<mods:titleid="369D8DFD23B8ED1D090E45C4EDD2021C">First description of the male Quelaestrygon puetzi Smetana, 1999 (Coleoptera, Staphylinidae, Staphylinini) from China</mods:title>
<mods:affiliationid="DC2C66245BC54C87917BC0A0A52D7624">Morphological Laboratory, Guizhou University of Traditional Chinese Medicine, Guiyang, 550025, Guizhou, China</mods:affiliation>
<mods:affiliationid="0629BB407180D0F51B4041CDF67F419F">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, China & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing, 100049, China</mods:affiliation>
<figureCitationid="6861AABBAC25D14AD682A29EBCF3D07C"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Male of Quelaestrygon puetzi Smetana A head in dorsal view B pronotum in dorsal view C elytra in dorsal view, showing sculpture D head in lateral view, showing postmandibular ridge (a) and infraorbital ridge (b) E antenna, detail. Scale bars: 2.00 mm."figureDoi="10.3897/zookeys.944.53482.figure1"httpUri="https://binary.pensoft.net/fig/426934"pageId="0"pageNumber="121">Figs 1</figureCitation>
<figureCitationid="6E737591ADB9E74AB4E7C571F2C5B8FC"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Male of Quelaestrygon puetzi Smetana, morphology A body in dorsal view B aedeagus, underside of paramere C aedeagus, apical portion of median lobe, parameral view. Scale bars: 5.00 mm (A), 0.50 mm (B, C)."figureDoi="10.3897/zookeys.944.53482.figure2"httpUri="https://binary.pensoft.net/fig/426935"pageId="0"pageNumber="121">, 2</figureCitation>
<figureCitationid="EC0B9EBC5A032657D9B93DFBFA7A422C"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">, 3</figureCitation>
<bibRefCitationid="A435AA8B9F2D69FF9D39A8A8A54C0E8B"author="Smetana, A"journalOrPublisher="Elytra"pageId="0"pageNumber="121"pagination="241 - 248"refId="B4"refString="Smetana, A, 1999. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 14. Quelaestrygon puetzi gen. nov., sp. nov. from Sichuan. Elytra 27 (1): 241 - 248"title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 14. Quelaestrygon puetzi gen. nov., sp. nov. from Sichuan."volume="27"year="1999">Smetana (1999</bibRefCitation>
Male with first four segments of foretarsus strongly dilated, sub-bilobed, each heavily covered with tenent setae ventrally, segment II slightly wider than apex of tibia; tergite VIII (Fig.
<figureCitationid="527783385B88C5FFAB2C14154DDAC357"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3A</figureCitation>
) with basal ridge complete, nearly straight, with one long seta on each side, apical margin with shallow and narrow medioapical emargination; sternite VIII (Fig.
<figureCitationid="373F447C09D4DFF622D3F0E007D3F891"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3B</figureCitation>
) with basal ridge complete, slightly sinuate, with one long seta on each side, apical margin with shallow and wide medioapical emargination, a very small triangular area in front of the emargination impunctate; sternite IX (Fig.
<figureCitationid="F705BCD99C2CDADF62DA062CF34D317A"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3C</figureCitation>
<figureCitationid="D108CDC95005729364E64C59E886F2C7"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3D</figureCitation>
) with basal side broadly and shallowly concave, apical margin complete, forming a right angle; aedeagus robust and strongly sclerotized, in lateral view (Fig.
<figureCitationid="AA116742A56350A22F7CAB39BEA51DE0"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3E</figureCitation>
) with apex of paramere distinctly not reaching that of median lobe, apical 1/4 of median lobe strongly bent toward parameral side, without any process at apex; aedeagus in parameral view (Fig.
<figureCitationid="D8E92379690962E3A63D6C4D97ED25A6"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3F</figureCitation>
) with paramere distinctly narrower than median lobe, wide at base, then gradually narrowed into obtuse apex, forming a near triangle shape, median lobe parallel-sided laterally, distinctly constricted at apex, apex somewhat pointed (Figs
<figureCitationid="5851D5351A84F214DBE5A6FCD420D404"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Male of Quelaestrygon puetzi Smetana, morphology A body in dorsal view B aedeagus, underside of paramere C aedeagus, apical portion of median lobe, parameral view. Scale bars: 5.00 mm (A), 0.50 mm (B, C)."figureDoi="10.3897/zookeys.944.53482.figure2"httpUri="https://binary.pensoft.net/fig/426935"pageId="0"pageNumber="121">2C</figureCitation>
<figureCitationid="F6408C6F7A85BD93E182558D921AD59D"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3F</figureCitation>
); apical portion of paramere with four moderately long apical setae, and two similar subapical setae on each lateral side below apex, underside with seven small tooth-shaped processes along the more strongly sclerotized and pigmented midline, and 6-7 spike-like sensory peg setae arranged in cluster far below apex on each side (Figs
<figureCitationid="799D7D6A712D23AA4399EE4300EBE6F7"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Male of Quelaestrygon puetzi Smetana, morphology A body in dorsal view B aedeagus, underside of paramere C aedeagus, apical portion of median lobe, parameral view. Scale bars: 5.00 mm (A), 0.50 mm (B, C)."figureDoi="10.3897/zookeys.944.53482.figure2"httpUri="https://binary.pensoft.net/fig/426935"pageId="0"pageNumber="121">2B</figureCitation>
<figureCitationid="00B8E680074228080338CFE7C007AD12"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Male terminalia and genitalia of Quelaestrygon puetzi Smetana A male tergite VIII B male sternite VIII C male sternite IX D male tergite X E aedeagus, lateral view F aedeagus, parameral view G aedeagus, underside of paramere. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.944.53482.figure3"httpUri="https://binary.pensoft.net/fig/426936"pageId="0"pageNumber="121">3G</figureCitation>
Smetana is at present known only from several mountain areas in Sichuan Province of China: Mt. Gongga, Mt. Jinping and Mt. Emei. The examined male specimen was hand-collected at a parking lot after landing on the
