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<document id="309A9F1760068FA5D574AE4965342ED6" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7833FFA8FEECFE9E98BFFDB1" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra liv Erwin, new species" docType="treatment" docVersion="12" lastPageNumber="40" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="39" updateTime="1698190273211" updateUser="plazi">
<mods:mods id="D9811B5C0F9015268EC8214560AAE833" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E553615C13A49C36C08EF745C100AD87">
<mods:title id="3265921B4F5401B5F45B7B3673EFEA24">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="34524217CD3833A257071D1403D82B9B" type="personal">
<mods:role id="46C1AE92BB73670C2E664BA7BEFC9F0C">
<mods:roleTerm id="A16AA3354241854E7D5687C6C2494E47">Author</mods:roleTerm>
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<mods:namePart id="B2BBF36FAB054120570637959243B577">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="37BFC31A60F2F67EC7C65C66F3EFE05F">text</mods:typeOfResource>
<mods:relatedItem id="E10DFA348FC4326906C7323166A35C61" type="host">
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<mods:title id="AFEFC8599CD9F256AF2933D8239D4DA2">Zootaxa</mods:title>
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<mods:part id="3ADC5C49DE5A7D86FE0FD4642EFC5D7B">
<mods:date id="5D53FB3BF71A7A20DDB4F01992379909">2002</mods:date>
<mods:detail id="50B619536C53065AA8F27E3F3043F085" type="volume">
<mods:number id="377203C7B3BC5103B4A953BF9196A114">119</mods:number>
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<mods:classification id="A5CF57DF603FBC26BF71D6C99D401E54">journal article</mods:classification>
<mods:identifier id="1D4B3DE418064D141219348CB91A68C1" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="9D81D0F98F15DE6A0F76C05DAB9A2373" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
<mods:identifier id="BE0F20A73AC96B435C4E4E2A76D14DB5" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="9841709310C86FF9F8AC69E1D3EFE9FB" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA7833FFA8FEECFE9E98BFFDB1" ID-DOI="http://doi.org/10.5281/zenodo.6277994" ID-GBIF-Taxon="119330625" ID-Zenodo-Dep="6277994" LSID="urn:lsid:plazi:treatment:03A287CA7833FFA8FEECFE9E98BFFDB1" httpUri="http://treatment.plazi.org/id/03A287CA7833FFA8FEECFE9E98BFFDB1" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<subSubSection id="C31165577833FFA9FEECFE9E9E01FEA6" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8BB436DC7833FFA9FEECFE9E9E69FECA" blockId="38.[264,605,283,345]" box="[264,605,283,310]" pageId="38" pageNumber="39">
<heading id="D0FC81B07833FFA9FEECFE9E9E69FECA" bold="true" box="[264,605,283,310]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="B97FEACE7833FFA9FEECFE9E9E69FECA" bold="true" box="[264,605,283,310]" pageId="38" pageNumber="39">
<taxonomicName id="4C0B4D5F7833FFA9FEECFE9E9DF4FECA" ID-CoL="B2QF" authority="Erwin" authorityName="Erwin" box="[264,448,283,310]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="liv" status="sp. nov.">
<emphasis id="B97FEACE7833FFA9FEECFE9E9D5DFEC9" bold="true" box="[264,361,284,310]" italics="true" pageId="38" pageNumber="39">Agra liv</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B57833FFA9FE29FE999E69FECA" box="[461,605,283,309]" pageId="38" pageNumber="39" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC7833FFA9FEECFEBD9E01FEA6" blockId="38.[264,605,283,345]" box="[264,565,319,345]" pageId="38" pageNumber="39">
(
<figureCitation id="13302A597833FFA9FEF5FEBD9D9EFEA6" box="[273,426,319,345]" captionStart="FIGURE 15. A. liv" captionStartId="38.[264,368,1791,1815]" captionTargetBox="[443,1135,1024,1758]" captionTargetId="figure@38.[442,1144,1004,1765]" captionTargetPageId="38" captionText="FIGURE 15. A. liv, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 2.0; male SW = 3.6, female SW = 3.8; AL = 4.0." httpUri="https://zenodo.org/record/162824/files/figure.png" pageId="38" pageNumber="39">Figs. 15a, 15</figureCitation>
b, 15c, 15d)
</paragraph>
</subSubSection>
<subSubSection id="C31165577833FFA9FEECFE0C982BFE2E" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BB436DC7833FFA9FEECFE0C982BFE2E" blockId="38.[264,1324,398,945]" pageId="38" pageNumber="39">
<emphasis id="B97FEACE7833FFA9FEECFE0C9DB5FE57" bold="true" box="[264,385,398,424]" pageId="38" pageNumber="39">
<typeStatus id="54B0887E7833FFA9FEECFE0C9D48FE57" box="[264,380,398,424]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female,
<emphasis id="B97FEACE7833FFA9FE15FE0C9E85FE57" bold="true" box="[497,689,398,424]" pageId="38" pageNumber="39">
<collectingCountry id="F31C764C7833FFA9FE15FE0C9E98FE57" box="[497,684,398,424]" name="Costa Rica" pageId="38" pageNumber="39">COSTA RICA</collectingCountry>
:
</emphasis>
Puntarenas, Manual Antonio National Park, Quepos,
<quantity id="4CF39B397833FFA9FEECFE359D0BFE2E" box="[264,319,439,465]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="38" pageNumber="39" unit="m" value="80.0">80m</quantity>
,
<geoCoordinate id="EE3F501B7833FFA9FEAFFE359D98FE2E" box="[331,428,439,465]" direction="north" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="9.383333">09°23'N</geoCoordinate>
<geoCoordinate id="EE3F501B7833FFA9FE57FE359E2EFE2E" box="[435,538,439,465]" direction="west" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="-84.15">84°09'W</geoCoordinate>
, January, (Bogenberger)(
<collectionCode id="ED1AAE197833FFA9FCA2FE359FB9FE2E" box="[838,909,439,465]" httpUri="http://grbio.org/cool/wzwk-ye3h" name="Zoologische Sammlung des Bayerischen Staates" pageId="38" pageNumber="39">ZSBS</collectionCode>
: ADP6893).
</paragraph>
</subSubSection>
<subSubSection id="C31165577833FFA9FEDCFE5C986AFDDE" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8BB436DC7833FFA9FEDCFE5C986AFDDE" blockId="38.[264,1324,398,945]" pageId="38" pageNumber="39">
<emphasis id="B97FEACE7833FFA9FEDCFE5C9D84FE07" bold="true" box="[312,432,478,504]" pageId="38" pageNumber="39">Diagnosis</emphasis>
. This is the only species of the
<emphasis id="B97FEACE7833FFA9FCD5FE5D9FB6FE07" box="[817,898,479,504]" italics="true" pageId="38" pageNumber="39">famula</emphasis>
species­group in
<collectingCountry id="F31C764C7833FFA9FBB3FE5D98EEFE06" box="[1111,1242,479,505]" name="Costa Rica" pageId="38" pageNumber="39">Costa Rica</collectingCountry>
that is entirely black with pale tibiae and tarsi and with normal elytral intervals.
</paragraph>
</subSubSection>
<subSubSection id="C31165577833FFA9FEDCFDAC9EC0FC4E" pageId="38" pageNumber="39" type="description">
<paragraph id="8BB436DC7833FFA9FEDCFDAC9EC0FC4E" blockId="38.[264,1324,398,945]" pageId="38" pageNumber="39">
<emphasis id="B97FEACE7833FFA9FEDCFDAC9DFDFDB7" bold="true" box="[312,457,558,584]" pageId="38" pageNumber="39">Description</emphasis>
. Color and luster: Black. Tips of mandibles, sides of antennomeres 5­11, and in some specimens sides of abdominal sterna rufopiceous. Surface shiny. Form: Labrum entire. Head (
<figureCitation id="13302A597833FFA9FDF1FDFD9E4EFD66" box="[533,634,639,665]" captionStart="FIGURE 15. A. liv" captionStartId="38.[264,368,1791,1815]" captionTargetBox="[443,1135,1024,1758]" captionTargetId="figure@38.[442,1144,1004,1765]" captionTargetPageId="38" captionText="FIGURE 15. A. liv, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 2.0; male SW = 3.6, female SW = 3.8; AL = 4.0." httpUri="https://zenodo.org/record/162824/files/figure.png" pageId="38" pageNumber="39">Fig. 15a</figureCitation>
) behind eyes elongate, rounded in both sexes. Prothorax subtubular, short, robust, sutures effaced, widest at middle, dorsal surface moderately convex and sparsely punctate, punctures markedly small. Elytron (
<figureCitation id="13302A597833FFA9FC1BFD4D9861FD16" box="[1023,1109,719,745]" captionStart="FIGURE 15. A. liv" captionStartId="38.[264,368,1791,1815]" captionTargetBox="[443,1135,1024,1758]" captionTargetId="figure@38.[442,1144,1004,1765]" captionTargetPageId="38" captionText="FIGURE 15. A. liv, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 2.0; male SW = 3.6, female SW = 3.8; AL = 4.0." httpUri="https://zenodo.org/record/162824/files/figure.png" pageId="38" pageNumber="39">Fig. 15</figureCitation>
b) with apex with more or less similar acute dentiform projections, interneurs with small regularly spaced and slightly cribriform punctulae, adjacent intervals
<taxonomicName id="4C0B4D5F7833FFA9FC9AFC9D9F97FCC6" box="[894,931,799,825]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
catenate. Sternum VI (
<figureCitation id="13302A597833FFA9FB59FC9D9920FCC6" box="[1213,1300,799,825]" captionStart="FIGURE 15. A. liv" captionStartId="38.[264,368,1791,1815]" captionTargetBox="[443,1135,1024,1758]" captionTargetId="figure@38.[442,1144,1004,1765]" captionTargetPageId="38" captionText="FIGURE 15. A. liv, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 2.0; male SW = 3.6, female SW = 3.8; AL = 4.0." httpUri="https://zenodo.org/record/162824/files/figure.png" pageId="38" pageNumber="39">Fig. 15</figureCitation>
c) moderately deeply v­notched in male, shallowly v­notched in female. Aedeagus (
<figureCitation id="13302A597833FFA9FB5BFCC59926FC9E" box="[1215,1298,839,865]" captionStart="FIGURE 15. A. liv" captionStartId="38.[264,368,1791,1815]" captionTargetBox="[443,1135,1024,1758]" captionTargetId="figure@38.[442,1144,1004,1765]" captionTargetPageId="38" captionText="FIGURE 15. A. liv, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 2.0; male SW = 3.6, female SW = 3.8; AL = 4.0." httpUri="https://zenodo.org/record/162824/files/figure.png" pageId="38" pageNumber="39">Fig. 15</figureCitation>
d) ostium elongate, apex a double scimitar, parameres glabrous. Size: medium, 14.0 to
<quantity id="4CF39B397833FFA9FEECFC159D5DFC4E" box="[264,361,919,945]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="38" pageNumber="39" unit="mm" value="15.5">15.5mm</quantity>
in length,
<quantity id="4CF39B397833FFA9FE03FC159EB7FC4E" box="[487,643,919,945]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.6" metricValueMin="3.4" pageId="38" pageNumber="39" unit="mm" value="4.0" valueMax="4.6" valueMin="3.4">3.4 to 4.6mm</quantity>
in width.
</paragraph>
</subSubSection>
<caption id="DF7466547833FFA9FEECF97D98E2F8A3" httpUri="https://zenodo.org/record/162824/files/figure.png" pageId="38" pageNumber="39" targetBox="[443,1135,1024,1758]" targetPageId="38">
<paragraph id="8BB436DC7833FFA9FEECF97D98E2F8A3" blockId="38.[264,1323,1791,1884]" pageId="38" pageNumber="39">
<emphasis id="B97FEACE7833FFA9FEECF97D9DAEF8E8" bold="true" box="[264,410,1791,1815]" pageId="38" pageNumber="39">FIGURE 15.</emphasis>
<taxonomicName id="4C0B4D5F7833FFA9FE40F8829DE9F8E8" box="[420,477,1792,1815]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="liv">
<emphasis id="B97FEACE7833FFA9FE40F8829DE9F8E8" box="[420,477,1792,1815]" italics="true" pageId="38" pageNumber="39">A. liv</emphasis>
</taxonomicName>
, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 2.0; male SW = 3.6, female SW = 3.8; AL = 4.0.
</paragraph>
</caption>
<subSubSection id="C31165577832FFA8FEDCFE999E5AFE01" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BB436DC7832FFA8FEDCFE999E1DFEA1" blockId="39.[264,1324,283,590]" pageId="39" pageNumber="40">
<emphasis id="B97FEACE7832FFA8FEDCFE999EA1FECA" bold="true" box="[312,661,283,309]" pageId="39" pageNumber="40">Other specimens examined.</emphasis>
<typeStatus id="54B0887E7832FFA8FD7BFE9E9F38FEC9" box="[671,780,284,310]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
above, and a male and female from
<collectingCountry id="F31C764C7832FFA8FB29FE9E991EFEC9" box="[1229,1322,284,310]" name="Panama" pageId="39" pageNumber="40">Panamá</collectingCountry>
are labeled as
<typeStatus id="54B0887E7832FFA8FE4BFEC69E11FEA1" box="[431,549,324,350]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
.
</paragraph>
<paragraph id="8BB436DC7832FFA8FEDCFEE99E5AFE01" blockId="39.[264,1324,283,590]" pageId="39" pageNumber="40">
<emphasis id="B97FEACE7832FFA8FEDCFEE99E35FE7A" bold="true" box="[312,513,363,389]" pageId="39" pageNumber="40">Specific epithet.</emphasis>
The specific epithet,
<taxonomicName id="4C0B4D5F7832FFA8FCEFFEEE9F12FE7A" box="[779,806,364,389]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="liv">
<emphasis id="B97FEACE7832FFA8FCEFFEEE9F12FE7A" box="[779,806,364,389]" italics="true" pageId="39" pageNumber="40">liv</emphasis>
</taxonomicName>
, is a noun in apposition based on the first name of the actress
<taxonomicName id="4C0B4D5F7832FFA8FE0BFE169E2CFE51" box="[495,536,404,430]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="liv">Liv</taxonomicName>
Tyler, starlet of the movie, Armageddon. The existence of this species of elegant beetle is dependent upon the rainforest
<taxonomicName id="4C0B4D5F7832FFA8FC4EFE3E9FFBFE29" box="[938,975,444,470]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
undergoing an Armageddon which is too much to hope for.
</paragraph>
</subSubSection>
<subSubSection id="C31165577832FFA8FEDCFD8998BFFDB1" pageId="39" pageNumber="40" type="description">
<paragraph id="8BB436DC7832FFA8FEDCFD8998BFFDB1" blockId="39.[264,1324,283,590]" pageId="39" pageNumber="40">
<emphasis id="B97FEACE7832FFA8FEDCFD899D4AFDDA" bold="true" box="[312,382,523,549]" pageId="39" pageNumber="40">Notes</emphasis>
. The single Costa Rican record above is from the Pacifico Central Conservation Area. Famula­group stylomere 2 can be found at the
<taxonomicName id="4C0B4D5F7832FFA8FC90FDB69838FDB2" box="[884,1036,564,589]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fortuna">
<emphasis id="B97FEACE7832FFA8FC90FDB69838FDB2" box="[884,1036,564,589]" italics="true" pageId="39" pageNumber="40">Agra fortuna</emphasis>
</taxonomicName>
web page.
</paragraph>
</subSubSection>
</treatment>
</document>
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