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151 lines
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<document id="767B87C2E8A1E506440082231DDC4D2C" ID-CLB-Dataset="35618" ID-DOI="http://dx.doi.org/10.3897/zookeys.611.9058" ID-GBIF-Dataset="ded1c4ec-7600-49bf-8f39-d20eb6b00849" ID-PMC="PMC4992807" ID-Pensoft-Pub="1313-2970-611-57" ID-PubMed="27594799" ID-ZooBank="91F749DA0AFE41C69B4D626B147EA0DA" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-611-57" ModsDocOrigin="ZooKeys 611" ModsDocTitle="Revisions of the genera Lurama Schaus, 1928 and Ulmara Schaus, 1928 (Lepidoptera, Mimallonoidea, Mimallonidae) with the descriptions of three new Ulmara species and a new genus" checkinTime="1471317591627" checkinUser="pensoft" docAuthor="St. Laurent, Ryan A." docDate="2016" docId="66B35535BD7BD60CAD82DBDBFB76834F" docLanguage="en" docName="ZooKeys 611: 57-92" docOrigin="ZooKeys 611" docSource="http://dx.doi.org/10.3897/zookeys.611.9058" docTitle="Lurama penia Dognin 1919" docType="treatment" docVersion="7" lastPageNumber="62" masterDocId="180EFFC04F09FFB5FFAEFF9DA6627021" masterDocTitle="Revisions of the genera Lurama Schaus, 1928 and Ulmara Schaus, 1928 (Lepidoptera, Mimallonoidea, Mimallonidae) with the descriptions of three new Ulmara species and a new genus" masterLastPageNumber="92" masterPageNumber="57" pageNumber="61" updateTime="1701377410659" updateUser="ExternalLinkService">
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<mods:title id="4692560CD867EC6D5F819E4E66982AEC">Revisions of the genera Lurama Schaus, 1928 and Ulmara Schaus, 1928 (Lepidoptera, Mimallonoidea, Mimallonidae) with the descriptions of three new Ulmara species and a new genus</mods:title>
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<mods:namePart id="8E0F3F0992554C003FFED6E759812EAF">St. Laurent, Ryan A.</mods:namePart>
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<mods:typeOfResource id="902AC7B599DAE2944E125C3E74172433">text</mods:typeOfResource>
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<mods:title id="F79C2CCDE5B22776E168DBD87C812E51">ZooKeys</mods:title>
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<mods:date id="E6A6BB98AA10798DDF416765321C9D9C">2016</mods:date>
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<mods:detail id="1D5CBCF2D1133CEB46D14AFAC0B2CCCE" type="volume">
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<mods:number id="8FE43B638D06BDE76B23882999D7B946">611</mods:number>
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<mods:start id="0C42911717247EFBABCA7387832DC506">57</mods:start>
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<mods:end id="51AB2F81CCDC0AB1460537F8D5B32086">92</mods:end>
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<mods:url id="902AFFFA83310B57CD4001EDBCE8EBB4">http://dx.doi.org/10.3897/zookeys.611.9058</mods:url>
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</mods:location>
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<mods:classification id="5B960A283DA8AC165AD609C267F64733">journal article</mods:classification>
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<mods:identifier id="1C7C54BE651B131B014CBBEAAD29B0E4" type="DOI">http://dx.doi.org/10.3897/zookeys.611.9058</mods:identifier>
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<mods:identifier id="874D0CABB70BDBDB89859D2B51AEF4CD" type="Pensoft-Pub">1313-2970-611-57</mods:identifier>
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</mods:mods>
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<treatment id="66B35535BD7BD60CAD82DBDBFB76834F" ID-GBIF-Taxon="127862675" LSID="urn:lsid:plazi:treatment:66B35535BD7BD60CAD82DBDBFB76834F" httpUri="http://treatment.plazi.org/id/66B35535BD7BD60CAD82DBDBFB76834F" lastPageId="5" lastPageNumber="62" pageId="4" pageNumber="61">
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<subSubSection id="90E739D5327642DF0425BFB3968047BC" pageId="4" pageNumber="61" type="multiple">
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<paragraph id="026EBE813B0571AD5D97A7D0D41C8D48" pageId="4" pageNumber="61">
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<pageBreakToken id="8596E30DC75C686F017FC009FC047F44" pageId="4" pageNumber="61" start="start">Taxon</pageBreakToken>
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classification Animalia Lepidoptera Mimallonidae
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</paragraph>
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</subSubSection>
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<subSubSection id="12BB48C45E67C3D5839316FBD6CC4902" pageId="4" pageNumber="61" type="nomenclature">
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<paragraph id="F25CBBFB8831676102E7BBB287AD491F" pageId="4" pageNumber="61">
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<taxonomicName id="A48B705666BB426748204369E838B0FC" ID-CoL="3WFTY" authority="Dognin, 1919" authorityName="Dognin" authorityYear="1919" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia (Dognin, 1919)</taxonomicName>
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Fig. 1; Map 1
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</paragraph>
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</subSubSection>
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<subSubSection id="13486F3856C7CA7185E6BBD5D0EA4A35" pageId="4" pageNumber="61" type="reference_group">
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<paragraph id="20F7AF16F9C45E69990DD3F4FF3E34BD" pageId="4" pageNumber="61">
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<taxonomicName id="6981EC79F28150FB55BCA26F3CA8FCA6" class="Insecta" family="Mimallonidae" genus="Perophora" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Perophora penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Perophora penia</taxonomicName>
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Dognin, 1919: 6, 7
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</paragraph>
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<paragraph id="323B26AAF9BC2E49D19253AA39E4B536" pageId="4" pageNumber="61">
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<taxonomicName id="3B3706816B9364BA16ED6E041413FEA0" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia</taxonomicName>
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;
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<bibRefCitation id="A467DFCA3C31F1F22DE54057DBEA1782" author="Schaus, W" editor="Seitz, A" journalOrPublisher="A. Kernen, Stuttgart" pageId="29" pageNumber="86" pagination="635 - 672" title="Familie Mimallonidae." volumeTitle="Die GrossSchmetterlinge der Erde. 6. Die amerikanischen Spinner und Schwa ̈ rmer" year="1928">Schaus 1928</bibRefCitation>
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</paragraph>
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<paragraph id="9F1E53D57B5853922436B84D814E4D03" pageId="4" pageNumber="61">
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<taxonomicName id="B69A478B5D44AE79EA6022F958D9BE32" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia</taxonomicName>
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;
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<bibRefCitation id="420F8EA2734A0F2DCC76D23C4786849E" author="Gaede, M" editor="Strand, E" journalOrPublisher="W. Junk, Berlin" pageId="29" pageNumber="86" pagination="2 - 19" title="Pars 50: Mimallonidae." volumeTitle="Lepidopterorum Catalogus" year="1931">Gaede 1931</bibRefCitation>
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</paragraph>
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<paragraph id="CC85047AB623FC3EF28B57C7409602C7" pageId="4" pageNumber="61">
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<taxonomicName id="D201F82A9266190720F309451C63D697" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia</taxonomicName>
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;
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<bibRefCitation id="F27B22BB4536ED4E200B402218A4E71D" author="Becker, VO" journalOrPublisher="Association for Tropical Lepidoptera & Scientific Publishers, Gainesville, Florida" pageId="29" pageNumber="86" pagination="17 - 19" title="Atlas of Neotropical Lepidoptera, Checklist. Part 4 B. Drepanoidea, Bombycoidea, Sphingoidea" year="1996">Becker 1996</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="322320949C816AF89A521C419C4D543D" pageId="4" pageNumber="61" type="type material">
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<paragraph id="01416EC91405EA2A503392A66583653F" pageId="4" pageNumber="61">Type material.</paragraph>
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<paragraph id="21AEDA665D1C64BD486BC9FB53925AD7" pageId="4" pageNumber="61">
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Holotype, ♂. COLOMBIA: Cundinamarca/Distrito Capital: Bogota [
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<normalizedToken id="FB51C6A52E75798EBFD38D7966B94A3B" originalValue="Bogotá">Bogota</normalizedToken>
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], Colombia, 28-3200 m, Coll. Fassl / Dognin, Collection/
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<taxonomicName id="2BD6DCD88FEC844D91BF97AA27CD1671" class="Insecta" family="Mimallonidae" genus="Perophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perophora penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Perophora penia</taxonomicName>
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, type ♂ Dgn./ ♂ genitalia slide. 6 June '28, C.H. #5. [genitalia prep. not located]/ [label with wing vein sketch]/ [Holo]Type No.: 29698 U.S.N.M./ USNM-Mimal: 1108/ (USNM, examined). Type locality: Colombia: Cundinamarca/Distrito Capital:
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<normalizedToken id="523FD68D3EE91D8B5A12D13CE688994C" originalValue="Bogotá">Bogota</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F8085976B03AF43207BB0793E27323EC" pageId="4" pageNumber="61" type="diagnosis">
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<paragraph id="BB991BDFC5A5B0465AF6BE87980B76F2" pageId="4" pageNumber="61">Diagnosis.</paragraph>
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<paragraph id="1F3D162287FF74034081B7765CE6C4B8" pageId="4" pageNumber="61">
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<taxonomicName id="22F8D91A08C0E6484FF8F4CDA42256B5" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia</taxonomicName>
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can be distinguished from
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<taxonomicName id="3239B9476BC4A830AF9D15753F9D412F" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama quindiuna" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="quindiuna">Lurama quindiuna</taxonomicName>
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by the smaller size, shorter, more rounded wings, and somewhat more diffuse, broader postmedial and antemedial lines on the wings. The lines are also closer together and farther from the wing margin than in
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<taxonomicName id="7627FEE9A6B102D384E0AB1E893C1C4E" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama quindiuna" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="quindiuna">Lurama quindiuna</taxonomicName>
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. Additionally, the forewing postmedial line has its apical angle intersecting Rs4 very near the fork of Rs3+Rs4, rather than being much more distant from this fork in the other species.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D841BA0A6AA14A3814AEE31768AD355" pageId="4" pageNumber="61" type="description">
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<paragraph id="397F6BD0DA9DD7BDEEA27E1D6E6C8E6B" pageId="4" pageNumber="61">Description.</paragraph>
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<paragraph id="EB1FA77BFED29662DD7E9113011C9878" pageId="4" pageNumber="61">Male.Head: As for genus but tan brown. Thorax: Coloration as for head, darker-brown scales present on prothorax. Legs: Coloration as for thorax, poorly preserved. Forewing dorsum: Forewing length: 14.5 mm, wingspan: ~27 mm, n = 1. Short, vaguely triangular, margin nearly straight from apex until after passing M3 where wing smoothly curves toward anal margin. Ground color yellowish tan, overall lightly speckled by dark petiolate scales, especially postmedially. Antemedial line brown, relatively wide, somewhat diffuse, slightly curving outward. Postmedial line nearly straight from anal margin to Rs4 where it abruptly angles toward costa perpendicularly, coloration and width as for antemedial line. Antemedial, medial, and postmedial areas concolorous. Costa and outer wing margin darker brown. Discal spot a dark streak spanning width of discal cell, mesally slightly angled inward toward cell. Forewing ventrum: Similar to dorsum but with more concentrated speckling postmedially; antemedial line nearly absent, postmedial line fainter. Hindwing dorsum: Coloration as for forewing dorsum, antemedial line absent, postmedial line not reaching anterior wing margin. Hindwing ventrum: Following same pattern as forewing ventrum but postmedial line curved outward rather than straight as on dorsum. Abdomen: Partially missing, but anterior segments concolorous with thorax. Genitalia: Not examined. Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="0E33D6DDF08447E6899AD1A019C72812" pageId="4" pageNumber="61" type="distribution">
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<paragraph id="4EF578F7E0B071D41D7ED9CD57E1B8AF" pageId="4" pageNumber="61">Distribution</paragraph>
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<paragraph id="C8DA07EB173DBECC168AE38099AB3CB3" pageId="4" pageNumber="61">
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(Map 1).
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<taxonomicName id="062185B90F2740010E51A9D57C462AA2" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia</taxonomicName>
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is known only from the holotype, collected between 2800 and 3200 m near
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<normalizedToken id="AE7FAA687CBD12B65A8A6F51DDC02317" originalValue="Bogotá">Bogota</normalizedToken>
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, Colombia.
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</paragraph>
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<caption id="4742FC92DB4BD8CBEFD781E39ABB7FBB" pageId="4" pageNumber="61">
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<paragraph id="F082CEB6EB1C3663F758753EB7D75A48" pageId="4" pageNumber="61">
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Map 1. Known distribution of
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<taxonomicName id="2308D6D040925BCFA95808BF084BC481" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Lurama</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="E4EC6B1F74EF946B5FBB9DEE3A10FC84" lastPageId="5" lastPageNumber="62" pageId="4" pageNumber="61" type="remarks">
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<paragraph id="039CEC37BE2D02BD815C8D21292BBDD0" pageId="4" pageNumber="61">Remarks.</paragraph>
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<paragraph id="EF60662C5C57080B7D234A49EE0DFE90" pageId="4" pageNumber="61">
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This species is apparently very rarely collected, as it is known only from the male holotype. Unfortunately, the genitalia preparation of the holotype, probably made by Carl Heinrich (R. Hutchings pers. comm.), is lost and could not be located by the individuals tasked with trying to find it at the USNM.
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<bibRefCitation id="4A63C72EF911493D211793541DD3C1BC" author="Schaus, W" editor="Seitz, A" journalOrPublisher="A. Kernen, Stuttgart" pageId="29" pageNumber="86" pagination="635 - 672" title="Familie Mimallonidae." volumeTitle="Die GrossSchmetterlinge der Erde. 6. Die amerikanischen Spinner und Schwa ̈ rmer" year="1928">Schaus (1928)</bibRefCitation>
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compared the genitalia of
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<taxonomicName id="E172D92FE7FD9E824798CE91E7155CBF" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Lurama</taxonomicName>
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to that of
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<taxonomicName id="2A5E74C71104690C94DC9894CBC249BE" class="Insecta" family="Mimallonidae" genus="Ulmara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulmara" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Ulmara</taxonomicName>
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, and considered them similar. I note that among
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<normalizedToken id="337C3FDFFED11535EFCB7594C22DD2A9" originalValue="Schaus’s">Schaus's</normalizedToken>
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examined
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<taxonomicName id="CE1BDCE01433124ECF3EF1496666470B" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Lurama</taxonomicName>
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material at the USNM and MNHU, the only dissected male
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<taxonomicName id="DAF295976EA7775ACB67E3F9C5FB1C39" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Lurama</taxonomicName>
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specimen is the holotype of
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<taxonomicName id="101109151B7FE9B2EF305E5BC0169733" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia</taxonomicName>
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; therefore it is reasonable to assume that the missing genitalia of
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<taxonomicName id="B58C4E5F0C15956EEBD12976CF203EAB" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama penia" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="penia">Lurama penia</taxonomicName>
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are similar to those of
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<taxonomicName id="7C099C656DC27821CFC668488D5C1693" class="Insecta" family="Mimallonidae" genus="Ulmara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulmara" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Ulmara</taxonomicName>
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because apparently Schaus did not look at any other
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<taxonomicName id="19A05FD7B364658FEAF0D819C9A2F3B4" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Lurama</taxonomicName>
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dissections. This is expected given the similarity between
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<taxonomicName id="24950FBECAD306882170B376B9891A54" class="Insecta" family="Mimallonidae" genus="Ulmara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulmara" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">Ulmara</taxonomicName>
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male genitalia and those of
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<taxonomicName id="E35C6E31EAF2C94E27735185BB263BCE" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama quindiuna" order="Lepidoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="quindiuna">Lurama quindiuna</taxonomicName>
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, for which the male genitalia are available for examination.
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</paragraph>
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<paragraph id="675AAAF6F59C2539832D3E22A96021EA" pageId="5" pageNumber="62">
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<pageBreakToken id="CB2A052AD800AFB2922C1749A6DB91C6" pageId="5" pageNumber="62" start="start">As</pageBreakToken>
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for
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<taxonomicName id="52DA025E29EE5901891A463A3186905E" class="Insecta" family="Mimallonidae" genus="Lurama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lurama quindiuna" order="Lepidoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="quindiuna">Lurama quindiuna</taxonomicName>
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below, the issue of missing type genitalia for these two species results in some difficulties determining the identity of non-type specimens. Fortunately, external diagnostic characteristics are ample enough to maintain both currently described species as valid, but prevents me from describing new species in the genus at this time. It is vital that more material be located for this and the following species at each of their respective type localities so that more conclusive diagnostic characters can be given, particularly in regards to the genitalia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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