<documentid="A4E71D88D4A3743D5D29460AE8BA9444"ID-CLB-Dataset="299696"ID-DOI="10.5281/zenodo.11512952"ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c"ID-ISSN="0068-547X"ID-Zenodo-Dep="11512952"IM.bibliography_approvedBy="julia"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="julia"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="julia"IM.treatmentCitations_approvedBy="julia"IM.treatments_approvedBy="julia"checkinTime="1717721037103"checkinUser="felipe"docAuthor="Kavanaugh, David H. & Cueva-Dabkoski, Mollie"docDate="2023"docId="1C1187FCEB7CED53FF9EFB9162F083CC"docLanguage="en"docName="ProcCalifAcadSci.67.21.493-537.pdf"docOrigin="Proceedings of the California Academy of Sciences 67 (21)"docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article"docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B"docStyleName="ProcCalifAcadSci.2015-.journal_article"docStyleVersion="2"docTitle="Cyclosomus pallidus Kavanaugh & Cueva-Dabkoski 2023, sp. nov."docType="treatment"docVersion="5"lastPageNumber="515"masterDocId="E028FF84EB68ED45FFE5FFAF6327867B"masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia."masterLastPageNumber="537"masterPageNumber="493"pageNumber="513"updateTime="1720721490288"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="873E3769127BC6253730654ECD3F253B">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
<mods:namePartid="A6F07B712E3C79930A9EE240E7591581">Kavanaugh, David H.</mods:namePart>
<mods:affiliationid="67FEF3C793DDA690A39F907D46015280">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
<mods:affiliationid="D1CB55058CCC60956B3D9B01782307F4">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
<taxonomicNameid="53B84D69EB7CED51FF9EFB9161EB822D"authority="Kavanaugh & Cueva-Dabkoski"authorityName="Kavanaugh & Cueva-Dabkoski"authorityYear="2023"box="[123,716,1086,1110]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="20"pageNumber="513"phylum="Arthropoda"rank="species"species="pallidus"status="sp. nov.">
<figureCitationid="0C832A6FEB7CED51FF9EFBF163F0820F"box="[123,215,1118,1140]"captionStart="FIGURE"captionStartId="21.[171,183,1155,1174]"captionTargetBox="[120,1176,216,1138]"captionTargetId="figure-233@21.[120,1176,216,1147]"captionTargetPageId="21"captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512976"httpUri="https://zenodo.org/record/11512976/files/figure.png"pageId="20"pageNumber="513">
<figureCitationid="0C832A6FEB7CED51FF07FBF1622D820F"box="[226,266,1118,1140]"captionStart="FIGURE"captionStartId="33.[171,183,1573,1592]"captionTargetBox="[139,1158,216,1560]"captionTargetId="figure-14@33.[135,1162,216,1560]"captionTargetPageId="33"captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512988"httpUri="https://zenodo.org/record/11512988/files/figure.png"pageId="20"pageNumber="513">14E</figureCitation>
</geoCoordinate>
</figureCitation>
,
<figureCitationid="0C832A6FEB7CED51FEF0FBF1620A820F"box="[277,301,1118,1140]"captionStart="FIGURE"captionStartId="39.[171,183,1593,1612]"captionTargetBox="[168,1128,979,1582]"captionTargetId="figure-84@39.[168,1128,979,1582]"captionTargetPageId="39"captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km."figureDoi="http://doi.org/10.5281/zenodo.11513002"httpUri="https://zenodo.org/record/11513002/files/figure.png"pageId="20"pageNumber="513">21</figureCitation>
<materialsCitationid="24D03CB7EB7CED51FEB0FB2C639D8280"ID-GBIF-Occurrence="4905152332"collectionCode="NHMUK"collectorName=" D. H. Kavanaugh & M. Cueva-Dabkoski"determinerName="H. E. Andrewes"pageId="20"pageNumber="513"specimenCount="2"typeStatus="holotype">
<collectionCodeid="F2A9AE2FEB7CED51FD38FB2B606282E0"box="[733,837,1156,1179]"country="United Kingdom"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665"lsid="urn:lsid:biocol.org:col:34665"name="Natural History Museum, London"pageId="20"pageNumber="513"type="Museum">NHMUK</collectionCode>
, labeled: “
<collectorNameid="394D533CEB7CED51FC56FB2C673A82E0"box="[947,1053,1155,1179]"pageId="20"pageNumber="513">T. R. Bell</collectorName>
, Karachi.”/ “
<collectorNameid="394D533CEB7CED51FF63FB0C624B82C0"box="[134,364,1187,1211]"pageId="20"pageNumber="513">H. E. Andrewes Coll.</collectorName>
<taxonomicNameid="53B84D69EB7CED51FEFBFB6C612782A0"authority="Kavanaugh & Cueva-Dabkoski, 2023"authorityName="Kavanaugh & Cueva-Dabkoski"authorityYear="2023"box="[286,512,1219,1243]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="20"pageNumber="513"phylum="Arthropoda"rank="species"species="pallidus"status="sp. nov.">
<materialsCitationid="24D03CB7EB7CED51FF20FB4B60C38327"ID-GBIF-Occurrence="4905152368"collectionCode="NHMUK"collectorName="D. H. Kavanaugh & M. Cueva-Dabkoski"determinerName="H. E. Andrewes"pageId="20"pageNumber="513"specimenCount="1"specimenCount-female="1"typeStatus="paratype">
<collectionCodeid="F2A9AE2FEB7CED51FE17FB4B617D8280"box="[498,602,1252,1275]"country="United Kingdom"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665"lsid="urn:lsid:biocol.org:col:34665"name="Natural History Museum, London"pageId="20"pageNumber="513"type="Museum">NHMUK</collectionCode>
, labeled: “
<collectorNameid="394D533CEB7CED51FD2FFB4C60128280"box="[714,821,1251,1275]"pageId="20"pageNumber="513">T. R. Bell</collectorName>
, Karachi.”/ “In loose sand where it hid itself by burrowing …[illegible]…28 6 04” [handwritten label]/ “
<taxonomicNameid="53B84D69EB7CED51FBD2FA8C62248327"authority="Kavanaugh & Cueva-Dabkoski, 2023"authorityName="Kavanaugh & Cueva-Dabkoski"authorityYear="2023"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="20"pageNumber="513"phylum="Arthropoda"rank="species"species="pallidus"status="sp. nov.">
<taxonomicNameid="53B84D69EB7CED51FD05FA2B601183E7"authorityName="Kavanaugh & Cueva-Dabkoski"authorityYear="2023"box="[736,822,1412,1436]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="20"pageNumber="513"phylum="Arthropoda"rank="species"species="pallidus">
<taxonomicNameid="53B84D69EB7CED51FE6AFA6A612283A7"authorityName="Kavanaugh & Cueva-Dabkoski"authorityYear="2023"box="[399,517,1476,1500]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="20"pageNumber="513"phylum="Arthropoda"rank="species"species="pallidus">
in Asia by the following combination of character states: Body size slightly small for genus, BL females =
<quantityid="53409B0FEB7CED51FF0AF9AB62AD8067"box="[239,394,1540,1564]"metricMagnitude="-3"metricUnit="m"metricValue="6.85"metricValueMax="6.9"metricValueMin="6.8"pageId="20"pageNumber="513"unit="mm"value="6.85"valueMax="6.9"valueMin="6.8">6.8 to 6.9 mm</quantity>
(male unknown); body form (
<figureCitationid="0C832A6FEB7CED51FD3CF9AB60088067"box="[729,815,1540,1564]"captionStart="FIGURE"captionStartId="21.[171,183,1155,1174]"captionTargetBox="[120,1176,216,1138]"captionTargetId="figure-233@21.[120,1176,216,1147]"captionTargetPageId="21"captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512976"httpUri="https://zenodo.org/record/11512976/files/figure.png"pageId="20"pageNumber="513">Fig. 9A</figureCitation>
) shorter (ratio BL/EW = 1.38 to 1.39), roundly ovoid, with elytra widest at mid-length; dorsal surface (
<figureCitationid="0C832A6FEB7CED51FC9BF98A60F38046"box="[894,980,1573,1597]"captionStart="FIGURE"captionStartId="21.[171,183,1155,1174]"captionTargetBox="[120,1176,216,1138]"captionTargetId="figure-233@21.[120,1176,216,1147]"captionTargetPageId="21"captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512976"httpUri="https://zenodo.org/record/11512976/files/figure.png"pageId="20"pageNumber="513">Fig. 9A</figureCitation>
) mainly pale tan-orange, with only pronotal disc, elytral basal dark band and longitudinal dark band on elytral interval 1 slightly darker brown, middle dark band and preapical dark spot absent; pronotum (
<figureCitationid="0C832A6FEB7CED51FBD5F9CA67A98006"box="[1072,1166,1637,1661]"captionStart="FIGURE"captionStartId="33.[171,183,1573,1592]"captionTargetBox="[139,1158,216,1560]"captionTargetId="figure-14@33.[135,1162,216,1560]"captionTargetPageId="33"captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512988"httpUri="https://zenodo.org/record/11512988/files/figure.png"pageId="20"pageNumber="513">Fig. 14E</figureCitation>
) short and very broad (ratio PWM/PL = 2.71 to 2.79), more broadened basally (ratio PWM/PWA = 1.70 to 1.73), very broadly flattened laterally (especially anteriorly), lateral pale bands very broad and only slightly contrasting with color on disc, anterior angles broader, lateral margin not sinuate near anterior angles; free apex of prosternal intercoxal process short (as in
<figureCitationid="0C832A6FEB7DED50FC4DFAAF672B8363"box="[936,1036,1280,1304]"captionStart="FIGURE"captionStartId="34.[171,183,869,888]"captionTargetBox="[140,1176,216,846]"captionTargetId="figure-9@34.[120,1176,216,846]"captionTargetPageId="34"captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512990"httpUri="https://zenodo.org/record/11512990/files/figure.png"pageId="21"pageNumber="514">Fig. 15D</figureCitation>
); elytra with striae moderately impressed and intervals flat or nearly so; elytral epipleura with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; males unknown; specimen from southeastern
<figureCitationid="0C832A6FEB7DED50FDFDFACF614E8303"box="[536,617,1376,1400]"captionStart="FIGURE"captionStartId="39.[171,183,1593,1612]"captionTargetBox="[168,1128,979,1582]"captionTargetId="figure-84@39.[168,1128,979,1582]"captionTargetPageId="39"captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km."figureDoi="http://doi.org/10.5281/zenodo.11513002"httpUri="https://zenodo.org/record/11513002/files/figure.png"pageId="21"pageNumber="514">Fig. 21</figureCitation>
<taxonomicNameid="53B84D69EB7DED50FEECFB2C61C182ED"authority="Kavanaugh and Cueva-Dabkoski"authorityName="Kavanaugh and Cueva-Dabkoski"authorityYear="2023"box="[265,742,1155,1174]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="514"phylum="Arthropoda"rank="species"species="pallidus"status="sp. nov.">
<paragraphid="940736EAEB7DED50FF9EFB33616A82CB"blockId="21.[123,1174,1155,1200]"box="[123,589,1180,1200]"pageId="21"pageNumber="514">Labels associated with holotype. Scale line = 1.0 mm.</paragraph>
<taxonomicNameid="53B84D69EB7DED50FED1FA2E628E83E3"authorityName="Kavanaugh and Cueva-Dabkoski"authorityYear="2023"box="[308,425,1408,1432]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="514"phylum="Arthropoda"rank="species"species="pallidus">
<taxonomicNameid="53B84D69EB7DED50FBC2FA2E638183C3"authorityName="Kavanaugh & Cueva-Dabkoski"authorityYear="2023"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="514"phylum="Arthropoda"rank="species"species="vespertilio">
<quantityid="53409B0FEB7DED50FCDBFA6F60FB83A3"box="[830,988,1472,1496]"metricMagnitude="-3"metricUnit="m"metricValue="6.85"metricValueMax="6.9"metricValueMin="6.8"pageId="21"pageNumber="514"unit="mm"value="6.85"valueMax="6.9"valueMin="6.8">6.8 to 6.9 mm</quantity>
; body form (
<figureCitationid="0C832A6FEB7DED50FB8FFA6F63BA8383"captionStart="FIGURE"captionStartId="21.[171,183,1155,1174]"captionTargetBox="[120,1176,216,1138]"captionTargetId="figure-233@21.[120,1176,216,1147]"captionTargetPageId="21"captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512976"httpUri="https://zenodo.org/record/11512976/files/figure.png"pageId="21"pageNumber="514">Fig. 9A</figureCitation>
) shorter (ratio BL/EW = 1.38 to 1.39), roundly ovoid, with elytra widest at mid-length.
</paragraph>
<paragraphid="940736EAEB7DED50FF4EF9AF671080C3"blockId="21.[123,1174,1248,1721]"pageId="21"pageNumber="514">Color. Head with dorsum, venter, and all mouthparts and other appendages light brown. Pronotum with disc light brown and lateral pale bands very wide and pale tan-orange. Elytra mainly yellow-tan, with elytral base, scutellum, and longitudinal dark band on elytral interval 1 darker brown, basal dark band diffusely and very slightly darker than pale disc, middle dark band and preapical dark spot absent. Venter light brown, proepipleura tan-orange, elytral epipleura yellow-tan, and apical portion of last abdominal sternite pale yellow-tan. Legs pale, light brown.</paragraph>
<paragraphid="940736EAEB7EED53FF4EFF7460028728"blockId="22.[123,1174,219,1463]"pageId="22"pageNumber="515">Reflection, luster, and microsculpture. Dorsum and venter without metallic reflection. Head and pronotum dull, with microsculpture moderately impressed and comprised of isodiametric meshes; elytra slightly shiny with isodiametric meshes slightly less deeply impressed; venter with isodiametric to slightly transverse meshes shallowly impressed.</paragraph>
<paragraphid="940736EAEB7EED53FF4EFEF4620987A8"blockId="22.[123,1174,219,1463]"pageId="22"pageNumber="515">Head. Eyes large, hemispheric; antennae slightly short, extended only to basal one-fifth of elytra; clypeus bisetose; labrum with apical margin deeply emarginate, three pairs of setae present; mentum asetose, with a broad, apically emarginate medial tooth; submentum anteriorly with a single pair of setae.</paragraph>
<figureCitationid="0C832A6FEB7EED53FE7FFE7362DE878F"box="[410,505,476,500]"captionStart="FIGURE"captionStartId="33.[171,183,1573,1592]"captionTargetBox="[139,1158,216,1560]"captionTargetId="figure-14@33.[135,1162,216,1560]"captionTargetPageId="33"captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512988"httpUri="https://zenodo.org/record/11512988/files/figure.png"pageId="22"pageNumber="515">Fig. 14E</figureCitation>
) trapezoidal, almost as wide as elytra at humeral angles, short and very broad (ratio PWM/PL = 2.71 to 2.79), more broadened basally (ratio PWM/PWA = 1.70 to 1.73), very broadly flattened laterally (especially anteriorly); anterior angles broad, narrowly rounded apically; anterior margin between anterior angles nearly straight at middle; lateral margins gently arcuate, not sinuate near anterior angles; basal margin moderately bisinuate, faintly lobate medially; anterior and basal margination distinct laterally, obsolete medially; lateral margination very slender and finely impressed; midlateral setae present, inserted just medial to lateral margination and at about one-third the distance between the anterior and basal angles along the lateral margin; basolateral setae present, inserted on lateral edge of pronotum and just anterior to posterior angles. Prosternal intercoxal process moderately long and slender, lanceolate, with complete margination, free apex of process short (as in
<figureCitationid="0C832A6FEB7EED53FD34FCB26014854E"box="[721,819,797,821]"captionStart="FIGURE"captionStartId="34.[171,183,869,888]"captionTargetBox="[140,1176,216,846]"captionTargetId="figure-9@34.[120,1176,216,846]"captionTargetPageId="34"captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512990"httpUri="https://zenodo.org/record/11512990/files/figure.png"pageId="22"pageNumber="515">Fig. 15D</figureCitation>
<figureCitationid="0C832A6FEB7EED53FE4FFC926123852E"box="[426,516,829,853]"captionStart="FIGURE"captionStartId="21.[171,183,1155,1174]"captionTargetBox="[120,1176,216,1138]"captionTargetId="figure-233@21.[120,1176,216,1147]"captionTargetPageId="21"captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512976"httpUri="https://zenodo.org/record/11512976/files/figure.png"pageId="22"pageNumber="515">Fig. 9A</figureCitation>
), slightly shorter than wide (ratio EL/EW = 0.90), broadly rounded apically, humeri slightly obtuse, angulate; elytral striae moderately impressed throughout; elytral intervals flat or nearly so, smooth; parascutellar setiferous pore present at base of interval 1 near junction of striae 1 and 2; two discal setiferous pores present on interval 3 adjacent to stria 2, one inserted just posterior to elytral mid-length and the other inserted near apical one-third; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 12 to 13 setae.
Legs. Middle and hind tibiae with medial tibial spurs long and serrulate (as in
<figureCitationid="0C832A6FEB7EED53FBCAFC5267AE826E"box="[1071,1161,1021,1045]"captionStart="FIGURE"captionStartId="2.[171,183,897,916]"captionTargetBox="[120,1176,216,878]"captionTargetId="figure-193@2.[120,1176,216,878]"captionTargetPageId="2"captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.11512958"httpUri="https://zenodo.org/record/11512958/files/figure.png"pageId="22"pageNumber="515">Fig. 2A</figureCitation>
). Females with front tarsomeres 1 to 3 markedly expanded laterally (as in
<figureCitationid="0C832A6FEB7EED53FC47FBB26724824E"box="[930,1027,1053,1077]"captionStart="FIGURE"captionStartId="2.[171,183,897,916]"captionTargetBox="[120,1176,216,878]"captionTargetId="figure-193@2.[120,1176,216,878]"captionTargetPageId="2"captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.11512958"httpUri="https://zenodo.org/record/11512958/files/figure.png"pageId="22"pageNumber="515">Figs. 2C</figureCitation>
), but without adhesive setae ventrally, and middle tarsomeres slender and without adhesive setae ventrally; tarsal claws smooth.
<taxonomicNameid="53B84D69EB7EED53FF9EFB3063D282CD"authorityName="Kavanaugh and Cueva-Dabkoski"authorityYear="2023"box="[123,245,1182,1206]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="515"phylum="Arthropoda"rank="species"species="pallidus">
states that it was collected from “loose sand” and that it “hid by burrowing…,” presumably into that sand. There are several large rivers with sandy shores in the Karachi region, as well as extensive sandy sea beaches and associated dune systems, so the species may occur in either or both of these habitat types.
<figureCitationid="0C832A6FEB7EED53FE04FAB1610A834D"box="[481,557,1310,1334]"captionStart="FIGURE"captionStartId="39.[171,183,1593,1612]"captionTargetBox="[168,1128,979,1582]"captionTargetId="figure-84@39.[168,1128,979,1582]"captionTargetPageId="39"captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km."figureDoi="http://doi.org/10.5281/zenodo.11513002"httpUri="https://zenodo.org/record/11513002/files/figure.png"pageId="22"pageNumber="515">Fig. 21</figureCitation>
<taxonomicNameid="53B84D69EB7EED53FF72FAD0622C83EC"authorityName="Kavanaugh and Cueva-Dabkoski"authorityYear="2023"box="[151,267,1407,1431]"class="Insecta"family="Carabidae"genus="Cyclosomus"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="515"phylum="Arthropoda"rank="species"species="pallidus">
<collectionCodeid="F2A9AE2FEB7EED53FC74FAD060DD83EC"box="[913,1018,1407,1431]"country="United Kingdom"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665"lsid="urn:lsid:biocol.org:col:34665"name="Natural History Museum, London"pageId="22"pageNumber="515"type="Museum">NHMUK</collectionCode>
