diagnostic male gonopods was not known. Our detailed examination of the specimens of
<taxonomicName id="4C0B53CEFFDFFFF6215FFAC4FA69147C" authorityName="Miers" authorityYear="1884" box="[1343,1436,1318,1344]" class="Malacostraca" family="Pilumnidae" genus="Ceratoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B97FF45FFFDFFFF6215FFAC4FA69147C" box="[1343,1436,1318,1344]" italics="true" pageId="1" pageNumber="393">P. laevis</emphasis>
</taxonomicName>
at hand corroborates its placement in the
<taxonomicName id="4C0B53CEFFDFFFF62609FAA9FD191458" authorityName="Samouelle" authorityYear="1819" box="[617,748,1354,1380]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="family">Pilumnidae</taxonomicName>
. We currently retain
<taxonomicName id="4C0B53CEFFDFFFF627B7FAA8FBAC1458" authorityName="Ward" authorityYear="1936" box="[983,1113,1355,1380]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF627B7FAA8FBAC1458" box="[983,1113,1355,1380]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
</taxonomicName>
in
<taxonomicName id="4C0B53CEFFDFFFF6201EFAA9FB0B1458" authorityName="Stimpson" authorityYear="1858" box="[1150,1278,1354,1380]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="392" phylum="Arthropoda" rank="subFamily" subFamily="Rhizopinae">Rhizopinae</taxonomicName>
following
<bibRefCitation id="EF9A55BCFFDFFFF6211AFAA9FF1114B4" author="Ng, P. K. L." pageId="1" pageNumber="393" pagination="69 - 111" refId="ref4856" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng (1987)</bibRefCitation>
, Ng
<emphasis id="B97FF45FFFDFFFF6257BFA8CFEA014B4" box="[283,341,1390,1416]" italics="true" pageId="1" pageNumber="393">et al.</emphasis>
(2008) and
<bibRefCitation id="EF9A55BCFFDFFFF62580FA8DFD0314B4" author="Poore, G. C. B. &amp; Ahyong, S. T." box="[480,758,1390,1416]" pageId="1" pageNumber="393" refId="ref5388" refString="Poore, G. C. B. &amp; Ahyong, S. T. (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Clayton South, xii + 916 pp. https: // doi. org / 10.1071 / 9781486311798" type="book" year="2023">Poore &amp; Ahyong (2023)</bibRefCitation>
, because of its close similarity to other rhizopines, many of which lack (or have minimally developed) anterolateral carapace spines and have a male pleon in which somites 1 and 3 are strongly widened and somite 2 narrow. The monophyly of
<taxonomicName id="4C0B53CEFFDFFFF627E0FA55FBF714EC" authorityName="Stimpson" authorityYear="1858" box="[896,1026,1462,1488]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="392" phylum="Arthropoda" rank="subFamily" subFamily="Rhizopinae">Rhizopinae</taxonomicName>
, however, has not been demonstrated (
<bibRefCitation id="EF9A55BCFFDFFFF624FFFA39FEFD14C8" author="Ng, P. K. L." box="[159,264,1498,1524]" pageId="1" pageNumber="393" pagination="69 - 111" refId="ref4856" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987</bibRefCitation>
; Ng
<emphasis id="B97FF45FFFDFFFF62524FA38FE7514C8" box="[324,384,1498,1524]" italics="true" pageId="1" pageNumber="393">et al.</emphasis>
2008) and as noted by
<bibRefCitation id="EF9A55BCFFDFFFF626F9FA39FC4014C8" author="Poore, G. C. B. &amp; Ahyong, S. T." box="[665,949,1498,1524]" pageId="1" pageNumber="393" refId="ref5388" refString="Poore, G. C. B. &amp; Ahyong, S. T. (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Clayton South, xii + 916 pp. https: // doi. org / 10.1071 / 9781486311798" type="book" year="2023">Poore &amp; Ahyong (2023)</bibRefCitation>
,
<taxonomicName id="4C0B53CEFFDFFFF627A4FA39FBB114C8" authorityName="Stimpson" authorityYear="1858" box="[964,1092,1498,1524]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="392" phylum="Arthropoda" rank="subFamily" subFamily="Rhizopinae">Rhizopinae</taxonomicName>
and Pilumninae are currently best considered as heuristic groupings that probably reflect a degree of phylogenetic and ecological reality, but which require refinement to be taxonomically effective. The
<taxonomicName id="4C0B53CEFFDFFFF6273AF9C1FC2F1700" authorityName="Stimpson" authorityYear="1858" box="[858,986,1570,1596]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="392" phylum="Arthropoda" rank="subFamily" subFamily="Rhizopinae">Rhizopinae</taxonomicName>
and Pilumninae remain to be properly diagnosed.
</paragraph>
<paragraph id="8BB4284DFFDFFFF624A7F989FC5516BC" blockId="1.[151,1437,489,2028]" pageId="1" pageNumber="393">
The resemblance between
<taxonomicName id="4C0B53CEFFDFFFF6259FF988FD7417B8" authorityName="Ward" authorityYear="1936" box="[511,641,1643,1668]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF6259FF988FD7417B8" box="[511,641,1643,1668]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B53CEFFDFFFF626A0F988FCE817B8" authorityName="A. Milne-Edwards" authorityYear="1873" box="[704,797,1643,1668]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF626A0F988FCE817B8" box="[704,797,1643,1668]" italics="true" pageId="1" pageNumber="393">Notonyx</emphasis>
</taxonomicName>
is superficial.
<taxonomicName id="4C0B53CEFFDFFFF627B0F988FAA217B8" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[976,1367,1642,1668]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF627B0F988FBD817B8" box="[976,1069,1643,1668]" italics="true" pageId="1" pageNumber="393">Notonyx</emphasis>
A.
<bibRefCitation id="EF9A55BCFFDFFFF6203EF989FAA217B8" author="Milne-Edwards, A." box="[1118,1367,1642,1668]" pageId="1" pageNumber="393" pagination="155 - 332" refId="ref4606" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54B096EFFFDFFFF6210AF989FA6917B8" box="[1386,1436,1642,1668]" pageId="1" pageNumber="393">type</typeStatus>
species
<taxonomicName id="4C0B53CEFFDFFFF62492F96CFD391794" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[242,716,1678,1704]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B97FF45FFFDFFFF62492F96CFE531794" box="[242,422,1678,1704]" italics="true" pageId="1" pageNumber="393">Notonyx nitidus</emphasis>
A.
<bibRefCitation id="EF9A55BCFFDFFFF625B4F96DFD391794" author="Milne-Edwards, A." box="[468,716,1678,1704]" pageId="1" pageNumber="393" pagination="155 - 332" refId="ref4606" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
</taxonomicName>
) is now known to belong to the
<taxonomicName id="4C0B53CEFFDFFFF6202FF96DFB191794" authorityName="MacLeay" authorityYear="1838" box="[1103,1260,1678,1704]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
, its sternal and gonopodal characters all being characteristic for the family (sensu
<bibRefCitation id="EF9A55BCFFDFFFF627E6F951FBE717F0" author="Castro, P." box="[902,1042,1714,1740]" pageId="1" pageNumber="393" pagination="609 - 773" refId="ref3991" refString="Castro, P. (2007) A reappraisal of the family Goneplacidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) and revision of the subfamily Goneplacinae, with the description of ten new genera and eighteen new species. Zoosystema, 29 (4), 609 - 773." type="journal article" year="2007">Castro 2007</bibRefCitation>
;
<bibRefCitation id="EF9A55BCFFDFFFF6207EF951FB1E17F0" author="Castro, P. &amp; Guinot, D. &amp; Ng, P. K. L." box="[1054,1259,1714,1740]" pageId="1" pageNumber="393" pagination="36 - 56" refId="ref4044" refString="Castro, P., Guinot, D. &amp; Ng, P. K. L. (2010) A new family for Sotoplax robertsi Guinot, 1984, with a diagnosis and key to the Goneplacoidea MacLeay, 1838 (Crustacea: Decapoda: Brachyura). Zootaxa, 2356 (1), 36 - 56. https: // doi. org / 10.11646 / zootaxa. 2356.1.2" type="journal article" year="2010">
Castro
<emphasis id="B97FF45FFFDFFFF6200FF950FB5D17F0" box="[1135,1192,1714,1740]" italics="true" pageId="1" pageNumber="393">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EF9A55BCFFDFFFF62096F951FED017CC" author="Ng, P. K. L. &amp; Manuel-Santos, M. R." pageId="1" pageNumber="393" pagination="39 - 68" refId="ref5161" refString="Ng, P. K. L. &amp; Manuel-Santos, M. R. (2007) Establishment of the Vultocinidae, a new family for an unusual new genus and new species of Indo-West Pacific crab (Crustacea: Decapoda: Brachyura: Goneplacoidea), with comments on the taxonomy of the Goneplacidae. Zootaxa, 1558, 39 - 68. https: // doi. org / 10.11646 / zootaxa. 1558.1.3" type="journal article" year="2007">Ng &amp; Manuel-Santos 2007</bibRefCitation>
). The G1 of all 12 known species of
<taxonomicName id="4C0B53CEFFDFFFF626DCF934FCEC17CC" authorityName="A. Milne-Edwards" authorityYear="1873" box="[700,793,1751,1776]" class="Malacostraca" family="Goneplacidae" genus="Notonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF626DCF934FCEC17CC" box="[700,793,1751,1776]" italics="true" pageId="1" pageNumber="393">Notonyx</emphasis>
</taxonomicName>
is relatively short and stout, the distal parts are lined with prominent spines, and the G2 is elongate, being always longer than the G1 (see
<bibRefCitation id="EF9A55BCFFDFFFF62067F919FB3F1629" author="Clark, P. F. &amp; Ng, P. K. L." box="[1031,1226,1786,1813]" pageId="1" pageNumber="393" pagination="539 - 551" refId="ref4117" refString="Clark, P. F. &amp; Ng, P. K. L. (2005) A new species of Notonyx A. Milne-Edwards, 1873 (Crustacea, Brachyura, Goneplacidae) from the intertidal zone of Phuket, Thailand. Zoosystema, 28 (2), 539 - 551." type="journal article" year="2005">Clark &amp; Ng 2005</bibRefCitation>
;
<bibRefCitation id="EF9A55BCFFDFFFF620B5F919FA6D1628" author="Ng, P. K. L. &amp; Clark P. F." box="[1237,1432,1786,1812]" pageId="1" pageNumber="393" pagination="20 - 26" refId="ref4914" refString="Ng, P. K. L. &amp; Clark P. F. (2008) A new species of Notonyx A. Milne-Edwards, 1873 (Crustacea: Decapoda: Brachyura: Goneplacidae) from Indonesia. Zootaxa, 1897 (1), 20 - 26. https: // doi. org / 10.11646 / zootaxa. 1897.1.2" type="journal article" year="2008">Ng &amp; Clark 2008</bibRefCitation>
, 2010, 2011;
<bibRefCitation id="EF9A55BCFFDFFFF62542F8FDFDAD1604" author="Naruse, T. &amp; Maenosono, T." box="[290,600,1822,1848]" pageId="1" pageNumber="393" pagination="183 - 189" refId="ref4660" refString="Naruse, T. &amp; Maenosono, T. (2009) Notonyx kumi, a new species of goneplacid crab (Decapoda: Brachyura) from the Ryukyu Islands, Japan, and Lesser Sunda Islands, Indonesia. Bulletin of the National Museum of Nature and Science, Tokyo, Series A (Zoology), Supplement 3, 183 - 189." type="journal article" year="2009">Naruse &amp; Maenosono 2009</bibRefCitation>
;
<bibRefCitation id="EF9A55BCFFDFFFF62603F8FDFC921604" author="Naruse, T. &amp; Takeda, M." box="[611,871,1822,1848]" pageId="1" pageNumber="393" pagination="71 - 74" refId="ref4725" refString="Naruse, T. &amp; Takeda, M. (2010) Notonyx angulatus, a new species of goneplacid crab (Crustacea, Decapoda, Brachyura) from Amami-Oshima, the Ryukyu Islands, Japan. Bulletin of the National Museum of Nature and Science, Tokyo, (A) (Zoology), 36 (3), 71 - 74." type="journal article" year="2010">Naruse &amp; Takeda 2010</bibRefCitation>
;
<bibRefCitation id="EF9A55BCFFDFFFF62711F8FDFBAC1604" author="Rahayu, D. L. &amp; Ng, P. K. L." box="[881,1113,1822,1848]" pageId="1" pageNumber="393" pagination="51 - 60" refId="ref5478" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2010 a) The species of Notonyx A. Milne-Edwards, 1873, from Lesser Sunda and Maluku Islands, Indonesia, with description of a new species (Crustacea: Decapoda: Brachyura: Goneplacidae). Zootaxa, 2570 (1), 51 - 60. https: // doi. org / 10.11646 / zootaxa. 2570.1.3" type="journal article" year="2010">Rahayu &amp; Ng 2010a</bibRefCitation>
, b;
<bibRefCitation id="EF9A55BCFFDFFFF620E0F8FDFAED1604" author="Rahayu, D. L." box="[1152,1304,1822,1848]" pageId="1" pageNumber="393" pagination="33 - 39" refId="ref5442" refString="Rahayu, D. L. (2011) A new species of Notonyx (Crustacea: Decapoda: Brachyura: Goneplacidae) from Maluku, Indonesia. Zootaxa, 2982, 33 - 39." type="journal article" year="2011">Rahayu 2011</bibRefCitation>
). The males of
<taxonomicName id="4C0B53CEFFDFFFF624D5F8A0FEE41660" authorityName="Miers" authorityYear="1884" box="[181,273,1858,1884]" class="Malacostraca" family="Pilumnidae" genus="Ceratoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B97FF45FFFDFFFF624D5F8A0FEE41660" box="[181,273,1858,1884]" italics="true" pageId="1" pageNumber="393">P. laevis</emphasis>
</taxonomicName>
examined in this study show conclusively that
<taxonomicName id="4C0B53CEFFDFFFF62742F8A0FC511660" authorityName="Ward" authorityYear="1936" box="[802,932,1859,1884]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF62742F8A0FC511660" box="[802,932,1859,1884]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C0B53CEFFDFFFF6202DF8A1FB241660" authorityName="Samouelle" authorityYear="1819" box="[1101,1233,1858,1884]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="family">Pilumnidae</taxonomicName>
: the G1 is sinuous and slender, and the G2 is very short and sigmoid in form (
<figureCitation id="133034C8FFDFFFF6274BF885FC6716BC" box="[811,914,1894,1920]" captionStart="FIGURE 4" captionStartId="6.[151,250,1719,1743]" captionTargetBox="[153,1434,599,1693]" captionTargetId="figure-203@6.[151,1436,597,1695]" captionTargetPageId="6" captionText="FIGURE 4. Pronotonyx laevis (Miers, 1884): A–B, female (cl 5.5 mm, cw 7.3 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; C–D, female (cl 3.8 mm, cw 5.0 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; E, female holotype (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7; F–J, male (cl 10.3, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, cephalothorax, anterior view; B–C, carapace, right anterolateral portion, dorsal view; D, front, dorsal view; E, left maxilliped 3, anterior view; F, median buccal margin; G, anterior thoracic sternum; H, pleon; I, left G1, pleonal view; J, left G2, pleonal view. Scale: A, I, J = 1.0 mm; B–D, F–H = 2.0; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7960060" httpUri="https://zenodo.org/record/7960060/files/figure.png" pageId="1" pageNumber="393">Fig. 4I, J</figureCitation>
).
</paragraph>
<paragraph id="8BB4284DFFDFFFF524A7F869FC351229" blockId="1.[151,1437,489,2028]" lastBlockId="2.[151,1437,150,933]" lastPageId="2" lastPageNumber="394" pageId="1" pageNumber="393">
Of the known rhizopine genera (sensu
<bibRefCitation id="EF9A55BCFFDFFFF6261FF869FD101698" author="Ng, P. K. L." box="[639,741,1930,1956]" pageId="1" pageNumber="393" pagination="69 - 111" refId="ref4856" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987</bibRefCitation>
; Ng
<emphasis id="B97FF45FFFDFFFF6277CF868FCA01698" box="[796,853,1930,1956]" italics="true" pageId="1" pageNumber="393">et al.</emphasis>
2008;
<bibRefCitation id="EF9A55BCFFDFFFF627C5F869FB5C1698" author="Poore, G. C. B. &amp; Ahyong, S. T." box="[933,1193,1930,1956]" pageId="1" pageNumber="393" refId="ref5388" refString="Poore, G. C. B. &amp; Ahyong, S. T. (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Clayton South, xii + 916 pp. https: // doi. org / 10.1071 / 9781486311798" type="book" year="2023">Poore &amp; Ahyong 2023</bibRefCitation>
),
<taxonomicName id="4C0B53CEFFDFFFF620DDF868FACA1698" authorityName="Ward" authorityYear="1936" box="[1213,1343,1931,1956]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF620DDF868FACA1698" box="[1213,1343,1931,1956]" italics="true" pageId="1" pageNumber="393">Pronotonyx</emphasis>
</taxonomicName>
appears to be morphologically closest to
<taxonomicName id="4C0B53CEFFDFFFF62667F84DFC7816F4" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" box="[519,909,1966,1992]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF62667F84DFCF516F4" box="[519,768,1966,1992]" italics="true" pageId="1" pageNumber="393">Pseudocryptocoeloma</emphasis>
<bibRefCitation id="EF9A55BCFFDFFFF62767F84DFC7816F4" author="Ward, M." box="[775,909,1966,1992]" pageId="1" pageNumber="393" pagination="1 - 13" refId="ref5805" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936</bibRefCitation>
</taxonomicName>
, which was established for one new species,
<taxonomicName id="4C0B53CEFFDFFFF621E8F84CFE9E16D0" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B97FF45FFFDFFFF621E8F84CFF1616D0" italics="true" pageId="1" pageNumber="393">P. parvus</emphasis>
<bibRefCitation id="EF9A55BCFFDFFFF62488F831FE9E16D0" author="Ward, M." box="[232,363,2002,2028]" pageId="1" pageNumber="393" pagination="1 - 13" refId="ref5805" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936</bibRefCitation>
</taxonomicName>
, also from Queensland.
<taxonomicName id="4C0B53CEFFDFFFF62611F831FC9F16D0" authorityName="Ward" authorityYear="1936" box="[625,874,2002,2028]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDFFFF62611F831FC9F16D0" box="[625,874,2002,2028]" italics="true" pageId="1" pageNumber="393">Pseudocryptocoeloma</emphasis>
</taxonomicName>
was a poorly known genus, the original description being brief and the figures uninformative.
<bibRefCitation id="EF9A55BCFFDCFFF52618FF74FCA5118D" author="Edmondson, C. H." box="[632,848,150,177]" pageId="2" pageNumber="394" pagination="183 - 243" refId="ref4422" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson (1951)</bibRefCitation>
interpreted the genus as including a new species he recognised from
<collectingCountry id="F31C68DDFFDCFFF5253BFF58FE5F11E9" box="[347,426,187,213]" name="Samoa" pageId="2" pageNumber="394">Samoa</collectingCountry>
,
<taxonomicName id="4C0B53CEFFDCFFF525D8FF58FD7511E9" authorityName="Edmondson" authorityYear="1951" box="[440,640,187,213]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="symmetrinudus">
<emphasis id="B97FF45FFFDCFFF525D8FF58FD7511E9" box="[440,640,187,213]" italics="true" pageId="2" pageNumber="394">P. symmetrinudus</emphasis>
</taxonomicName>
, a taxon with distinct areolae on the carapace.
<bibRefCitation id="EF9A55BCFFDCFFF520C6FF58FA6911E9" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[1190,1436,187,213]" pageId="2" pageNumber="394" pagination="428 - 454" refId="ref5307" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5), 428 - 454. https: // doi. org / 10.11646 / zootaxa. 5244.5.2" type="journal article" year="2023">Ng &amp; Rahayu (2023)</bibRefCitation>
redescribed the genus and species, restricting
<taxonomicName id="4C0B53CEFFDCFFF526D2FF3CFC5E11C5" authorityName="Ward" authorityYear="1936" box="[690,939,223,249]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF526D2FF3CFC5E11C5" box="[690,939,223,249]" italics="true" pageId="2" pageNumber="394">Pseudocryptocoeloma</emphasis>
</taxonomicName>
to the
<typeStatus id="54B096EFFFDCFFF52065FF3CFBC011C5" box="[1029,1077,223,249]" pageId="2" pageNumber="394">type</typeStatus>
species,
<taxonomicName id="4C0B53CEFFDCFFF520C3FF3CFE801021" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="parvum">
<emphasis id="B97FF45FFFDCFFF520C3FF3CFF181020" italics="true" pageId="2" pageNumber="394">Pseudocryptocoeloma parvum</emphasis>
<bibRefCitation id="EF9A55BCFFDCFFF52492FEE0FE801021" author="Ward, M." box="[242,373,259,285]" pageId="2" pageNumber="394" pagination="1 - 13" refId="ref5805" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936</bibRefCitation>
</taxonomicName>
, and transferring
<bibRefCitation id="EF9A55BCFFDCFFF52655FEE0FCE81021" author="Edmondson, C. H." box="[565,797,258,285]" pageId="2" pageNumber="394" pagination="183 - 243" refId="ref4422" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson’s (1951)</bibRefCitation>
species to
<taxonomicName id="4C0B53CEFFDCFFF527F5FEE0FB6F1021" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[917,1178,259,285]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF527F5FEE0FBF81021" box="[917,1037,259,285]" italics="true" pageId="2" pageNumber="394">Lophoplax</emphasis>
<bibRefCitation id="EF9A55BCFFDCFFF52071FEE0FB6F1021" author="Tesch, J. J." box="[1041,1178,259,285]" pageId="2" pageNumber="394" pagination="149 - 295" refId="ref5770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C0B53CEFFDCFFF520C3FEE0FA691021" authorityName="Ward" authorityYear="1936" box="[1187,1436,259,285]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF520C3FEE0FA691021" box="[1187,1436,259,285]" italics="true" pageId="2" pageNumber="394">Pseudocryptocoeloma</emphasis>
</taxonomicName>
can easily be distinguished from
<taxonomicName id="4C0B53CEFFDCFFF5266FFEC4FD64107C" authorityName="Ward" authorityYear="1936" box="[527,657,295,320]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF5266FFEC4FD64107C" box="[527,657,295,320]" italics="true" pageId="2" pageNumber="394">Pronotonyx</emphasis>
</taxonomicName>
in possessing a more quadrate carapace with the regions weakly but visibly demarcated (
<bibRefCitation id="EF9A55BCFFDCFFF52519FEA8FDA21059" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[377,599,331,357]" pageId="2" pageNumber="394" pagination="428 - 454" refId="ref5307" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5), 428 - 454. https: // doi. org / 10.11646 / zootaxa. 5244.5.2" type="journal article" year="2023">Ng &amp; Rahayu 2023</bibRefCitation>
: fig. 13A) (versus proportionately wider with the no regions visible and the surface appearing smooth and polished; anterolateral teeth are low but indicated by distinct notches versus entire or scarcely discernible;
<figureCitation id="133034C8FFDCFFF525E3FE70FE2F1091" box="[387,474,403,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,880,904]" captionTargetBox="[151,1436,181,856]" captionTargetId="figure-18@3.[151,1436,181,856]" captionTargetPageId="3" captionText="FIGURE 1. Pronotonyx laevis (Miers, 1884), holotype female (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7. A, overall dorsal habitus; B, frontal view of cephalothorax; C, ventral view of cephalothorax; D, right chela (outer view)." figureDoi="http://doi.org/10.5281/zenodo.7960050" httpUri="https://zenodo.org/record/7960050/files/figure.png" pageId="2" pageNumber="394">Fig. 1A</figureCitation>
,
<figureCitation id="133034C8FFDCFFF52586FE70FDDE1091" box="[486,555,403,429]" captionStart="FIGURE 2" captionStartId="4.[151,250,1908,1932]" captionTargetBox="[168,1420,181,1884]" captionTargetId="figure-17@4.[168,1420,181,1884]" captionTargetPageId="4" captionText="FIGURE 2. Pronotonyx laevis (Miers, 1884), male (cl 10.3 mm, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, overall dorsal habitus; B, dorsal view of carapace; C, left maxilliped 3; D, frontal view of cephalothorax; E, frontal view showing epistome, eyes, antennules, antenna and orbit; F, outer view of chelae; G, buccal cavity, anterior thoracic sternites and pleon; H, anterior thoracic sternites and pleon; I, posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.7960054" httpUri="https://zenodo.org/record/7960054/files/figure.png" pageId="2" pageNumber="394">2A, B</figureCitation>
,
<figureCitation id="133034C8FFDCFFF52656FE70FD881091" box="[566,637,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1640,1664]" captionTargetBox="[151,1436,181,1616]" captionTargetId="figure-18@5.[151,1436,181,1616]" captionTargetPageId="5" captionText="FIGURE 3. Pronotonyx laevis (Miers, 1884), Lindeman Island, Queensland: A, female (cl 3.8 mm, cw 5.0 mm), NHM 1937.7.15.18; B, female (cl 5.5 mm, cw 7.3 mm), NHM 1937.7.15.19; C–H, female (cl 9.5 mm, cw 13.2 mm), AM P93048. A–C, overall dorsal habitus; D, dorsal view of carapace; E, outer view of chelae; F, buccal cavity, anterior thoracic sternites and pleon; G, posterior thoracic sternites and pleon; H, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7960056" httpUri="https://zenodo.org/record/7960056/files/figure.png" pageId="2" pageNumber="394">3A–D</figureCitation>
,
<figureCitation id="133034C8FFDCFFF526E8FE70FD391091" box="[648,716,403,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1719,1743]" captionTargetBox="[153,1434,599,1693]" captionTargetId="figure-203@6.[151,1436,597,1695]" captionTargetPageId="6" captionText="FIGURE 4. Pronotonyx laevis (Miers, 1884): A–B, female (cl 5.5 mm, cw 7.3 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; C–D, female (cl 3.8 mm, cw 5.0 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; E, female holotype (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7; F–J, male (cl 10.3, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, cephalothorax, anterior view; B–C, carapace, right anterolateral portion, dorsal view; D, front, dorsal view; E, left maxilliped 3, anterior view; F, median buccal margin; G, anterior thoracic sternum; H, pleon; I, left G1, pleonal view; J, left G2, pleonal view. Scale: A, I, J = 1.0 mm; B–D, F–H = 2.0; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7960060" httpUri="https://zenodo.org/record/7960060/files/figure.png" pageId="2" pageNumber="394">4B, C</figureCitation>
); the posterior margin of the epistome is almost straight with the median lobe low and the lateral parts almost straight (
<bibRefCitation id="EF9A55BCFFDCFFF5268BFE54FC3F10ED" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[747,970,439,465]" pageId="2" pageNumber="394" pagination="428 - 454" refId="ref5307" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5), 428 - 454. https: // doi. org / 10.11646 / zootaxa. 5244.5.2" type="journal article" year="2023">Ng &amp; Rahayu 2023</bibRefCitation>
: figs. 14A, 15C) (versus margin distinctly sinuous with the lateral parts distinctly convex;
<figureCitation id="133034C8FFDCFFF526D8FE38FCFA10C9" box="[696,783,475,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,880,904]" captionTargetBox="[151,1436,181,856]" captionTargetId="figure-18@3.[151,1436,181,856]" captionTargetPageId="3" captionText="FIGURE 1. Pronotonyx laevis (Miers, 1884), holotype female (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7. A, overall dorsal habitus; B, frontal view of cephalothorax; C, ventral view of cephalothorax; D, right chela (outer view)." figureDoi="http://doi.org/10.5281/zenodo.7960050" httpUri="https://zenodo.org/record/7960050/files/figure.png" pageId="2" pageNumber="394">Fig. 1B</figureCitation>
,
<figureCitation id="133034C8FFDCFFF5277CFE38FCB410C9" box="[796,833,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1908,1932]" captionTargetBox="[168,1420,181,1884]" captionTargetId="figure-17@4.[168,1420,181,1884]" captionTargetPageId="4" captionText="FIGURE 2. Pronotonyx laevis (Miers, 1884), male (cl 10.3 mm, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, overall dorsal habitus; B, dorsal view of carapace; C, left maxilliped 3; D, frontal view of cephalothorax; E, frontal view showing epistome, eyes, antennules, antenna and orbit; F, outer view of chelae; G, buccal cavity, anterior thoracic sternites and pleon; H, anterior thoracic sternites and pleon; I, posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.7960054" httpUri="https://zenodo.org/record/7960054/files/figure.png" pageId="2" pageNumber="394">2D</figureCitation>
,
<figureCitation id="133034C8FFDCFFF5272EFE38FC9A10C9" box="[846,879,475,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1640,1664]" captionTargetBox="[151,1436,181,1616]" captionTargetId="figure-18@5.[151,1436,181,1616]" captionTargetPageId="5" captionText="FIGURE 3. Pronotonyx laevis (Miers, 1884), Lindeman Island, Queensland: A, female (cl 3.8 mm, cw 5.0 mm), NHM 1937.7.15.18; B, female (cl 5.5 mm, cw 7.3 mm), NHM 1937.7.15.19; C–H, female (cl 9.5 mm, cw 13.2 mm), AM P93048. A–C, overall dorsal habitus; D, dorsal view of carapace; E, outer view of chelae; F, buccal cavity, anterior thoracic sternites and pleon; G, posterior thoracic sternites and pleon; H, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7960056" httpUri="https://zenodo.org/record/7960056/files/figure.png" pageId="2" pageNumber="394">3E</figureCitation>
,
<figureCitation id="133034C8FFDCFFF5271CFE38FC3410C9" box="[892,961,475,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1719,1743]" captionTargetBox="[153,1434,599,1693]" captionTargetId="figure-203@6.[151,1436,597,1695]" captionTargetPageId="6" captionText="FIGURE 4. Pronotonyx laevis (Miers, 1884): A–B, female (cl 5.5 mm, cw 7.3 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; C–D, female (cl 3.8 mm, cw 5.0 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; E, female holotype (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7; F–J, male (cl 10.3, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, cephalothorax, anterior view; B–C, carapace, right anterolateral portion, dorsal view; D, front, dorsal view; E, left maxilliped 3, anterior view; F, median buccal margin; G, anterior thoracic sternum; H, pleon; I, left G1, pleonal view; J, left G2, pleonal view. Scale: A, I, J = 1.0 mm; B–D, F–H = 2.0; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7960060" httpUri="https://zenodo.org/record/7960060/files/figure.png" pageId="2" pageNumber="394">4A, F</figureCitation>
); the chelae are distinctly stouter with the pollex clearly shorter than the palm with the dactylus angled obliquely downwards at an angle of about 60° to the vertical (
<bibRefCitation id="EF9A55BCFFDCFFF5249AFDC0FE291301" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[250,476,547,573]" pageId="2" pageNumber="394" pagination="428 - 454" refId="ref5307" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5), 428 - 454. https: // doi. org / 10.11646 / zootaxa. 5244.5.2" type="journal article" year="2023">Ng &amp; Rahayu 2023</bibRefCitation>
: figs. 13B, 14C, D) (versus chelae longer, with the pollex as long as the palm and the dactylus inclined at an angle not exceeding 45° to the vertical;
<figureCitation id="133034C8FFDCFFF5273FFDA4FC42135D" box="[863,951,583,609]" captionStart="FIGURE 1" captionStartId="3.[151,250,880,904]" captionTargetBox="[151,1436,181,856]" captionTargetId="figure-18@3.[151,1436,181,856]" captionTargetPageId="3" captionText="FIGURE 1. Pronotonyx laevis (Miers, 1884), holotype female (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7. A, overall dorsal habitus; B, frontal view of cephalothorax; C, ventral view of cephalothorax; D, right chela (outer view)." figureDoi="http://doi.org/10.5281/zenodo.7960050" httpUri="https://zenodo.org/record/7960050/files/figure.png" pageId="2" pageNumber="394">Fig. 1D</figureCitation>
,
<figureCitation id="133034C8FFDCFFF527A4FDA4FC17135D" box="[964,994,583,609]" captionStart="FIGURE 2" captionStartId="4.[151,250,1908,1932]" captionTargetBox="[168,1420,181,1884]" captionTargetId="figure-17@4.[168,1420,181,1884]" captionTargetPageId="4" captionText="FIGURE 2. Pronotonyx laevis (Miers, 1884), male (cl 10.3 mm, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, overall dorsal habitus; B, dorsal view of carapace; C, left maxilliped 3; D, frontal view of cephalothorax; E, frontal view showing epistome, eyes, antennules, antenna and orbit; F, outer view of chelae; G, buccal cavity, anterior thoracic sternites and pleon; H, anterior thoracic sternites and pleon; I, posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.7960054" httpUri="https://zenodo.org/record/7960054/files/figure.png" pageId="2" pageNumber="394">2F</figureCitation>
,
<figureCitation id="133034C8FFDCFFF5278EFDA4FBE4135D" box="[1006,1041,583,609]" captionStart="FIGURE 3" captionStartId="5.[151,250,1640,1664]" captionTargetBox="[151,1436,181,1616]" captionTargetId="figure-18@5.[151,1436,181,1616]" captionTargetPageId="5" captionText="FIGURE 3. Pronotonyx laevis (Miers, 1884), Lindeman Island, Queensland: A, female (cl 3.8 mm, cw 5.0 mm), NHM 1937.7.15.18; B, female (cl 5.5 mm, cw 7.3 mm), NHM 1937.7.15.19; C–H, female (cl 9.5 mm, cw 13.2 mm), AM P93048. A–C, overall dorsal habitus; D, dorsal view of carapace; E, outer view of chelae; F, buccal cavity, anterior thoracic sternites and pleon; G, posterior thoracic sternites and pleon; H, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7960056" httpUri="https://zenodo.org/record/7960056/files/figure.png" pageId="2" pageNumber="394">3E</figureCitation>
); the P2–P5 are clearly shorter and stouter (
<bibRefCitation id="EF9A55BCFFDCFFF52492FD88FE2613B9" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[242,467,619,645]" pageId="2" pageNumber="394" pagination="428 - 454" refId="ref5307" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5), 428 - 454. https: // doi. org / 10.11646 / zootaxa. 5244.5.2" type="journal article" year="2023">Ng &amp; Rahayu 2023</bibRefCitation>
: fig. 13A, 14E) (versus P2–P5 distinctly longer and more slender;
<figureCitation id="133034C8FFDCFFF520DEFD88FAC213B9" box="[1214,1335,619,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,880,904]" captionTargetBox="[151,1436,181,856]" captionTargetId="figure-18@3.[151,1436,181,856]" captionTargetPageId="3" captionText="FIGURE 1. Pronotonyx laevis (Miers, 1884), holotype female (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7. A, overall dorsal habitus; B, frontal view of cephalothorax; C, ventral view of cephalothorax; D, right chela (outer view)." figureDoi="http://doi.org/10.5281/zenodo.7960050" httpUri="https://zenodo.org/record/7960050/files/figure.png" pageId="2" pageNumber="394">Fig. 1A, B</figureCitation>
,
<figureCitation id="133034C8FFDCFFF52123FD88FA9213B9" box="[1347,1383,619,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1908,1932]" captionTargetBox="[168,1420,181,1884]" captionTargetId="figure-17@4.[168,1420,181,1884]" captionTargetPageId="4" captionText="FIGURE 2. Pronotonyx laevis (Miers, 1884), male (cl 10.3 mm, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, overall dorsal habitus; B, dorsal view of carapace; C, left maxilliped 3; D, frontal view of cephalothorax; E, frontal view showing epistome, eyes, antennules, antenna and orbit; F, outer view of chelae; G, buccal cavity, anterior thoracic sternites and pleon; H, anterior thoracic sternites and pleon; I, posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.7960054" httpUri="https://zenodo.org/record/7960054/files/figure.png" pageId="2" pageNumber="394">2A</figureCitation>
,
<figureCitation id="133034C8FFDCFFF52113FD88FF591395" captionStart="FIGURE 3" captionStartId="5.[151,250,1640,1664]" captionTargetBox="[151,1436,181,1616]" captionTargetId="figure-18@5.[151,1436,181,1616]" captionTargetPageId="5" captionText="FIGURE 3. Pronotonyx laevis (Miers, 1884), Lindeman Island, Queensland: A, female (cl 3.8 mm, cw 5.0 mm), NHM 1937.7.15.18; B, female (cl 5.5 mm, cw 7.3 mm), NHM 1937.7.15.19; C–H, female (cl 9.5 mm, cw 13.2 mm), AM P93048. A–C, overall dorsal habitus; D, dorsal view of carapace; E, outer view of chelae; F, buccal cavity, anterior thoracic sternites and pleon; G, posterior thoracic sternites and pleon; H, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7960056" httpUri="https://zenodo.org/record/7960056/files/figure.png" pageId="2" pageNumber="394">3A, B</figureCitation>
); and the male pleonal somites 1 and 3 are proportionally narrower (
<bibRefCitation id="EF9A55BCFFDCFFF527CFFD6CFB651395" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[943,1168,655,681]" pageId="2" pageNumber="394" pagination="428 - 454" refId="ref5307" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5), 428 - 454. https: // doi. org / 10.11646 / zootaxa. 5244.5.2" type="journal article" year="2023">Ng &amp; Rahayu 2023</bibRefCitation>
: figs. 13B, 14F, 15E, F) (versus male somites distinctly wider;
<figureCitation id="133034C8FFDCFFF5265FFD50FD7F13F1" box="[575,650,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1908,1932]" captionTargetBox="[168,1420,181,1884]" captionTargetId="figure-17@4.[168,1420,181,1884]" captionTargetPageId="4" captionText="FIGURE 2. Pronotonyx laevis (Miers, 1884), male (cl 10.3 mm, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, overall dorsal habitus; B, dorsal view of carapace; C, left maxilliped 3; D, frontal view of cephalothorax; E, frontal view showing epistome, eyes, antennules, antenna and orbit; F, outer view of chelae; G, buccal cavity, anterior thoracic sternites and pleon; H, anterior thoracic sternites and pleon; I, posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.7960054" httpUri="https://zenodo.org/record/7960054/files/figure.png" pageId="2" pageNumber="394">Fig. 2I</figureCitation>
,
<figureCitation id="133034C8FFDCFFF526F4FD50FD4E13F1" box="[660,699,691,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1719,1743]" captionTargetBox="[153,1434,599,1693]" captionTargetId="figure-203@6.[151,1436,597,1695]" captionTargetPageId="6" captionText="FIGURE 4. Pronotonyx laevis (Miers, 1884): A–B, female (cl 5.5 mm, cw 7.3 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; C–D, female (cl 3.8 mm, cw 5.0 mm), Lindeman Island, Queensland, NHM 1937.7.15.18–19; E, female holotype (cl 5.0 mm, cw 6.5 mm), Arafura Sea, NHM 1882.7; F–J, male (cl 10.3, cw 13.5 mm), Lindeman Island, Queensland, AM P93048. A, cephalothorax, anterior view; B–C, carapace, right anterolateral portion, dorsal view; D, front, dorsal view; E, left maxilliped 3, anterior view; F, median buccal margin; G, anterior thoracic sternum; H, pleon; I, left G1, pleonal view; J, left G2, pleonal view. Scale: A, I, J = 1.0 mm; B–D, F–H = 2.0; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7960060" httpUri="https://zenodo.org/record/7960060/files/figure.png" pageId="2" pageNumber="394">4H</figureCitation>
). In addition, the carapace and pereopods of
<taxonomicName id="4C0B53CEFFDCFFF520C3FD50FA6913F1" authorityName="Ward" authorityYear="1936" box="[1187,1436,691,717]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF520C3FD50FA6913F1" box="[1187,1436,691,717]" italics="true" pageId="2" pageNumber="394">Pseudocryptocoeloma</emphasis>
</taxonomicName>
are distinctly more setose than those of
<taxonomicName id="4C0B53CEFFDCFFF52636FD34FD2D13CC" authorityName="Ward" authorityYear="1936" box="[598,728,727,752]" class="Malacostraca" family="Pilumnidae" genus="Pronotonyx" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF52636FD34FD2D13CC" box="[598,728,727,752]" italics="true" pageId="2" pageNumber="394">Pronotonyx</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9A55BCFFDCFFF52687FD34FC3D13CD" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[743,968,727,753]" pageId="2" pageNumber="394" pagination="428 - 454" refId="ref5307" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5), 428 - 454. https: // doi. org / 10.11646 / zootaxa. 5244.5.2" type="journal article" year="2023">Ng &amp; Rahayu 2023</bibRefCitation>
: fig. 13 versus
<figureCitation id="133034C8FFDCFFF52013FD34FB1813CD" box="[1139,1261,727,753]" captionStart="FIGURE 3" captionStartId="5.[151,250,1640,1664]" captionTargetBox="[151,1436,181,1616]" captionTargetId="figure-18@5.[151,1436,181,1616]" captionTargetPageId="5" captionText="FIGURE 3. Pronotonyx laevis (Miers, 1884), Lindeman Island, Queensland: A, female (cl 3.8 mm, cw 5.0 mm), NHM 1937.7.15.18; B, female (cl 5.5 mm, cw 7.3 mm), NHM 1937.7.15.19; C–H, female (cl 9.5 mm, cw 13.2 mm), AM P93048. A–C, overall dorsal habitus; D, dorsal view of carapace; E, outer view of chelae; F, buccal cavity, anterior thoracic sternites and pleon; G, posterior thoracic sternites and pleon; H, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.7960056" httpUri="https://zenodo.org/record/7960056/files/figure.png" pageId="2" pageNumber="394">Fig. 3C, D</figureCitation>
), with the setae longer and coarser as well as covering the anterior parts of the carapace.
</paragraph>
<paragraph id="8BB4284DFFDCFFF524AFFCFCFA801299" blockId="2.[151,1437,150,933]" pageId="2" pageNumber="394">
The very wide male pleonal somites 1 and 3 is a feature also shared with rhizopine genera such as
<taxonomicName id="4C0B53CEFFDCFFF52120FCFCFEBC1261" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" class="Malacostraca" family="Pilumnidae" genus="Rhizopa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF52120FCFCFA691205" box="[1344,1436,799,825]" italics="true" pageId="2" pageNumber="394">Rhizopa</emphasis>
<bibRefCitation id="EF9A55BCFFDCFFF524F7FCA0FEBC1261" author="Stimpson, W." box="[151,329,834,861]" pageId="2" pageNumber="394" pagination="93 - 110" refId="ref5698" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 [39 - 56]. https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0B53CEFFDCFFF52533FCA0FDD01261" authority="Rathbun, 1931" authorityName="Rathbun" authorityYear="1931" box="[339,549,835,861]" class="Malacostraca" family="Pilumnidae" genus="Ser" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF52533FCA0FE8D1261" box="[339,376,835,861]" italics="true" pageId="2" pageNumber="394">Ser</emphasis>
<bibRefCitation id="EF9A55BCFFDCFFF5251FFCA0FDD01261" author="Rathbun, M. J." box="[383,549,835,861]" pageId="2" pageNumber="394" pagination="75 - 125" refId="ref5664" refString="Rathbun, M. J. (1931) New and rare Chinese crabs. Lingnan Science Journal, 1929 (8), 75 - 125, pls. 5 - 15." type="journal article" year="1931">Rathbun, 1931</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0B53CEFFDCFFF52650FCA0FCCA1261" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" box="[560,831,835,861]" class="Malacostraca" family="Pilumnidae" genus="Cryptolutea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF52650FCA0FD431261" box="[560,694,835,861]" italics="true" pageId="2" pageNumber="394">Cryptolutea</emphasis>
<bibRefCitation id="EF9A55BCFFDCFFF526DBFCA0FCCA1261" author="Ward, M." box="[699,831,835,861]" pageId="2" pageNumber="394" pagination="1 - 13" refId="ref5805" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C0B53CEFFDCFFF52718FCA0FB841261" authority="Ng, 1987" authorityName="Ng" authorityYear="1987" box="[888,1137,835,861]" class="Malacostraca" family="Pilumnidae" genus="Rhizopoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF52718FCA0FBF41261" box="[888,1025,835,861]" italics="true" pageId="2" pageNumber="394">Rhizopoides</emphasis>
<bibRefCitation id="EF9A55BCFFDCFFF52067FCA0FB841261" author="Ng, P. K. L." box="[1031,1137,835,861]" pageId="2" pageNumber="394" pagination="69 - 111" refId="ref4856" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng, 1987</bibRefCitation>
</taxonomicName>
, as well as some species of the heterogeneous
<taxonomicName id="4C0B53CEFFDCFFF5250AFC84FD3312BD" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[362,710,870,897]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B97FF45FFFDCFFF5250AFC84FDE812BD" box="[362,541,871,897]" italics="true" pageId="2" pageNumber="394">Heteropilumnus</emphasis>
De
<bibRefCitation id="EF9A55BCFFDCFFF5262CFC84FD3312BD" author="Man, J. G. De" box="[588,710,870,897]" pageId="2" pageNumber="394" pagination="485 - 609" refId="ref4461" refString="Man, J. G. De (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 485 - 609, pls. 12 - 14. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">Man, 1895</bibRefCitation>
</taxonomicName>
(cf.
<bibRefCitation id="EF9A55BCFFDCFFF5269AFC84FC9512BD" author="Ng, P. K. L." box="[762,864,870,897]" pageId="2" pageNumber="394" pagination="627 - 633" refId="ref4790" refString="Ng, P. K. L. (1985) On a new species of pilumnid crab, Rhizopa yangae sp. nov. from Singapore, with notes on the genus Rhizopa Stimpson, 1858 (Decapoda: Brachyura: Pilumnidae). Journal of Natural History, 19, 627 - 633. https: // doi. org / 10.1080 / 00222938500770391" type="journal article" year="1985">Ng 1985</bibRefCitation>
: fig. 2F;
<bibRefCitation id="EF9A55BCFFDCFFF527A5FC84FBDF12BD" author="Ng, P. K. L." box="[965,1066,871,897]" pageId="2" pageNumber="394" pagination="69 - 111" refId="ref4856" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987</bibRefCitation>
: fig. 2A;
<bibRefCitation id="EF9A55BCFFDCFFF520F4FC84FA9512BD" author="Ng, P. K. L. &amp; Davie, P. J. F." box="[1172,1376,871,897]" pageId="2" pageNumber="394" pagination="517 - 524" refId="ref5042" refString="Ng, P. K. L. &amp; Davie, P. J. F. (1991) The Indo-Pacific Pilumnidae VII. Notes on the taxonomy of Heteropilumnus sasekumari (Serene, 1971) and Cryptolutea Ward, 1936 (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 30 (3), 517 - 524." type="journal article" year="1991">Ng &amp; Davie 1991</bibRefCitation>
: figs. 2C, 4H;
<bibRefCitation id="EF9A55BCFFDCFFF52497FC68FE201299" author="Ng, P. K. L. &amp; Rahayu, D. L." box="[247,469,907,933]" pageId="2" pageNumber="394" pagination="1 - 100" refId="ref5238" refString="Ng, P. K. L. &amp; Rahayu, D. L. (2020) A synopsis of Typhlocarcinops Rathbun, 1909 (Crustacea: Decapoda: Brachyura: Pilumnidae), with descriptions of nine new species from the Indo-West Pacific. Zootaxa, 4788 (1), 1 - 100. https: // doi. org / 10.11646 / zootaxa. 4788.1.1" type="journal article" year="2020">Ng &amp; Rahayu 2020</bibRefCitation>
: figs. 4B), but these genera have very different carapace and maxilliped 3 features.
</paragraph>
</subSubSection>
</treatment>
</document>
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