treatments-xml/data/0D/73/87/0D7387A3ED38296DFF5EFB074920FBAD.xml

313 lines
39 KiB
XML
Raw Normal View History

2024-06-21 12:29:19 +02:00
<document id="448C206594D8B08C339855B7844974B8" ID-CLB-Dataset="22410" ID-DOI="10.11646/zootaxa.4768.3.5" ID-GBIF-Dataset="d08524b6-1aac-443c-b6f3-dad1c62d9473" ID-ISSN="1175-5326" ID-Zenodo-Dep="3784074" ID-ZooBank="B1334A2D-6272-4A7A-9856-3F0A6C952E43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588576438311" checkinUser="plazi" docAuthor="Decker, Brenna L., Pitts, James P., Yuan, David &amp; Rodriguez, Juanita" docDate="2020" docId="0D7387A3ED38296DFF5EFB074920FBAD" docLanguage="en" docName="zootaxa.4768.3.5.pdf" docOrigin="Zootaxa 4768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Minagenia Banks 1934" docType="treatment" docVersion="6" lastPageNumber="385" masterDocId="F14AFFDBED39296FFFC9FFF24C07FFF6" masterDocTitle="Re-examination of Australian and Oriental species of Minagenia Banks, 1934 (Hymenoptera: Pompilidae), with a new record for the genus in Australia and a new species description" masterLastPageNumber="394" masterPageNumber="383" pageNumber="384" updateTime="1698838363314" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0AEC3F6F44B3AB22A224BD3F4F086FEB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5453361B00F86A11AD2FC5363B6F12BC">
<mods:title id="5C34AAD86DE7BA7162A4F2DF10E466FC">Re-examination of Australian and Oriental species of Minagenia Banks, 1934 (Hymenoptera: Pompilidae), with a new record for the genus in Australia and a new species description</mods:title>
</mods:titleInfo>
<mods:name id="1E4DD941E10E0286A8F5E5EC826EDFA5" type="personal">
<mods:role id="3C68E4BD8BB2C9BB72686897C2FD92E7">
<mods:roleTerm id="6FC519BAB464D375479B940AD46EBFFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="309281F762BB2647DF312079F8E0B2A1">Decker, Brenna L.</mods:namePart>
<mods:affiliation id="6EFEA84D93582B797DE4B4BF5525CC07">Utah State University, Department of Biology, 5305 Old Main Hill, Logan, UT 84322 - 5305, USA. brennaldecker @ gmail. com brennaldecker @ gmail. com; https: // orcid. org / 0000 - 0002 - 9878 - 1903</mods:affiliation>
</mods:name>
<mods:name id="52CBA8E2E59CECF9477F0D472245B7C2" type="personal">
<mods:role id="F775B2CFB7BA5A257E671E9EC37E98FA">
<mods:roleTerm id="0447F087D1156603ECC2EBFBB5284ABE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0FC64980E10C3D15765A6889065A479">Pitts, James P.</mods:namePart>
<mods:affiliation id="80BDA894FBB50109E812CE524E9411C5">Utah State University, Department of Biology, 5305 Old Main Hill, Logan, UT 84322 - 5305, USA. brennaldecker @ gmail. com brennaldecker @ gmail. com; https: // orcid. org / 0000 - 0002 - 9878 - 1903</mods:affiliation>
</mods:name>
<mods:name id="1EA6A435F4E3BE45F1B27A168ED1B320" type="personal">
<mods:role id="5019430D1DF3D29B4E8B9F5CDF23AB46">
<mods:roleTerm id="A289E75C8EEA3751DDB33090F1FC3334">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D38BD8CB17BEB41F522D2B1038378D10">Yuan, David</mods:namePart>
<mods:affiliation id="B35136334D3A66409B723B5D8ED1F480">Australian National Insect Collection, CSIRO National Research Collections Australian, Canberra, ACT 2601, Australia. david. yuan @ csiro. au; https: // orcid. org / 0000 - 0002 - 4864 - 6615</mods:affiliation>
</mods:name>
<mods:name id="6AE101B8F8836719AC5F71E28EDC3924" type="personal">
<mods:role id="00A9B6C7965F09364AC19F029590C64E">
<mods:roleTerm id="EF4079D24EA05CD8DEF95161C968791B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="837B59D4AF71EF6CF0731D9B38A5CC89">Rodriguez, Juanita</mods:namePart>
<mods:affiliation id="80254BF28353038618EACAD4ACC8913D">Australian National Insect Collection, CSIRO National Research Collections Australian, Canberra, ACT 2601, Australia. david. yuan @ csiro. au; https: // orcid. org / 0000 - 0002 - 4864 - 6615</mods:affiliation>
</mods:name>
<mods:typeOfResource id="36FB81CF4C16D155BDAF3BDFF5B09CFF">text</mods:typeOfResource>
<mods:relatedItem id="DFC27B584816B99DCB7370DF3E8A9C35" type="host">
<mods:titleInfo id="7791BD316A7EB7A7064BC9A575C475CC">
<mods:title id="DBAE64F75A74C8C33CECC52CF091806D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2490ED0760507242D349D6EF39D91DE3">
<mods:date id="7EE7518723530904A62A8F116B81300A">2020</mods:date>
<mods:detail id="E1C270C93D87E0013E593E74887540BF" type="pubDate">
<mods:number id="0A28F15FF6A90D72A77074480CE2248F">2020-05-04</mods:number>
</mods:detail>
<mods:detail id="15CA306EEA4F086E037E2EB551825638" type="volume">
<mods:number id="EF95BB0EF8B2FB3021F85C9A6DF24724">4768</mods:number>
</mods:detail>
<mods:detail id="82589DD0A4C84857F5BB5A90FD61677B" type="issue">
<mods:number id="4EB965B9ED8F6BD3C30C699E882EB67D">3</mods:number>
</mods:detail>
<mods:extent id="93330644C3C4124C43DBBA5C5C284226" unit="page">
<mods:start id="ADD9F83C785EFC37CDBA8B6D4C901B6B">383</mods:start>
<mods:end id="C113CD84E37E03F9BA4B06852A1AEA85">394</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ED9A26158EC668AAF015FB6BC27D7D0E">journal article</mods:classification>
<mods:identifier id="A5576AD6E13651706F0554AFDE545DFB" type="CLB-Dataset">22410</mods:identifier>
<mods:identifier id="B3FEF30B25F85784310C26000369C5A9" type="DOI">10.11646/zootaxa.4768.3.5</mods:identifier>
<mods:identifier id="9B83417B4AD2C74A84978058B8A4F687" type="GBIF-Dataset">d08524b6-1aac-443c-b6f3-dad1c62d9473</mods:identifier>
<mods:identifier id="CBB2D69C5E1D331E43DBCC69A832691E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0D9FAD59276AE4B5F133480743A60072" type="Zenodo-Dep">3784074</mods:identifier>
<mods:identifier id="34AED834B146B433CE02677C7D2BDE0B" type="ZooBank">B1334A2D-6272-4A7A-9856-3F0A6C952E43</mods:identifier>
</mods:mods>
<treatment id="0D7387A3ED38296DFF5EFB074920FBAD" ID-DOI="http://doi.org/10.5281/zenodo.3794812" ID-GBIF-Taxon="163886009" ID-Zenodo-Dep="3794812" LSID="urn:lsid:plazi:treatment:0D7387A3ED38296DFF5EFB074920FBAD" httpUri="http://treatment.plazi.org/id/0D7387A3ED38296DFF5EFB074920FBAD" lastPageId="2" lastPageNumber="385" pageId="1" pageNumber="384">
<subSubSection id="CDC0653EED38296EFF5EFB074DBDFAF9" box="[151,442,1269,1295]" pageId="1" pageNumber="384" type="nomenclature">
<paragraph id="856536B5ED38296EFF5EFB074DBDFAF9" blockId="1.[151,442,1269,1295]" box="[151,442,1269,1295]" pageId="1" pageNumber="384">
<heading id="DE2D81D9ED38296EFF5EFB074DBDFAF9" bold="true" box="[151,442,1269,1295]" fontSize="11" level="1" pageId="1" pageNumber="384" reason="1">
<taxonomicName id="42DA4D36ED38296EFF5EFB074DBDFAF9" ID-CoL="5T9R" ID-ENA="1747266" authority="Banks, 1934" authorityName="Banks" authorityYear="1934" box="[151,442,1269,1295]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED38296EFF5EFB074DBDFAF9" bold="true" box="[151,442,1269,1295]" pageId="1" pageNumber="384">
<emphasis id="B7AEEAA7ED38296EFF5EFB074D1CFAF9" bold="true" box="[151,283,1269,1295]" italics="true" pageId="1" pageNumber="384">Minagenia</emphasis>
<bibRefCitation id="E14B4B44ED38296EFEEAFB074DBDFAF9" author="Banks, N." box="[291,442,1269,1295]" pageId="1" pageNumber="384" pagination="1 - 117" refId="ref8911" refString="Banks, N. (1934) The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences, 69, 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED38296EFF5EFAC84936FAA7" box="[151,1329,1336,1362]" pageId="1" pageNumber="384" type="reference_group">
<paragraph id="856536B5ED38296EFF5EFAC84936FAA7" blockId="1.[151,1436,1336,1545]" box="[151,1329,1336,1362]" pageId="1" pageNumber="384">
<taxonomicName id="42DA4D36ED38296EFF5EFAC84D8BFAA7" ID-CoL="5T9R" authority="Banks, 1934" authorityName="Banks" authorityYear="1934" box="[151,396,1336,1361]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED38296EFF5EFAC84D02FAA7" box="[151,261,1338,1361]" italics="true" pageId="1" pageNumber="384">Minagenia</emphasis>
<bibRefCitation id="E14B4B44ED38296EFEC2FACA4D8BFAA7" author="Banks, N." box="[267,396,1336,1361]" pageId="1" pageNumber="384" pagination="1 - 117" refId="ref8911" refString="Banks, N. (1934) The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences, 69, 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
</taxonomicName>
, (as a subgenus of
<taxonomicName id="42DA4D36ED38296EFD85FACB4F4BFAA7" ID-CoL="6YLZ" authority="Kohl, 1884" authorityName="Kohl" authorityYear="1884" box="[588,844,1336,1361]" class="Insecta" family="Pompilidae" genus="Pseudagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED38296EFD85FACB4EC8FAA7" box="[588,719,1337,1361]" italics="true" pageId="1" pageNumber="384">Pseudagenia</emphasis>
<bibRefCitation id="E14B4B44ED38296EFD1CFACA4F4BFAA7" author="Kohl, F." box="[725,844,1336,1361]" pageId="1" pageNumber="384" pagination="33 - 58" refId="ref9565" refString="Kohl, F. (1884) Die gattungen der Pompiliden. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 34, 33 - 58. https: // doi. org / 10.5962 / bhl. part. 26501" type="journal article" year="1884">Kohl, 1884</bibRefCitation>
</taxonomicName>
): Proc. Amer. Acad. Arts. Sci., v. 69, pg. 40, 64.
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED38296EFF5EFAAA482DFA86" box="[151,1066,1367,1392]" pageId="1" pageNumber="384" type="materials_examined">
<paragraph id="856536B5ED38296EFF5EFAAA482DFA86" blockId="1.[151,1436,1336,1545]" box="[151,1066,1367,1392]" pageId="1" pageNumber="384">
<typeStatus id="5A618817ED38296EFF5EFAAA4CCEFA86" box="[151,201,1368,1392]" pageId="1" pageNumber="384">Type</typeStatus>
species:
<taxonomicName id="42DA4D36ED38296EFEEFFAAA4F29FA86" authority="Banks, 1934" authorityName="Banks" authorityYear="1934" box="[294,814,1367,1392]" class="Insecta" family="Pompilidae" genus="Pseudagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="brevicornis" subGenus="Minagenia">
<emphasis id="B7AEEAA7ED38296EFEEFFAAA4EA0FA86" box="[294,679,1368,1392]" italics="true" pageId="1" pageNumber="384">Pseudagenia (Minagenia) brevicornis</emphasis>
<bibRefCitation id="E14B4B44ED38296EFD64FAA54F29FA86" author="Banks, N." box="[685,814,1367,1392]" pageId="1" pageNumber="384" pagination="1 - 117" refId="ref8911" refString="Banks, N. (1934) The Psammocharidae of the Philippines. Proceedings of the American Academy of Arts and Sciences, 69, 1 - 117. https: // doi. org / 10.2307 / 20023025" type="journal article" year="1934">Banks, 1934</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED38296EFF5EFA844F3DFA79" box="[151,826,1398,1423]" pageId="1" pageNumber="384" type="reference_group">
<paragraph id="856536B5ED38296EFF5EFA844F3DFA79" blockId="1.[151,1436,1336,1545]" box="[151,826,1398,1423]" pageId="1" pageNumber="384">
<taxonomicName id="42DA4D36ED38296EFF5EFA844DA7FA79" authority="Banks, 1944" authorityName="Banks" authorityYear="1944" box="[151,416,1398,1423]" class="Insecta" family="Pompilidae" genus="Nannochilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED38296EFF5EFA844D10FA78" box="[151,279,1398,1422]" italics="true" pageId="1" pageNumber="384">Nannochilus</emphasis>
<bibRefCitation id="E14B4B44ED38296EFED7FA844DA7FA79" author="Banks, N." box="[286,416,1398,1423]" pageId="1" pageNumber="384" pagination="165 - 187" refId="ref8981" refString="Banks, N. (1944) Psammocharidae (spider-wasps), notes and descriptions. Bulletin of the Museum of Comparative Zoology, 94, 165 - 187." type="journal article" year="1944">Banks, 1944</bibRefCitation>
</taxonomicName>
: Bull. Mus. Comp. Zool., v. 94, pg. 171.
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED38296EFF5EFA6448FAFA5B" box="[151,1277,1428,1454]" pageId="1" pageNumber="384" type="materials_examined">
<paragraph id="856536B5ED38296EFF5EFA6448FAFA5B" blockId="1.[151,1436,1336,1545]" box="[151,1277,1428,1454]" pageId="1" pageNumber="384">
<typeStatus id="5A618817ED38296EFF5EFA644CCEFA58" box="[151,201,1430,1454]" pageId="1" pageNumber="384">Type</typeStatus>
species:
<taxonomicName id="42DA4D36ED38296EFEEFFA674E84FA5B" authority="Banks, 1910" authorityName="Banks" authorityYear="1910" box="[294,643,1428,1453]" class="Insecta" family="Pompilidae" genus="Pseudagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="externa">
<emphasis id="B7AEEAA7ED38296EFEEFFA674DFBFA5B" box="[294,508,1429,1453]" italics="true" pageId="1" pageNumber="384">Pseudagenia externa</emphasis>
<bibRefCitation id="E14B4B44ED38296EFDCBFA664E84FA5B" author="Banks, N." box="[514,643,1428,1453]" pageId="1" pageNumber="384" pagination="114 - 126" refId="ref8859" refString="Banks, N. (1910) New Species of Psammocharidae. Journal of the New York Entomological Society, 18, 114 - 126." type="journal article" year="1910">Banks, 1910</bibRefCitation>
</taxonomicName>
, by original designation. Synonymized by
<bibRefCitation id="E14B4B44ED38296EFBFDFA6448FFFA5B" author="Townes, H." box="[1076,1272,1428,1453]" pageId="1" pageNumber="384" pagination="226 - 237" refId="ref10058" refString="Townes, H. (1957) Nearctic wasps of the subfamilies Pepsinae and Ceropalinae. United States National Museum Bulletin, 209, 226 - 237. https: // doi. org / 10.5479 / si. 03629236.209.1" type="journal article" year="1957">Townes, 1957: 226</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED38296EFF5EFA464F31FA3A" box="[151,822,1459,1484]" pageId="1" pageNumber="384" type="reference_group">
<paragraph id="856536B5ED38296EFF5EFA464F31FA3A" blockId="1.[151,1436,1336,1545]" box="[151,822,1459,1484]" pageId="1" pageNumber="384">
<taxonomicName id="42DA4D36ED38296EFF5EFA464DAEFA3A" ID-CoL="3SV9" authority="Haupt, 1959" authorityName="Haupt" authorityYear="1959" box="[151,425,1459,1484]" class="Insecta" family="Pompilidae" genus="Compsagenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED38296EFF5EFA464D25FA3D" box="[151,290,1460,1483]" italics="true" pageId="1" pageNumber="384">Compsagenia</emphasis>
<bibRefCitation id="E14B4B44ED38296EFEE0FA464DAEFA3A" author="Haupt, H." box="[297,425,1459,1484]" pageId="1" pageNumber="384" pagination="5 - 74" refId="ref9386" refString="Haupt, H. (1959) Elemente einer systematischen Aufteilung der Macromerinae m. (Hymenoptera-Sphecoidea). Nova Aeta Leopoldina, 141 (21), 5 - 74." type="journal article" year="1959">Haupt, 1959</bibRefCitation>
</taxonomicName>
: Nova Acta
<taxonomicName id="42DA4D36ED38296EFDECFA414E9BFA3A" authorityName="Hering" authorityYear="1934" box="[549,668,1459,1484]" class="Insecta" family="Arctiidae" genus="Leopoldina" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="genus">Leopoldina</taxonomicName>
. v. 141, pg. 66.
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED38296EFF5EFA214D7EF9FF" pageId="1" pageNumber="384" type="materials_examined">
<paragraph id="856536B5ED38296EFF5EFA214D7EF9FF" blockId="1.[151,1436,1336,1545]" pageId="1" pageNumber="384">
<typeStatus id="5A618817ED38296EFF5EFA214CCEFA1D" box="[151,201,1491,1515]" pageId="1" pageNumber="384">Type</typeStatus>
species:
<taxonomicName id="42DA4D36ED38296EFEE5FA214E9BFA1D" authority="Haupt, 1959" authorityName="Haupt" authorityYear="1959" box="[300,668,1490,1515]" class="Insecta" family="Pompilidae" genus="Compsagenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="laevipes">
<emphasis id="B7AEEAA7ED38296EFEE5FA214E14FA1C" box="[300,531,1490,1514]" italics="true" pageId="1" pageNumber="384">Compsagenia laevipes</emphasis>
<bibRefCitation id="E14B4B44ED38296EFDD5FA214E9BFA1D" author="Haupt, H." box="[540,668,1490,1515]" pageId="1" pageNumber="384" pagination="5 - 74" refId="ref9386" refString="Haupt, H. (1959) Elemente einer systematischen Aufteilung der Macromerinae m. (Hymenoptera-Sphecoidea). Nova Aeta Leopoldina, 141 (21), 5 - 74." type="journal article" year="1959">Haupt, 1959</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="42DA4D36ED38296EFD0CFA204844FA1D" authority="Banks, 1946" authorityName="Banks" authorityYear="1946" box="[709,1091,1490,1515]" class="Insecta" family="Pompilidae" genus="Nannnochilus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B7AEEAA7ED38296EFD0CFA204FB2FA1C" box="[709,949,1490,1514]" italics="true" pageId="1" pageNumber="384">Nannnochilus obscurus</emphasis>
<bibRefCitation id="E14B4B44ED38296EFC76FA204844FA1D" author="Banks, N." box="[959,1091,1490,1515]" pageId="1" pageNumber="384" pagination="311 - 525" refId="ref9045" refString="Banks, N. (1946) Studies of South American Psammocharidae: Part 1. Bulletin of the Museum of Comparative Zoology, 96 (4), 311 - 525." type="journal article" year="1946">Banks, 1946</bibRefCitation>
</taxonomicName>
), by monotypy. Synonymized by
<bibRefCitation id="E14B4B44ED38296EFF0EFA004D73F9FF" author="Evans, H. E." box="[199,372,1521,1545]" pageId="1" pageNumber="384" pagination="212 - 226" refId="ref9253" refString="Evans, H. E. (1973) Studies on Neotropical Prompilidae (Hymenoptera). IX. The genera of Auplopodini. Psyche, 80 (3), 212 - 226. https: // doi. org / 10.1155 / 1973 / 25131" type="journal article" year="1973">Evans 1973: 214</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED38296DFF5EF9C54D19FF0D" lastPageId="2" lastPageNumber="385" pageId="1" pageNumber="384" type="diagnosis">
<paragraph id="856536B5ED38296EFF5EF9C54EC1F914" blockId="1.[151,1437,1591,2014]" pageId="1" pageNumber="384">
<emphasis id="B7AEEAA7ED38296EFF5EF9C54D15F9A4" bold="true" box="[151,274,1591,1618]" pageId="1" pageNumber="384">Diagnosis.</emphasis>
<taxonomicName id="42DA4D36ED38296EFED3F9CA4D94F9A7" authorityName="Banks" authorityYear="1934" box="[282,403,1592,1617]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED38296EFED3F9CA4D94F9A7" box="[282,403,1592,1617]" italics="true" pageId="1" pageNumber="384">Minagenia</emphasis>
</taxonomicName>
, a member of the subfamily
<taxonomicName id="42DA4D36ED38296EFD2BF9C54F4FF9A7" authorityName="Lepeletier de Saint Fargeau" authorityYear="1845" box="[738,840,1591,1617]" class="Insecta" family="Pompilidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="subFamily" subFamily="Pepsinae">Pepsinae</taxonomicName>
, can be differentiated from other genera of the tribe
<taxonomicName id="42DA4D36ED38296EFF5EF9A94CEEF983" authorityName="Lepeletier de Saint Fargeau" authorityYear="1845" box="[151,233,1627,1653]" class="Insecta" family="Pompilidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="tribe" tribe="Pepsini">Pepsini</taxonomicName>
by the following combination of characters: the eyes are large, with the width of the eye in lateral view greater than 2 × the width of the gena; and the fore wing
<emphasis id="B7AEEAA7ED38296EFCE7F9724F68F96C" box="[814,879,1664,1690]" italics="true" pageId="1" pageNumber="384">2rs-m</emphasis>
and
<emphasis id="B7AEEAA7ED38296EFC60F9724FECF96C" box="[937,1003,1664,1690]" italics="true" pageId="1" pageNumber="384">3rs-m</emphasis>
veins similar in curvature, with some exceptions, including the
<emphasis id="B7AEEAA7ED38296EFE7EF9564DFFF948" box="[439,504,1700,1726]" italics="true" pageId="1" pageNumber="384">3rs-m</emphasis>
either having a median outward pocket or the
<emphasis id="B7AEEAA7ED38296EFBCBF9564843F948" box="[1026,1092,1700,1726]" italics="true" pageId="1" pageNumber="384">3rs-m</emphasis>
being straight and angled posteriorly (as in
<taxonomicName id="42DA4D36ED38296EFEFCF93A4EBEF914" authority="Tsuneki, 1989" authorityName="Tsuneki" authorityYear="1989" box="[309,697,1735,1762]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis id="B7AEEAA7ED38296EFEFCF93A4E08F917" box="[309,527,1736,1761]" italics="true" pageId="1" pageNumber="384">Minagenia taiwana</emphasis>
<bibRefCitation id="E14B4B44ED38296EFDDEF9354EBEF914" author="Tsuneki, K." box="[535,697,1735,1762]" pageId="1" pageNumber="384" pagination="161 - 172" refId="ref10098" refString="Tsuneki, K. (1989) A study of the Pompilidae of Taiwan (Hymenoptera). Special Publications of the Japan Hymenopterists Association, 35, 161 - 172." type="journal article" year="1989">Tsuneki, 1989</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="856536B5ED38296DFF0EF9194D19FF0D" blockId="1.[151,1437,1591,2014]" lastBlockId="2.[151,1437,153,1115]" lastPageId="2" lastPageNumber="385" pageId="1" pageNumber="384">
Other identifying characters for
<taxonomicName id="42DA4D36ED38296EFDFFF91E4EA8F8F3" authorityName="Banks" authorityYear="1934" box="[566,687,1772,1797]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED38296EFDFFF91E4EA8F8F3" box="[566,687,1772,1797]" italics="true" pageId="1" pageNumber="384">Minagenia</emphasis>
</taxonomicName>
include: the clypeus width being greater than 2 × its height, with the sides angling slightly inward before the truncate or gently concave apical margin; the pronotum with a distinct anterior face; the fore wing
<emphasis id="B7AEEAA7ED38296EFE1EF8C64E19F8B8" box="[471,542,1844,1870]" italics="true" pageId="1" pageNumber="384">3m-cu</emphasis>
meeting the SMC3 at or beyond the middle, the
<emphasis id="B7AEEAA7ED38296EFB98F8C6486FF8BB" box="[1105,1128,1844,1869]" italics="true" pageId="1" pageNumber="384">M</emphasis>
reaching the wing margin; the hind wing
<emphasis id="B7AEEAA7ED38296EFEF1F8AA4D6DF887" box="[312,362,1880,1905]" italics="true" pageId="1" pageNumber="384">cu-a</emphasis>
ending before the juncture of
<emphasis id="B7AEEAA7ED38296EFD75F8AA4ED4F887" box="[700,723,1880,1905]" italics="true" pageId="1" pageNumber="384">M</emphasis>
with
<emphasis id="B7AEEAA7ED38296EFCDAF8AA4F42F884" box="[787,837,1880,1906]" italics="true" pageId="1" pageNumber="384">CuA</emphasis>
; the hind wing anal lobe equal to or less than half the length of
<emphasis id="B7AEEAA7ED38296EFECDF88E4D67F860" box="[260,352,1916,1942]" italics="true" pageId="1" pageNumber="384">M+CuA</emphasis>
; the hind wing basal set of hamuli ending at the separation of the costal vein from
<emphasis id="B7AEEAA7ED38296EFAC7F88E492FF860" box="[1294,1320,1916,1942]" italics="true" pageId="1" pageNumber="384">Sc</emphasis>
,
<emphasis id="B7AEEAA7ED38296EFAFEF88E494FF863" box="[1335,1352,1916,1941]" italics="true" pageId="1" pageNumber="384">R</emphasis>
and
<emphasis id="B7AEEAA7ED38296EFA49F88E499BF863" box="[1408,1436,1916,1941]" italics="true" pageId="1" pageNumber="384">Rs</emphasis>
veins (except in
<taxonomicName id="42DA4D36ED38296EFE9BF8524E2AF84F" authorityName="Tsuneki" authorityYear="1989" box="[338,557,1952,1977]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="384" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis id="B7AEEAA7ED38296EFE9BF8524E2AF84F" box="[338,557,1952,1977]" italics="true" pageId="1" pageNumber="384">Minagenia taiwana</emphasis>
</taxonomicName>
); and the hind tibia smooth, without row of integument serrations. Additional characters specific for the males include: the fore tarsal claws being asymmetrical, and at least the mid-leg tarsal claws are bifid. Additional characters specific for the females include: the fore tarsal claws being symmetrical, and all tarsal claws being bifid; S6 being laterally compressed, usually with a ventral ridge present; and the stinger being straight.
</paragraph>
</subSubSection>
<subSubSection id="CDC0653EED3B296DFF0EFEF74920FBAD" pageId="2" pageNumber="385" type="discussion">
<paragraph id="856536B5ED3B296DFF0EFEF748B6FE91" blockId="2.[151,1437,153,1115]" pageId="2" pageNumber="385">
<emphasis id="B7AEEAA7ED3B296DFF0EFEF74D3CFEE9" bold="true" box="[199,315,261,287]" pageId="2" pageNumber="385">Remarks.</emphasis>
The name was first used as a subgenus of
<taxonomicName id="42DA4D36ED3B296DFCDFFEF74FA0FEE9" authorityName="Kohl" authorityYear="1884" box="[790,935,261,287]" class="Insecta" family="Pompilidae" genus="Pseudagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFCDFFEF74FA0FEE9" box="[790,935,261,287]" italics="true" pageId="2" pageNumber="385">Pseudagenia</emphasis>
</taxonomicName>
by Banks in 1934 to describe two Philippine species. The name was then raised to generic level by
<bibRefCitation id="E14B4B44ED3B296DFD3FFEDB4F99FEB5" author="Townes, H." box="[758,926,296,323]" pageId="2" pageNumber="385" pagination="226 - 237" refId="ref10058" refString="Townes, H. (1957) Nearctic wasps of the subfamilies Pepsinae and Ceropalinae. United States National Museum Bulletin, 209, 226 - 237. https: // doi. org / 10.5479 / si. 03629236.209.1" type="journal article" year="1957">Townes (1957)</bibRefCitation>
and synonymized with the New World genus
<taxonomicName id="42DA4D36ED3B296DFF5EFEBF4DBCFE91" authority="Banks, 1944" authorityName="Banks" authorityYear="1944" box="[151,443,333,359]" class="Insecta" family="Pompilidae" genus="Nannochilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFF5EFEBF4D22FE91" box="[151,293,333,359]" italics="true" pageId="2" pageNumber="385">Nannochilus</emphasis>
<bibRefCitation id="E14B4B44ED3B296DFEE5FEBF4DBCFE91" author="Banks, N." box="[300,443,333,359]" pageId="2" pageNumber="385" pagination="165 - 187" refId="ref8981" refString="Banks, N. (1944) Psammocharidae (spider-wasps), notes and descriptions. Bulletin of the Museum of Comparative Zoology, 94, 165 - 187." type="journal article" year="1944">Banks, 1944</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="42DA4D36ED3B296DFE0EFEBC4E47FE91" authorityName="Banks" authorityYear="1934" box="[455,576,334,359]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFE0EFEBC4E47FE91" box="[455,576,334,359]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
was first used at the generic level by
<bibRefCitation id="E14B4B44ED3B296DFC2FFEBF48AAFE91" author="Dreisbach, R. R." box="[998,1197,332,359]" pageId="2" pageNumber="385" pagination="832 - 845" refId="ref9158" refString="Dreisbach, R. R. (1953) New species in the genera Dipogon Fox and Minagenia Banks (Hymenoptera: Psammocharidae) with keys to species and photomicrographs of genital parts. American Midland Naturalist, 49 (3), 832 - 845. https: // doi. org / 10.2307 / 2485211" type="journal article" year="1953">Dreisbach (1953)</bibRefCitation>
.
</paragraph>
<paragraph id="856536B5ED3B296DFF0EFE804E49FDC9" blockId="2.[151,1437,153,1115]" pageId="2" pageNumber="385">
<taxonomicName id="42DA4D36ED3B296DFF0EFE804D47FE7D" authorityName="Banks" authorityYear="1934" box="[199,320,370,395]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFF0EFE804D47FE7D" box="[199,320,370,395]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
can be found throughout the New World, Africa, and the Australasian region. In the New World, a revision is desperately needed, as there are many undescribed species throughout the region (unpub. data). A single species is described from
<collectingCountry id="FDCD7625ED3B296DFE0AFE4B4E4BFE25" box="[451,588,441,467]" name="Madagascar" pageId="2" pageNumber="385">Madagascar</collectingCountry>
, first placed into the genus
<taxonomicName id="42DA4D36ED3B296DFC46FE4848B5FE25" authority="Arnold, 1934" authorityName="Arnold" authorityYear="1934" box="[911,1202,441,467]" class="Insecta" family="Pompilidae" genus="Micragenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFC46FE484815FE25" box="[911,1042,442,467]" italics="true" pageId="2" pageNumber="385">Micragenia</emphasis>
<bibRefCitation id="E14B4B44ED3B296DFBD3FE4B48B5FE25" author="Arnold, G." box="[1050,1202,441,467]" pageId="2" pageNumber="385" pagination="1 - 9" refId="ref8828" refString="Arnold, G. (1934) On three new and remarkable genera of the family Psammocharidae. Occasional Papers of the Rhodesian Museum, 3, 1 - 9." type="journal article" year="1934">Arnold, 1934</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="E14B4B44ED3B296DFB28FE4B497CFE25" author="Banks, N." box="[1249,1403,441,467]" pageId="2" pageNumber="385" pagination="335 - 362" refId="ref8950" refString="Banks, N. (1940) Some Psammocharidae from Madagascar (Hymenoptera). Proceedings of the Academy of Natural Sciences of Philadelphia, 92, 335 - 362." type="journal article" year="1940">Banks (1940)</bibRefCitation>
as
<taxonomicName id="42DA4D36ED3B296DFF5EFE2C4E0AFE01" authority="Banks, 1940" authorityName="Banks" authorityYear="1940" box="[151,525,477,503]" class="Insecta" family="Pompilidae" genus="Micragenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B7AEEAA7ED3B296DFF5EFE2C4D71FE01" box="[151,374,478,503]" italics="true" pageId="2" pageNumber="385">Micragenia minima</emphasis>
<bibRefCitation id="E14B4B44ED3B296DFEB4FE2F4E0AFE01" author="Banks, N." box="[381,525,477,503]" pageId="2" pageNumber="385" pagination="335 - 362" refId="ref8950" refString="Banks, N. (1940) Some Psammocharidae from Madagascar (Hymenoptera). Proceedings of the Academy of Natural Sciences of Philadelphia, 92, 335 - 362." type="journal article" year="1940">Banks, 1940</bibRefCitation>
</taxonomicName>
; later,
<bibRefCitation id="E14B4B44ED3B296DFD90FE2F4F05FE01" author="Townes, H." box="[601,770,476,503]" pageId="2" pageNumber="385" pagination="226 - 237" refId="ref10058" refString="Townes, H. (1957) Nearctic wasps of the subfamilies Pepsinae and Ceropalinae. United States National Museum Bulletin, 209, 226 - 237. https: // doi. org / 10.5479 / si. 03629236.209.1" type="journal article" year="1957">Townes (1957)</bibRefCitation>
created a new combination for
<taxonomicName id="42DA4D36ED3B296DFBA0FE2C4CDBFDED" authority="(Banks, 1940)" baseAuthorityName="Banks" baseAuthorityYear="1940" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B7AEEAA7ED3B296DFBA0FE2C4939FE01" box="[1129,1342,478,503]" italics="true" pageId="2" pageNumber="385">Minagenia minima</emphasis>
(
<bibRefCitation id="E14B4B44ED3B296DFA87FE2F4CD4FDED" author="Banks, N." pageId="2" pageNumber="385" pagination="335 - 362" refId="ref8950" refString="Banks, N. (1940) Some Psammocharidae from Madagascar (Hymenoptera). Proceedings of the Academy of Natural Sciences of Philadelphia, 92, 335 - 362." type="journal article" year="1940">Banks, 1940</bibRefCitation>
)
</taxonomicName>
. No other
<taxonomicName id="42DA4D36ED3B296DFE84FDF04DC1FDED" authorityName="Banks" authorityYear="1934" box="[333,454,514,539]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFE84FDF04DC1FDED" box="[333,454,514,539]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
species are recorded in the literature for Africa, but there are several undescribed species from mainland Africa and
<collectingCountry id="FDCD7625ED3B296DFE09FDD74E4EFDC9" box="[448,585,549,575]" name="Madagascar" pageId="2" pageNumber="385">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="856536B5ED3B296DFF0EFDBB4890FC3D" blockId="2.[151,1437,153,1115]" pageId="2" pageNumber="385">
Several other genera closely resemble
<taxonomicName id="42DA4D36ED3B296DFD48FDB84EFDFD95" authorityName="Banks" authorityYear="1934" box="[641,762,586,611]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFD48FDB84EFDFD95" box="[641,762,586,611]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
. Species of
<taxonomicName id="42DA4D36ED3B296DFC40FDB84805FD95" authorityName="Banks" authorityYear="1934" box="[905,1026,586,611]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFC40FDB84805FD95" box="[905,1026,586,611]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="42DA4D36ED3B296DFAF5FDB84D5FFD71" authority="Kohl, 1884" authorityName="Kohl" authorityYear="1884" class="Insecta" family="Pompilidae" genus="Epipompilus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFAF5FDB84CCAFD71" italics="true" pageId="2" pageNumber="385">Epipompilus</emphasis>
<bibRefCitation id="E14B4B44ED3B296DFF1EFD9F4D5FFD71" author="Kohl, F." box="[215,344,621,647]" pageId="2" pageNumber="385" pagination="33 - 58" refId="ref9565" refString="Kohl, F. (1884) Die gattungen der Pompiliden. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 34, 33 - 58. https: // doi. org / 10.5962 / bhl. part. 26501" type="journal article" year="1884">Kohl, 1884</bibRefCitation>
</taxonomicName>
by the following characteristics: the eyes are large and protuberant; the integumental puncture size is variable, but dense with the intervals dull; the hind wing
<emphasis id="B7AEEAA7ED3B296DFCAAFD604F92FD5D" box="[867,917,658,683]" italics="true" pageId="2" pageNumber="385">cu-a</emphasis>
is long and often curved, the angle at which it leaves the medial vein is broad and smooth, not sharp; the males have bifid claws on the middle leg, and the antennae is filamentous with appressed setae; females lack setae on eyes, have all claws bifid, and have the SGP laterally compressed. Males of
<taxonomicName id="42DA4D36ED3B296DFE5EFD0F4F04FCE1" authority="Gussakovskij, 1932" authorityName="Gussakovskij" authorityYear="1932" box="[407,771,765,791]" class="Insecta" family="Pompilidae" genus="Eopompilus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFE5EFD0F4E1AFCE1" box="[407,541,765,791]" italics="true" pageId="2" pageNumber="385">Eopompilus</emphasis>
<bibRefCitation id="E14B4B44ED3B296DFDECFD0F4F04FCE1" author="Gussakovskij, V. V." box="[549,771,765,791]" pageId="2" pageNumber="385" pagination="1 - 66" refId="ref9300" refString="Gussakovskij, V. V. (1932) Verzeichnis der von Herrn D: r R. Malaise im Ussuri und Kamtschatka gesammelten aculeaten Hymenopteren. Arkiv Fur Zoologi, A, 24 (10), 1 - 66." type="journal article" year="1932">Gussakovskij, 1932</bibRefCitation>
</taxonomicName>
look similar to
<taxonomicName id="42DA4D36ED3B296DFC70FD0C4835FCE1" authorityName="Banks" authorityYear="1934" box="[953,1074,766,791]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFC70FD0C4835FCE1" box="[953,1074,766,791]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
males, except that the antennae are highly serrated, the hind leg tibial groove along the brush is obvious and mostly devoid of setae, S6 has a deep median emargination fringed with long setae, and T1 is dorsally white. Upon further examination, one species that was previously labeled as
<taxonomicName id="42DA4D36ED3B296DFE75FC984E32FC75" authorityName="Banks" authorityYear="1934" box="[444,565,874,899]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFE75FC984E32FC75" box="[444,565,874,899]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
from
<collectingCountry id="FDCD7625ED3B296DFDB1FC9B4ECDFC75" box="[632,714,873,899]" name="Taiwan" pageId="2" pageNumber="385">Taiwan</collectingCountry>
likely belongs in a new genus, perhaps, near
<taxonomicName id="42DA4D36ED3B296DFB0DFC9B494DFC75" authorityName="Gussakovskij" authorityYear="1932" box="[1220,1354,873,899]" class="Insecta" family="Pompilidae" genus="Eopompilus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFB0DFC9B494DFC75" box="[1220,1354,873,899]" italics="true" pageId="2" pageNumber="385">Eopompilus</emphasis>
</taxonomicName>
, which will be discussed in a later paper. An additional two species from
<collectingCountry id="FDCD7625ED3B296DFC4AFC7F4FD2FC51" box="[899,981,909,935]" name="Taiwan" pageId="2" pageNumber="385">Taiwan</collectingCountry>
are members of other genera, discussed here, further demonstrating that differences between these genera can be easily overlooked.
</paragraph>
<paragraph id="856536B5ED3B296DFF0EFC274920FBAD" blockId="2.[151,1437,153,1115]" pageId="2" pageNumber="385">
Males of
<taxonomicName id="42DA4D36ED3B296DFEF8FC244DADFC19" authorityName="Banks" authorityYear="1934" box="[305,426,982,1007]" class="Insecta" family="Pompilidae" genus="Minagenia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B7AEEAA7ED3B296DFEF8FC244DADFC19" box="[305,426,982,1007]" italics="true" pageId="2" pageNumber="385">Minagenia</emphasis>
</taxonomicName>
are generally distinguished by genitalic features, while females lack such diagnostic characters. Female head and antennal measurement ratios and placement of sensory receptors on the antennae tend to greatly aid in identification, but there are several exceptions. In general, females have less morphological variation than males, and in some instances, it is difficult to identify morphological characters to separate species.
</paragraph>
</subSubSection>
</treatment>
</document>