117 lines
13 KiB
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117 lines
13 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.544.6074" ID-GBIF-Dataset="0e5e47a0-f8f3-4039-b0e1-c159fef0b9f6" ID-PMC="PMC4714384" ID-Pensoft-Pub="1313-2970-544-1" ID-PubMed="26798289" ID-ZBK="C5978BD0145B40F8ACDEB27371B7B9A4" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-544-1" ModsDocOrigin="ZooKeys 544" ModsDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" checkinTime="1451243640953" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2015" docId="84ED6BB059E71BCFA9897014E9D2B0F2" docLanguage="en" docName="ZooKeys 544: 1-407" docOrigin="ZooKeys 544" docSource="http://dx.doi.org/10.3897/zookeys.544.6074" docTitle="Mecyclothorax patulus Liebherr, 2015, sp. n." docType="treatment" docUuid="F43541E5-4856-4FF3-858D-73BE0BBBB639" docUuidSource="ZooBank" docVersion="5" lastPageNumber="117" masterDocId="FF92FFC2FF9FFFC7191EFFB8FF91AF5D" masterDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" masterLastPageNumber="407" masterPageNumber="1" pageNumber="116" updateTime="1668162300918" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>544</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>407</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.544.6074</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.544.6074</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-544-1</mods:identifier>
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<mods:identifier type="ZBK">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
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<mods:identifier type="ZooBank">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127853641" LSID="urn:lsid:zoobank.org:act:F43541E5-4856-4FF3-858D-73BE0BBBB639" httpUri="http://treatment.plazi.org/id/84ED6BB059E71BCFA9897014E9D2B0F2" lastPageId="116" lastPageNumber="117" pageId="115" pageNumber="116">
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<subSubSection pageId="115" pageNumber="116" type="multiple">
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<paragraph pageId="115" pageNumber="116">
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<pageBreakToken pageId="115" pageNumber="116" start="start">Taxon</pageBreakToken>
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classification Animalia Coleoptera Carabidae
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</paragraph>
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</subSubSection>
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<subSubSection pageId="115" pageNumber="116" type="nomenclature">
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<paragraph pageId="115" pageNumber="116">
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(044)
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<taxonomicName LSID="http://zoobank.org/F43541E5-4856-4FF3-858D-73BE0BBBB639" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax patulus" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="patulus">Mecyclothorax patulus</taxonomicName>
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<taxonomicNameLabel pageId="115" pageNumber="116">sp. n.</taxonomicNameLabel>
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Figs 60B, 64
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</paragraph>
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</subSubSection>
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<subSubSection pageId="115" pageNumber="116" type="diagnosis">
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<paragraph pageId="115" pageNumber="116">Diagnosis.</paragraph>
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<paragraph pageId="115" pageNumber="116">
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This second of the three bicolored species (Fig. 60B) exhibits more narrowly flavous elytral margins-only intervals 8-9-versus piceous discal intervals 2-7. The sutural interval is rufous basally, flavous apically. The bisetose pronotum is more transverse than that of
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax subtilis" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="subtilis">Mecyclothorax subtilis</taxonomicName>
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; MPW/PL = 1.28, and less constricted basally, MPW/BPW = 1.53. As in the next species below,
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax patagiatus" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="patagiatus">Mecyclothorax patagiatus</taxonomicName>
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, the parascutellar and subapical elytral setae are present, but the elytral humeri are narrower in this species-MEW/HuW = 2.06 versus 2.0-and the elytra more ellipsoid. The setal impressions of the dorsal elytral setae are larger in beetles of this species, as they span interval 3. Setal formula 2 1 2 1[sae]. Standardized body length 3.4 mm.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="116" lastPageNumber="117" pageId="115" pageNumber="116" type="description">
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<paragraph pageId="115" pageNumber="116">Description</paragraph>
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<paragraph lastPageId="116" lastPageNumber="117" pageId="115" pageNumber="116">
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(n = 1). Head capsule with frontal grooves broad near clypeus, a lateral carina extended to mesad anterior supraorbital seta; dorsal surface of neck flat; eyes moderately convex, ocular ratio = 1.47, not extended onto posterior portion of ocular lobe, ocular lobe ratio = 0.75; labral anterior margin broadly emarginate to 1/6 of length; antennae filiform, sparse setae on apex of antennomere 1 and shafts of antennomeres 2-3; mentum tooth with sides acute, apex tightly rounded. Pronotum transverse, MPW/PL = 1.28, bisetose, glabrous hind angles obtuse, rounded behind, lateral margin subparallel for short distance anterad angle; base moderately broad, MPW/BPW = 1.53; median base nearly coplanar with disc, ~10 sparsely distributed, isolated punctures each side; basal margin convexly expanded between laterobasal depressions; median longitudinal impression shallow, finely incised, crossed by fine longitudinal wrinkles; anterior transverse impression deep, finely incised, minute irregularities in deepest part; anterior callosity convex, glossy surface with minute longitudinal wrinkles; front angles slightly projected, tightly rounded; pronotal apical width greater than basal width, APW/BPW = 1.10; lateral marginal depression narrow, edge upturned, slightly broader at front angle; laterobasal depression with slightly irregular surface, continuous with lateral depression. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with narrow median impression, lateral margins broadly beaded between coxae. Elytra broadly subellipsoid, disc flat, sides moderately sloped; basal groove evenly and distinctly recurved to tightly rounded humeral angle, MEW/HuW = 2.06; parascutellar seta present; parascutellar striole with 3-4 punctures, very shallow between punctures; sutural interval more convex than lateral intervals, sutural juncture upraised; sutural and 2nd striae of subequal depth from base to disc, 2nd stria slightly shallower at apex; discal striae 1-5 moderately broad, evident, stria 6 shallower and stria 7 shallower still, interrupted along length; striae 1-4 with minute elongate punctures, stria 5 with shallower punctures and stria 6 with only irregularities along length; intervals 2-5 moderately convex, lateral intervals less so; 8th interval slightly more convex than fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.28
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<normalizedToken originalValue="–0.32×">-0.32x</normalizedToken>
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and 0.62
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<normalizedToken originalValue="×">x</normalizedToken>
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elytral length, setal impressions evident, spanning interval 3; apical elytral seta absent, subapical elytral seta present; lateral elytral setae arranged as
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<pageBreakToken pageId="116" pageNumber="117" start="start">anterior</pageBreakToken>
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series of 6 setae and posterior series of 5 setae; elytral marginal depression narrow, edge little upturned in basal half, margin beadlike near subapical sinuation; subapical sinuation very shallow, nearly obsolete. Mesepisternum with ~6 punctures in 1-2 rows; metepisternal width to length ratio = 0.80; metepisternum/metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1-5, lateral depressions on ventrites 3-6; suture between ventrites 2 and 3 complete; apical female abdominal ventrite with 4 equally spaced setae plus trapezoid of 4 subequal, short setae. Legs-metatarsomere 1/metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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medial tarsomere length, apical and subapical setae present; metatarsal dorsolateral sulci narrow, shallow, median surface broad. Microsculpture of vertex shallow isodiametric sculpticells in rows; pronotal disc with shallow transverse mesh, sculpticell breadth 3
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<normalizedToken originalValue="×">x</normalizedToken>
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length, to transverse lines not connected into mesh; pronotal median base glossy, obsolete transverse mesh between punctures; elytral disc and apex with shallow transverse mesh, sculpticell breadth 2
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<normalizedToken originalValue="–4×">-4x</normalizedToken>
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length; metasternum with transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, antennomeres 2-3 rufoflavous, 4-11 rufobrunneous; pronotal disc rufobrunneous with piceous cast, lateral margins, apex, and base rufoflavous; proepipleuron flavous, proepisternum rufobrunneous with piceous cast; elytral epipleuron dorsally flavous, ventrally rufoflavous, metepisternum rufobrunneous with piceous cast; abdomen with ventrites 1-2 medially, and 3-5 mediobasally rufopiceous, ventrites 3-6 apical and marginally rufoflavous; metafemur flavous; metatibia flavous with brunneous cast.
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</paragraph>
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<paragraph pageId="116" pageNumber="117">Female reproductive tract. The lone female holotype was not dissected.</paragraph>
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</subSubSection>
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<subSubSection pageId="116" pageNumber="117" type="holotype">
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<paragraph pageId="116" pageNumber="117">Holotype.</paragraph>
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<paragraph pageId="116" pageNumber="117">
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Female (NMNH) labeled: HI:Maui Koolau / F.R. Kula Pipeline Rd. /
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="20.809668">N20°48.58'</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-156.23833">W156°14.30'</geoCoordinate>
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, / 18-V-2003 lot09 el. 1305m / pyr. fog log D.A. Polhemus // HOLOTYPE /
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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/
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<taxonomicName lsidName="patulus" pageId="116" pageNumber="117" rank="species" species="patulus">patulus</taxonomicName>
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/ Liebherr / det. J.K. Liebherr 2015 (black-margined red label).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="116" pageNumber="117" type="etymology">
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<paragraph pageId="116" pageNumber="117">Etymology.</paragraph>
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<paragraph pageId="116" pageNumber="117">
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The Latin adjective
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<taxonomicName lsidName="patulus" pageId="116" pageNumber="117" rank="species" species="patulus">patulus</taxonomicName>
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means open, spread out, or broad (
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<bibRefCitation author="Brown, RW" journalOrPublisher="Smithsonian Institution Press, Washington, D. C." pageId="307" pageNumber="308" title="Composition of Scientific Words" year="1956">Brown 1956</bibRefCitation>
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), signifying the short, broad body of these beetles.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="116" pageNumber="117" type="distribution and habitat">
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<paragraph pageId="116" pageNumber="117">Distribution and habitat.</paragraph>
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<paragraph pageId="116" pageNumber="117">
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax patulus" order="Coleoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="patulus">Mecyclothorax patulus</taxonomicName>
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is known only from an
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<normalizedToken originalValue="‘Ōhi‘a">'Ōhi'a</normalizedToken>
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Wet Forest site uphill from the junction of the Kula Pipeline and Waikamoi Flume Roads (Fig. 64). The type specimen was found in a pyrethrin fog sample of a downed
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<normalizedToken originalValue="‘ōhi‘a">'ōhi'a</normalizedToken>
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log covered with moss.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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