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<document id="1C81E31E04354DD21FED2AFEA2EEF565" ID-CLB-Dataset="37307" ID-DOI="10.5281/zenodo.247206" ID-GBIF-Dataset="cd615cd7-ecb8-4cc3-b074-5ddfab94649a" ID-Zenodo-Dep="247206" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484571130511" checkinUser="plazi" docAuthor="Ivan Löbl & Ryo Ogawa" docDate="2016" docId="0C70F70CD710FF82FF7BAEA4DCC746B5" docLanguage="en" docName="LinzerbiolBeitr.48.2.1339-1492.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Scaphisoma kodadai Löbl & Ogawa, 2016, nov.sp." docType="treatment" docVersion="12" lastPageNumber="1371" masterDocId="F0498F74D70FFFA2FFC9AB07DD0F4264" masterDocTitle="On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH" masterLastPageNumber="1492" masterPageNumber="1339" pageNumber="1370" updateTime="1698435770511" updateUser="ExternalLinkService">
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<mods:titleInfo id="D6836702C2D5E6AC53809B24DEC1BB4C">
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<mods:title id="EE4891502102DBF51B39E161A92D90B6">On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH</mods:title>
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</mods:titleInfo>
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<mods:name id="A08AE2C0DD6A5E61E45DAFE777655FFE" type="personal">
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<mods:roleTerm id="7BA9EDCAEC7BD9F711CF9FF39A7DB57D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0EFA9359D78E06D0C107D14ABF165879">Ivan Löbl</mods:namePart>
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</mods:name>
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<mods:name id="F7A791D2A9B79068BF48E2225190DA0F" type="personal">
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<mods:roleTerm id="138FA34BA67B1A5C64576BAEA496D139">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8EC87BD2F978AAE1354D72DA4F5800E9">Ryo Ogawa</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A1417A9AE6F873C52BE5265FC77897D5">text</mods:typeOfResource>
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<mods:title id="FD2C229A7BD5BA5C22876491096F2DB6">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="EA4554442D6483655D2A51A7CD12E361">
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<mods:date id="EE5000EAA1B2E3017A5696FCB40D3234">2016</mods:date>
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<mods:detail id="85F0CBE10324EDB2004E3752F8DFF45D" type="volume">
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<mods:number id="A67411749BDB9A4A91915ABECBA12163">48</mods:number>
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</mods:detail>
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<mods:detail id="C05BDB2DC309BF99D2207C47BDDBC291" type="issue">
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<mods:number id="3E8F732AF8C2B9A926A8EA6F7FE2F685">2</mods:number>
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<mods:start id="9BB65213952308AA3EC2842D3B61AB07">1339</mods:start>
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<mods:end id="0616962C18507D787B3391B9DF7F2207">1492</mods:end>
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<mods:classification id="D5A180F433B0C1433D94155ACC25F774">journal article</mods:classification>
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<mods:identifier id="6F7B0B8916CA8956E7A1E83357730C8F" type="CLB-Dataset">37307</mods:identifier>
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<mods:identifier id="9BC30926A0E91786DFF98BCD36B378FF" type="DOI">10.5281/zenodo.247206</mods:identifier>
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<mods:identifier id="E82A08DDAD80EB30F0871D8FDD7A052A" type="GBIF-Dataset">cd615cd7-ecb8-4cc3-b074-5ddfab94649a</mods:identifier>
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<mods:identifier id="41DCA5A7B414419A525511709475889B" type="Zenodo-Dep">247206</mods:identifier>
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<treatment id="0C70F70CD710FF82FF7BAEA4DCC746B5" ID-DOI="http://doi.org/10.5281/zenodo.4390669" ID-GBIF-Taxon="127598820" ID-Zenodo-Dep="4390669" LSID="urn:lsid:plazi:treatment:0C70F70CD710FF82FF7BAEA4DCC746B5" httpUri="http://treatment.plazi.org/id/0C70F70CD710FF82FF7BAEA4DCC746B5" lastPageId="32" lastPageNumber="1371" pageId="31" pageNumber="1370">
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<subSubSection id="CCC31591D710FFBDFF7BAEA4DFB747DD" box="[178,696,1443,1465]" pageId="31" pageNumber="1370" type="nomenclature">
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<paragraph id="8466461AD710FFBDFF7BAEA4DFB747DD" blockId="31.[178,1086,1443,1601]" box="[178,696,1443,1465]" pageId="31" pageNumber="1370">
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<heading id="DF2EF176D710FFBDFF7BAEA4DFB747DD" bold="true" box="[178,696,1443,1465]" fontSize="9" level="3" pageId="31" pageNumber="1370" reason="6">
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<emphasis id="B6AD9A08D710FFBDFF7BAEA4DFB747DD" bold="true" box="[178,696,1443,1465]" pageId="31" pageNumber="1370">
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5.2.7.9.
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<taxonomicName id="43D93D99D710FFBDFECCAEA4DCD347DD" ID-CoL="6Y5C5" box="[261,476,1443,1465]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1370" phylum="Arthropoda" rank="species" species="kodadai" status="sp. nov.">
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<emphasis id="B6AD9A08D710FFBDFECCAEA4DCD347DD" bold="true" box="[261,476,1443,1465]" italics="true" pageId="31" pageNumber="1370">Scaphisoma kodadai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AD9E2773D710FFBDFE2AAEA4DF2047DC" box="[483,559,1443,1464]" pageId="31" pageNumber="1370">nov.sp.</taxonomicNameLabel>
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(
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<figureCitation id="1CE25A9FD710FFBDFDF5AEA4DFBE47DD" box="[572,689,1443,1465]" captionStart="Figs 83 - 88" captionStartId="123.[178,218,1522,1544]" captionTargetBox="[197,1070,217,1512]" captionTargetId="figure@123.[178,1085,212,1512]" captionTargetPageId="123" captionText="Figs 83 - 88: Aedeagi of Scaphisoma in dorsal and lateral views; (83, 84), S. densepunctatum nov. sp., scale = 0.2 mm; (85, 86), S. hexamerum nov. sp., scale 0.1 mm; (87, 88), S. kodadai nov. sp., scale = 0.1 mm." httpUri="https://zenodo.org/record/247213/files/figure.png" pageId="31" pageNumber="1370">Figs 87, 88</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCC31591D710FFBDFF7BAEC4DC304479" pageId="31" pageNumber="1370" type="materials_examined">
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<paragraph id="8466461AD710FFBDFF7BAEC4DC304479" blockId="31.[178,1086,1443,1601]" pageId="31" pageNumber="1370">T y p e m a t e r i a l: Holotype 6: Philippines, Luzon: Lagunas, Mt. Banahaw above Kinabuhayan, 600-700m, trail to Crystalino, 24.XI.1995, leg. J. Kodada & B. Rigová (MHNG). Paratype: Luzon: Lagunas, Mt. Banahaw above Kinabuhayan, 600-700 m, 24.XI.1995, leg. I. Löbl, 1 ex. (MHNG).</paragraph>
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</subSubSection>
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<subSubSection id="CCC31591D710FF82FF7BAD2CDC674675" lastPageId="32" lastPageNumber="1371" pageId="31" pageNumber="1370" type="description">
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<paragraph id="8466461AD710FF82FF7BAD2CDF58415F" blockId="31.[178,1086,1443,1601]" lastBlockId="32.[178,1086,219,1233]" lastPageId="32" lastPageNumber="1371" pageId="31" pageNumber="1370">D e s c r i p t i o n: Length 1.35 mm, width 0.98 mm. Head and body brown, appendages lighter than body, yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 10/7: IV 12/6: V 35/8: VI 37/9: VII 45/14: VIII 40/8: IX 43/10: X 45/10: XI 60/15. Pronotum finely and densely punctate, punctures well visible at 30 times magnification, lateral contours evenly rounded, lateral margin carinae visible in dorsal view, lateral striae impunctate. Exposed tip of scutellum minute. Elytra with lateral margin carinae exposed in dorsal view, apical margins slightly rounded, inner apical angles not prominent, situated in level with outer angles; sutural margin not raised, sutural striae fairly deep, bent at base, not extending along basal margins, parallel from level of scutellar tip to apices; adsutural areas narrow, very finely punctate. Lateral striae appearing impunctate. Elytral disc with punctation fine and dense, coarser than pronotal punctation, puncture intervals mostly twice to three times as large as puncture diameters. Hind wings fully developed. Hypomera and mesanepisterna lacking obvious microsculpture. Mesepimera about as long as intervals to mesocoxae and about four times as long as wide. Metaventrite lacking obvious microsculpture, very finely punctate. Median part of metaventrite convex, lacking impressions; antecoxal puncture rows absent. Submesocoxal areas 0.03 mm, about as fourth of intervals to metacoxae, submesocoxal lines weakly convex, distinctly punctate. Metanepisternum flat, narrow, narrowed anteriad, inner margin arcuate, suture shallow. Tibiae straight. Abdomen with distinct strigulate microsculpture. Sternite 1 with submetacoxal areas about 0.03 mm, about as fifth of intervals to apical margin, submetacoxal lines weakly convex, with distinct marginal punctures.</paragraph>
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<paragraph id="8466461AD72FFF82FF7BA84CDC674675" blockId="32.[178,1086,219,1233]" pageId="32" pageNumber="1371">
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M a l e: Protarsomeres 1 to 3 weakly widened. Aedeagus (
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<figureCitation id="1CE25A9FD72FFF82FCE9A84CDE994105" box="[800,918,843,865]" captionStart="Figs 83 - 88" captionStartId="123.[178,218,1522,1544]" captionTargetBox="[197,1070,217,1512]" captionTargetId="figure@123.[178,1085,212,1512]" captionTargetPageId="123" captionText="Figs 83 - 88: Aedeagi of Scaphisoma in dorsal and lateral views; (83, 84), S. densepunctatum nov. sp., scale = 0.2 mm; (85, 86), S. hexamerum nov. sp., scale 0.1 mm; (87, 88), S. kodadai nov. sp., scale = 0.1 mm." httpUri="https://zenodo.org/record/247213/files/figure.png" pageId="32" pageNumber="1371">Figs 87, 88</figureCitation>
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) 0.38 mm long, symmetrical, moderately sclerotized. Apical process of median lobe almost as long as basal bulb, inflexed, with concave ventral side, narrowed in apical section, with blunt tip; articular processes small, not prominent. Parameres evenly narrow and almost straight in both, dorsal and lateral views, reaching beyond tip of median lobe. Internal sac without rods or flagellum, with dense spine-like structures, latter larger and more sclerotized along left margin.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCC31591D72FFF82FF7BAF27DCC746B5" pageId="32" pageNumber="1371" type="etymology">
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<paragraph id="8466461AD72FFF82FF7BAF27DCC746B5" blockId="32.[178,1086,219,1233]" pageId="32" pageNumber="1371">
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E t y m o l o g y: The species is named in honour of a friend of the senior author, Jan Kodada, Bratislava, Slovakia, who collected this and a number of other new species. Co mp ar a t i v e n o t e s: The aedeagal characters suggest relationships with
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<emphasis id="B6AD9A08D72FFF82FBE3AF64DC2446F1" italics="true" pageId="32" pageNumber="1371">S. binaluanum</emphasis>
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and its allied described above. It differs from these species by the sutural striae extended along basal margins of the elytra and by the much shorter antennomere IV. The paratype is teneral.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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