treatments-xml/data/EB/68/87/EB6887C0A92F173DFAA7F968FCD4FD3E.xml

<document id="D25CE71E359DB07620EE1EEB30708FEA" ID="10.11646/zootaxa.4263.2.2SLASH11000" ID-CLB-Dataset="33089" ID-DOI="10.11646/zootaxa.4263.2.2" ID-GBIF-Dataset="55b5ae73-6fd6-4c60-8230-ab3d60f5f163" ID-ISSN="1175-5326" ID-Zenodo-Dep="573070" ID-ZooBank="0E6B1EAE-5346-4CAE-ADA1-8CA75D9D7038" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494314943239" checkinUser="plazi" docAuthor="Liu, Yubin, Hutchings, Pat &amp; Sun, Shichun" docDate="2017" docId="EB6887C0A92F173DFAA7F968FCD4FD3E" docLanguage="en" docName="zootaxa.4263.2.2SLASH11000.pdf" docOrigin="Zootaxa 4263 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marphysa tribranchiata Liu, Hutchings &amp; Sun, 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="240" masterDocId="1751FFB8A9271731FA30FFCBFFD1FF96" masterDocTitle="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934" masterLastPageNumber="250" masterPageNumber="228" pageNumber="236" updateTime="1698439380213" updateUser="ExternalLinkService">
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<mods:title id="7330AA1E6B8873CA5C7470483560320A">Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934</mods:title>
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<mods:namePart id="E19A71E95F1D40C12D29038A49FB25C7">Liu, Yubin</mods:namePart>
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<mods:namePart id="9422A9E0EFD7AD8BA8A17DF1BE206490">Hutchings, Pat</mods:namePart>
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<mods:namePart id="D78D66BCCD23CE8172AAB24F01B0D42E">Sun, Shichun</mods:namePart>
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<mods:title id="9EDD0052A031E59BB7B7C6736D8FECBE">Zootaxa</mods:title>
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<mods:date id="169D70E76D3EA47E97783B5AA29664DE">2017</mods:date>
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<treatment id="EB6887C0A92F173DFAA7F968FCD4FD3E" ID-DOI="http://doi.org/10.5281/zenodo.6032320" ID-GBIF-Taxon="130497660" ID-Zenodo-Dep="6032320" LSID="urn:lsid:plazi:treatment:EB6887C0A92F173DFAA7F968FCD4FD3E" httpUri="http://treatment.plazi.org/id/EB6887C0A92F173DFAA7F968FCD4FD3E" lastPageId="12" lastPageNumber="240" pageId="8" pageNumber="236">
<subSubSection id="2BDB655DA92F1739FAA7F968FE72F949" pageId="8" pageNumber="236" type="nomenclature">
<paragraph id="637E36D6A92F1739FAA7F968FDD0F928" blockId="8.[151,513,1699,1759]" box="[151,513,1699,1726]" pageId="8" pageNumber="236">
<heading id="383681BAA92F1739FAA7F968FDD0F928" bold="true" box="[151,513,1699,1726]" fontSize="11" level="1" pageId="8" pageNumber="236" reason="1">
<emphasis id="51B5EAC4A92F1739FAA7F968FDD0F928" bold="true" box="[151,513,1699,1726]" pageId="8" pageNumber="236">
<taxonomicName id="A4C14D55A92F1739FAA7F968FE66F92B" ID-CoL="3Y8BP" box="[151,439,1699,1725]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="236" phylum="Annelida" rank="species" species="tribranchiata" status="sp. nov.">
<emphasis id="51B5EAC4A92F1739FAA7F968FE66F92B" bold="true" box="[151,439,1699,1725]" italics="true" pageId="8" pageNumber="236">Marphysa tribranchiata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A8657BFA92F1739FB8FF96FFDD0F928" box="[447,513,1700,1726]" pageId="8" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="637E36D6A92F1739FAA7F90DFE72F949" blockId="8.[151,513,1699,1759]" box="[151,419,1734,1759]" pageId="8" pageNumber="236">
<figureCitation id="FBFA2A53A92F1739FAA7F90DFED1F949" box="[151,256,1734,1759]" captionStart="FIGURE 1" captionStartId="3.[151,250,1342,1364]" captionTargetBox="[151,1434,269,1318]" captionTargetId="figure@3.[151,1436,267,1318]" captionTargetPageId="3" captionText="FIGURE 1. Type localities of Marphysa tripectinata n. sp., M. multipectinata n. sp., M. tribranchiata n. sp. and M. sinensis." httpUri="https://zenodo.org/record/573071/files/figure.png" pageId="8" pageNumber="236">Figures 1</figureCitation>
, 
<figureCitation id="FBFA2A53A92F1739FB3EF90CFECFF949" box="[270,286,1735,1759]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="8" pageNumber="236">4</figureCitation>
, 
<tableCitation id="2E43036DA92F1739FB1AF90DFE5BF949" box="[298,394,1734,1759]" captionStart="TABLE 2" captionStartId="2.[151,239,908,931]" captionText="TABLE 2. Types of pectinate chaetae present in Chinese species of Marphysa." pageId="8" pageNumber="236">Tables 2</tableCitation>
, 
<tableCitation id="2E43036DA92F1739FBA5F90CFE72F949" box="[405,419,1735,1759]" captionStart="TABLE 3" captionStartId="14.[151,235,152,174]" captionText="TABLE 3. Morphological characters of species of Marphysa occurring in Chinese waters and nearby belonging to Group B (after Fauchald 1970). (M. stragulum Grube, is not included as there is insufficient data provided in the original description)." pageId="8" pageNumber="236">3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="2BDB655DA92F1738FAA7F8C5FB05FDAB" lastPageId="9" lastPageNumber="237" pageId="8" pageNumber="236" type="materials_examined">
<paragraph id="637E36D6A92F1738FAA7F8C5FDD4FF43" blockId="8.[151,1437,1806,2011]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="237" pageId="8" pageNumber="236">
<emphasis id="51B5EAC4A92F1739FAA7F8C5FE54F8B1" bold="true" box="[151,389,1806,1831]" pageId="8" pageNumber="236">Material examined.</emphasis>
Holotype: ASIZW0000348-2, collected by Shi-Kuei Wu from Wanli, 
<date id="177F1016A92F1739FEF0F8C5FA90F8B1" box="[1216,1345,1806,1831]" pageId="8" pageNumber="236" value="1965-04-22">22.04.1965</date>
, (Long. 121.67°E; Lat. 25.2°N). Paratypes: ASIZW0000348-3, width on chaetiger 10, 
<quantity id="A4399B33A92F1739FE18F8F8FB55F8DC" box="[1064,1156,1843,1867]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="236" unit="mm" value="3.3">3.3 mm</quantity>
, length from anterior to posterior margin of chaetiger 10 is 10.0 mm, complete with 181 chaetigers, collected by Shi-Kuei Wu from Wanli, 
<date id="177F1016A92F1739FAA7F8B1FEC9F805" box="[151,280,1914,1939]" pageId="8" pageNumber="236" value="1965-04-22">22.04.1965</date>
, (121.67°E; 25.2°N); ASIZW0000346, width on chaetiger 10, 
<quantity id="A4399B33A92F1739F9E3F8B0FBFBF804" box="[979,1066,1915,1939]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="236" unit="mm" value="2.7">2.7 mm</quantity>
, length from anterior to posterior margin of chaetiger 10 is 
<quantity id="A4399B33A92F1739FBE6F854FDE5F820" box="[470,564,1951,1975]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="236" unit="mm" value="7.4">7.4 mm</quantity>
, complete with 161 chaetigers, collected by Shi-Kuei Wu from Shimen, 0 8.08.1964, (121.67°E; 25.2°N); ASIZW0000347-1, width on chaetiger 10, 
<quantity id="A4399B33A92F1739FE2DF808FBAAF84C" box="[1053,1147,1987,2011]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="236" unit="mm" value="3.4">3.4 mm</quantity>
, length from anterior to posterior margin of chaetiger 10 is 
<quantity id="A4399B33A92E1738F816FF53FDAEFF39" box="[550,639,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="9" pageNumber="237" unit="mm" value="7.4">7.4 mm</quantity>
, complete with 160 chaetigers, collected by Shi-Kuei Wu from Wanli, 0 6.08.1965, (121.67°E; 25.2°N).
</paragraph>
<paragraph id="637E36D6A92E1738FAF7FF14FB05FDAB" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="237">
<emphasis id="51B5EAC4A92E1738FAF7FF14FDE1FF6E" bold="true" box="[199,560,223,248]" pageId="9" pageNumber="237">Additional material examined</emphasis>
ASIZW0000354-2, width on chaetiger 10, 
<quantity id="A4399B33A92E1738FE13FF14FB5AFF61" box="[1059,1163,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="9" pageNumber="237" unit="mm" value="2.05">2.05 mm</quantity>
, length from anterior to posterior margin of chaetiger 10, 
<quantity id="A4399B33A92E1738F83CFECFFDB5FE8A" box="[524,612,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="9" pageNumber="237" unit="mm" value="5.1">5.1 mm</quantity>
, complete with 85 chaetigers, collected by Shi-Kuei Wu, (no locality data or date of collection given); 
<materialsCitation id="D3A93C8BA92E1738FBE8FEECFD0EFE1E" ID-GBIF-Occurrence="1500390012" collectionCode="ASIZW" collectorName="Shi-Kuei Wu" location="All" pageId="9" pageNumber="237" specimenCode="ASIZW0000354-4" specimenCount="1">
<specimenCode id="33679EADA92E1738FBE8FEECFD7CFED6" box="[472,685,295,320]" collectionCode="ASIZW" pageId="9" pageNumber="237">
<collectionCode id="05D0AE13A92E1738FBE8FEECFDE2FED6" box="[472,563,295,320]" pageId="9" pageNumber="237">ASIZW</collectionCode>
0000354-4
</specimenCode>
, width on chaetiger 10, 
<quantity id="A4399B33A92E1738F98DFEECFBF5FEA9" box="[957,1060,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" pageId="9" pageNumber="237" unit="mm" value="3.65">3.65 mm</quantity>
, length from anterior to posterior margin of chaetiger 10, 
<quantity id="A4399B33A92E1738FB94FE87FE2DFEF2" box="[420,508,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="237" unit="mm" value="9.5">9.5 mm</quantity>
, complete with 195 chaetigers, collected by 
<collectorName id="CE345300A92E1738F9DFFE87FB57FEF2" box="[1007,1158,332,357]" pageId="9" pageNumber="237">Shi-Kuei Wu</collectorName>
, (no locality data or date of collection given). 
<location id="661E600DA92E1738FBAFFEA4FE12FE1E" LSID="urn:lsid:plazi:treatment:EB6887C0A92F173DFAA7F968FCD4FD3E:661E600DA92E1738FBAFFEA4FE12FE1E" box="[415,451,367,392]" name="All" pageId="9" pageNumber="237">All</location>
posteriorly incomplete
</materialsCitation>
, ASIZW0000345-4; ASIZW0000347-3, ASIZW0000347-4 collected by Shi-Kuei Wu from Wanli, 0 6.08.1965, (121.67°E; 25.2°N); ASIZW0000348-4; collected by Shi-Kuei Wu from Wanli, 
<date id="177F1016A92E1738FB6CFE7CFE0CFE46" box="[348,477,439,464]" pageId="9" pageNumber="237" value="1965-04-22">22.04.1965</date>
, (121.67°E; 25.2°N); ASIZW0000354-1, ASIZW0000354-3, collected by Shi-Kuei Wu, (no locality data or date of collection given); ASIZW0000348-2, ASIZW0000348-3, ASIZW0000348-4 were collected by Shi-Kuei Wu from Wanli, 
<date id="177F1016A92E1738F864FE34FD05FD8E" box="[596,724,511,536]" pageId="9" pageNumber="237" value="1965-04-22">22.04.1965</date>
, (121.67°E; 25.2°N); 
<materialsCitation id="D3A93C8BA92E1738F9FCFE34FB05FDAB" ID-GBIF-Occurrence="1500390018" collectionCode="ASIZW" collectorName="Shi-Kuei Wu &amp; Shimen" country="Taiwan" latitude="25.2" location="All" longLatPrecision="5578" longitude="121.67" pageId="9" pageNumber="237" specimenCode="ASIZW0000345-4" specimenCount="1">
<specimenCode id="33679EADA92E1738F9FCFE34FB4FFD8E" box="[972,1182,511,536]" collectionCode="ASIZW" pageId="9" pageNumber="237">
<collectionCode id="05D0AE13A92E1738F9FCFE34FBF7FD8E" box="[972,1062,511,536]" pageId="9" pageNumber="237">ASIZW</collectionCode>
0000345-4
</specimenCode>
collected by 
<collectorName id="CE345300A92E1738FF07FE34FF6FFDAA" pageId="9" pageNumber="237">Shi-Kuei Wu</collectorName>
from 
<collectorName id="CE345300A92E1738FB33FDEFFE8AFDAB" box="[259,347,548,573]" pageId="9" pageNumber="237">Shimen</collectorName>
, 0 8.08.1964, (
<geoCoordinate id="06F55011A92E1738FBCAFDEFFDB8FDAB" box="[506,617,548,573]" direction="east" orientation="longitude" pageId="9" pageNumber="237" precision="555" value="121.67">121.67°E</geoCoordinate>
; 
<geoCoordinate id="06F55011A92E1738F844FDEFFD18FDAB" box="[628,713,548,573]" direction="north" orientation="latitude" pageId="9" pageNumber="237" precision="5555" value="25.2">25.2°N</geoCoordinate>
). 
<location id="661E600DA92E1738F8EBFDEFFD2EFDAB" LSID="urn:lsid:plazi:treatment:EB6887C0A92F173DFAA7F968FCD4FD3E:661E600DA92E1738F8EBFDEFFD2EFDAB" box="[731,767,548,573]" country="Taiwan" latitude="25.2" longLatPrecision="5578" longitude="121.67" name="All" pageId="9" pageNumber="237">All</location>
material collected from 
<collectingCountry id="1BD67646A92E1738FE22FDEFFBB5FDAB" box="[1042,1124,548,573]" name="Taiwan" pageId="9" pageNumber="237">Taiwan</collectingCountry>
of 
<collectingCountry id="1BD67646A92E1738FEBAFDEFFB00FDAB" box="[1162,1233,548,573]" name="China" pageId="9" pageNumber="237">China</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2BDB655DA92E1738FAF7FD8CFC6EF8DA" pageId="9" pageNumber="237" type="description">
<paragraph id="637E36D6A92E1738FAF7FD8CFE5DFD31" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="237">
<emphasis id="51B5EAC4A92E1738FAF7FD8CFE88FDF6" bold="true" box="[199,345,583,608]" pageId="9" pageNumber="237">Description.</emphasis>
<typeStatus id="BC7A8874A92E1738FB50FD8CFE19FDF6" box="[352,456,583,608]" pageId="9" pageNumber="237" type="holotype">Holotype</typeStatus>
complete with 215 chaetigers. Body iridescent. Preserved specimen colour red or dark pink. Maximum width at chaetiger 10, 
<quantity id="A4399B33A92E1738F864FDA7FD7CFD12" box="[596,685,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="237" unit="mm" value="3.9">3.9 mm</quantity>
, excluding parapodia. Length from anterior to posterior margin of chaetiger 10, 
<quantity id="A4399B33A92E1738FB1EFD5BFE56FD31" box="[302,391,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="237" unit="mm" value="8.7">8.7 mm</quantity>
.
</paragraph>
<paragraph id="637E36D6A92E1738FAF7FD7FFED3FC5E" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="237">
Prostomium longer than length of peristomium, margins of prostomium dorsally flattened with shallow median sulcus. Eyes absent. Antennae arranged with median one inserted more posteriorly to lateral ones. Length of antennae varies, median longer and thicker than lateral ones, longest antennae reaching to chaetiger 3 (
<figureCitation id="FBFA2A53A92E1738FF03FD37FAA8FC82" box="[1331,1401,764,789]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
A). Ceratophores smooth, ceratostyles digitiform with faint articulations. Palps smooth, shorter than antennae. Ratio of length between antennae and palps is 1.5x. Peristomial rings well demarcated all around. Mandibles with calcareous plates longer than sclerotized matrix (
<figureCitation id="FBFA2A53A92E1738F88CFCACFCD1FC16" box="[700,768,871,896]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
D). Maxillae I more than twice length of carrier; maxilla III arched with anterior teeth in line with posterior ones. Maxillary formula: I=1+1, II=4+4, III=5+0, IV=4+8, V=1+1 (
<figureCitation id="FBFA2A53A92E1738FAAFFC64FF32FC5E" box="[159,227,943,968]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
F).
</paragraph>
<paragraph id="637E36D6A92E1738FAF7FC1FFB80F9C6" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="237">
Parapodia vary along the body, prechaetal lobe small throughout, anterior parapodia with short chaetal lobe, postchaetal lobe longer, median with postchaetal lobe decreasing in size and becoming same length as chaetal lobe, posterior with chaetal lobe same length as chaetal lobe (
<figureCitation id="FBFA2A53A92E1738F928FBD7FC8FFBA2" box="[792,862,1052,1077]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
C). Dorsal cirri conical, shorter than chaetal lobes. Ventral cirri globular throughout body, the base of ventral cirri inconspicuous, length similar throughout body. Length of dorsal cirrus and ventral cirrus differs along body; in anterior and median chaetigers, dorsal cirri longer than ventral cirri, in posterior chaetigers equal in length. Branchiae begin with chaetiger 26, finish at chaetiger 181, pectinate throughout. Number of branchial filaments in anterior branchiae with 1–2 simple filaments, with maximum number of branchial filaments 3, in median parapodia (
<figureCitation id="FBFA2A53A92E1738F9BCFB04FC03FB7E" box="[908,978,1231,1256]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
B). Length of branchial filaments where best developed is up to 2–2.5x length of dorsal cirrus. Parapodia 3, 20, 55, 90, 125, 160, 195, examined under the light microscope. Aciculae brown, with maximum number of 
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anterior parapodia (
<figureCitation id="FBFA2A53A92E1738FE50FADCFB74FAA6" box="[1120,1189,1303,1328]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
C). Subacicular hooks brown, unidentate and bidentate, present from chaetiger 20 onwards. Maximum number of subacicular hooks 6 per parapodia. Compound spinigers present, throughout. Number of compound spinigers within a parapodium 10–31. Compound falcigers absent. Pectinate chaetae present throughout body. Number of pectinate chaetae within a parapodium 
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anterior and nearly 
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median and posterior ones. Pectinate chaetae 
<specimenCount id="75C7FD5FA92E1738FEB2FA63FB03FA57" box="[1154,1234,1448,1473]" pageId="9" pageNumber="237" type="generic" typeStatus="types">3 types</specimenCount>
present (
<tableCitation id="2E43036DA92E1738FF08FA6CFA5FFA56" box="[1336,1422,1447,1472]" captionStart="TABLE 2" captionStartId="2.[151,239,908,931]" captionText="TABLE 2. Types of pectinate chaetae present in Chinese species of Marphysa." pageId="9" pageNumber="237">Table 2</tableCitation>
), with both asymmetrical and symmetrical teeth, outer teeth diverse, in some chaetigers outer teeth longer than inner teeth (
<figureCitation id="FBFA2A53A92E1738FAEDFA24FEF0F99E" box="[221,289,1519,1544]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
G). In anterior parapodia only symmetrical isodont pectinate chaetae with about 12 teeth present (
<figureCitation id="FBFA2A53A92E1738FF5FFA24FF77F9BA" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
H-11). In posterior 
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present: symmetrical anodont pectinate chaetae with about 14 teeth (
<figureCitation id="FBFA2A53A92E1738FEE0F9DFFAC5F9BA" box="[1232,1300,1556,1581]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
G-(1), 4H-I) and isodont symmetrical pectinate chaetae with about 17 teeth (
<figureCitation id="FBFA2A53A92E1738F96FF9FCFC73F9C6" box="[863,930,1591,1616]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
G-(II), 4H–VI).
</paragraph>
<paragraph id="637E36D6A92E1738FAF7F997FF6BF90E" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="237">Limbate capillaries present, throughout body, all similar. Number of limbate chaetae within a parapodium 11– 24.</paragraph>
<paragraph id="637E36D6A92E1738FAF7F96FFC23F976" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="237">
Pygidium with 2 pairs of pygidial cirri (
<figureCitation id="FBFA2A53A92E1738F897F96FFD21F92A" box="[679,752,1700,1725]" captionStart="FIGURE 4" captionStartId="10.[151,250,1704,1726]" captionTargetBox="[284,1306,198,1683]" captionTargetId="figure@10.[276,1312,193,1689]" captionTargetPageId="10" captionText="FIGURE 4. Marphysa tribranchiata n. sp. and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8 – 12 teeth; III. symmetrical isodont with about 14 – 16 teeth; IV. asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14 – 18 teeth; VIII. asymmetrical anodont with 5 – 7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW 0000348 - 2. D, E. ASIZW 0000348 - 3. H. ASIZW 0000348 - 2, ASIZW 0000345 - 1, AM W. 49069. Scale bars: A, B. = 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale." httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="9" pageNumber="237">Fig. 4</figureCitation>
F), one pair longer than the other and inserted posteriorly laterally, shortest pair about 0.2x length of longer pair and inserted ventrally.
</paragraph>
<paragraph id="637E36D6A92E1738FAF7F927FC6EF8DA" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="237">
Variation. The 
<typeStatus id="BC7A8874A92E1738FB5FF926FE0BF893" box="[367,474,1773,1797]" pageId="9" pageNumber="237" type="paratype">paratypes</typeStatus>
exhibit the following variation, branchiae starting from chaetigers 16 to 26, and with 21–34 of the posterior-most chaetigers without any branchiae, the maximum number of filaments1–3. The ratio of the length between the antennae and the palps varies between 1.3 to 1.7.
</paragraph>
</subSubSection>
<subSubSection id="2BDB655DA92E173DFAF7F89CFE66FE63" lastPageId="12" lastPageNumber="240" pageId="9" pageNumber="237" type="discussion">
<paragraph id="637E36D6A92E173DFAF7F89CFB1CFED6" blockId="9.[151,1437,151,2013]" lastBlockId="12.[151,1437,151,680]" lastPageId="12" lastPageNumber="240" pageId="9" pageNumber="237">
<materialsCitation id="D3A93C8BA92E173DFAF7F89CFB85FEA9" ID-GBIF-Occurrence="1500390014" country="Taiwan" county="Also" lastPageId="12" lastPageNumber="240" location="Shihmen" municipality="Using" pageId="9" pageNumber="237" specimenCount="1" stateProvince="Taipei" typeStatus="holotype">
<emphasis id="51B5EAC4A92E1738FAF7F89CFEEAF8E6" bold="true" box="[199,315,1879,1904]" pageId="9" pageNumber="237">Remarks.</emphasis>
<taxonomicName id="A4C14D55A92E1738FB76F893FD88F8E6" box="[326,601,1880,1904]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="237" phylum="Annelida" rank="species" species="tribranchiata" status="sp. nov.">
<emphasis id="51B5EAC4A92E1738FB76F893FD88F8E6" box="[326,601,1880,1904]" italics="true" pageId="9" pageNumber="237">Marphysa tribranchiata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92E1738F855F89CFD78F8E6" bold="true" box="[613,681,1879,1904]" pageId="9" pageNumber="237">
<taxonomicNameLabel id="4A8657BFA92E1738F855F89CFD78F8E6" box="[613,681,1879,1904]" pageId="9" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is described as a new species because of the poorly developed branchiae with only 2–3 filaments, two 
<typeStatus id="BC7A8874A92E1738F861F8B6FD5DF803" box="[593,652,1917,1941]" pageId="9" pageNumber="237">types</typeStatus>
of symmetrical isodont pectinate chaetae with either 12 or 17 teeth, as well as symmetrical anodont chaetae with 14 teeth and the large number of subacicular hooks present which begin from around chaetiger 20. 
<collectingCounty id="8A1F4E5AA92E1738FBF1F80FFE27F84B" box="[449,502,1988,2013]" pageId="9" pageNumber="237">Also</collectingCounty>
the W10 ranges for the two species differs, with 
<taxonomicName id="A4C14D55A92E1738FE2CF80EFB35F84A" box="[1052,1252,1988,2012]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="237" phylum="Annelida" rank="species" species="multipectinata" status="sp. nov.">
<emphasis id="51B5EAC4A92E1738FE2CF80EFB35F84A" box="[1052,1252,1988,2012]" italics="true" pageId="9" pageNumber="237">M. multipectinata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92E1738FEDAF80FFAF8F84B" bold="true" box="[1258,1321,1988,2013]" pageId="9" pageNumber="237">
<taxonomicNameLabel id="4A8657BFA92E1738FEDAF80FFAF8F84B" box="[1258,1321,1988,2013]" pageId="9" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
always being larger (4.05–5.7) than 
<taxonomicName id="A4C14D55A92B173DFBEAFF52FD44FF26" box="[474,661,152,176]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="tribranchiata" status="sp. nov.">
<emphasis id="51B5EAC4A92B173DFBEAFF52FD44FF26" box="[474,661,152,176]" italics="true" pageId="12" pageNumber="240">M. tribranchiata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92B173DF8ADFF5CFD0DFF26" bold="true" box="[669,732,151,176]" pageId="12" pageNumber="240">
<taxonomicNameLabel id="4A8657BFA92B173DF8ADFF5CFD0DFF26" box="[669,732,151,176]" pageId="12" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(2.05-3.65). 
<collectingMunicipality id="831AACACA92B173DF941FF5CFC65FF26" box="[881,948,151,176]" pageId="12" pageNumber="240">Using</collectingMunicipality>
all these characters the new species can be distinguished from both 
<taxonomicName id="A4C14D55A92B173DFB98FF76FDBEFF42" box="[424,623,188,212]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="multipectinata" status="sp. nov.">
<emphasis id="51B5EAC4A92B173DFB98FF76FDBEFF42" box="[424,623,188,212]" italics="true" pageId="12" pageNumber="240">M. multipectinata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92B173DF845FF77FD65FF43" bold="true" box="[629,692,188,213]" pageId="12" pageNumber="240">
<taxonomicNameLabel id="4A8657BFA92B173DF845FF77FD65FF43" box="[629,692,188,213]" pageId="12" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="A4C14D55A92B173DF8DAFF76FC43FF42" box="[746,914,189,212]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="tripectinata" status="sp. nov.">
<emphasis id="51B5EAC4A92B173DF8DAFF76FC43FF42" box="[746,914,189,212]" italics="true" pageId="12" pageNumber="240">M. tripectinata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92B173DF9A8FF77FC07FF43" bold="true" box="[920,982,188,213]" pageId="12" pageNumber="240">
<taxonomicNameLabel id="4A8657BFA92B173DF9A8FF77FC07FF43" box="[920,982,188,213]" pageId="12" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
which occur in 
<collectingCountry id="1BD67646A92B173DFEB9FF77FB0AFF43" box="[1161,1243,188,213]" name="Taiwan" pageId="12" pageNumber="240">Taiwan</collectingCountry>
and on mainland 
<collectingCountry id="1BD67646A92B173DFAA7FF14FF0BFF6E" box="[151,218,223,248]" name="China" pageId="12" pageNumber="240">China</collectingCountry>
(see 
<tableCitation id="2E43036DA92B173DFB2DFF14FEA5FF6E" box="[285,372,223,248]" captionStart="TABLE 3" captionStartId="14.[151,235,152,174]" captionText="TABLE 3. Morphological characters of species of Marphysa occurring in Chinese waters and nearby belonging to Group B (after Fauchald 1970). (M. stragulum Grube, is not included as there is insufficient data provided in the original description)." pageId="12" pageNumber="240">Table 3</tableCitation>
). 
<taxonomicName id="A4C14D55A92B173DFBBEFF2BFD71FF6E" box="[398,672,224,248]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="tribranchiata" status="sp. nov.">
<emphasis id="51B5EAC4A92B173DFBBEFF2BFD71FF6E" box="[398,672,224,248]" italics="true" pageId="12" pageNumber="240">Marphysa tribranchiata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92B173DF89BFF14FD3CFF6E" bold="true" box="[683,749,223,248]" pageId="12" pageNumber="240">
<taxonomicNameLabel id="4A8657BFA92B173DF89BFF14FD3CFF6E" box="[683,749,223,248]" pageId="12" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
co-occurs with 
<taxonomicName id="A4C14D55A92B173DF99CFF2AFBB3FF6E" box="[940,1122,224,248]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="multipectina" status="sp. nov.">
<emphasis id="51B5EAC4A92B173DF99CFF2AFBB3FF6E" box="[940,1122,224,248]" italics="true" pageId="12" pageNumber="240">M. multipectina</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92B173DFE5CFF14FB7EFF6E" bold="true" box="[1132,1199,223,248]" pageId="12" pageNumber="240">
<taxonomicNameLabel id="4A8657BFA92B173DFE5CFF14FB7EFF6E" box="[1132,1199,223,248]" pageId="12" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
at 
<location id="661E600DA92B173DFEE9FF14FAEEFF6E" LSID="urn:lsid:plazi:treatment:EB6887C0A92F173DFAA7F968FCD4FD3E:661E600DA92B173DFEE9FF14FAEEFF6E" box="[1241,1343,223,248]" country="Taiwan" county="Also" municipality="Using" name="Shihmen" pageId="12" pageNumber="240" stateProvince="Taipei">Shihmen</location>
, 
<collectingRegion id="A105F834A92B173DFF7EFF14FA46FF6E" box="[1358,1431,223,248]" country="Taiwan" name="Taipei" pageId="12" pageNumber="240">Taipei</collectingRegion>
; however, unfortunately no habitat details are available. 
<bibRefCitation id="07504B27A92B173DF934FECFFC04FE8B" box="[772,981,260,285]" pageId="12" pageNumber="240" refString="Zanol, J., da Silva, T. dos S. C. &amp; Hutchings, P. (2016) Integrative taxonomy of Marphysa (Eunicidae, polychaete, Annelida) species of the S anguinea-group from Australia. Invertebrate Biology, 135 (4), 328 - 344. https: // doi. org / 10.1111 / ivb. 12146" type="journal article">
<location id="661E600DA92B173DF934FECFFC94FE8B" LSID="urn:lsid:plazi:treatment:EB6887C0A92F173DFAA7F968FCD4FD3E:661E600DA92B173DF934FECFFC94FE8B" box="[772,837,260,285]" country="Taiwan" county="Also" municipality="Using" name="Zanol" pageId="12" pageNumber="240" stateProvince="Taipei">Zanol</location>
<emphasis id="51B5EAC4A92B173DF97CFECEFC55FE8A" box="[844,900,260,284]" italics="true" pageId="12" pageNumber="240">et al.</emphasis>
(2016)
</bibRefCitation>
also found two species of 
<taxonomicName id="A4C14D55A92B173DFF30FECFFAA3FE8A" box="[1280,1394,260,284]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="genus">
<emphasis id="51B5EAC4A92B173DFF30FECFFAA3FE8A" box="[1280,1394,260,284]" italics="true" pageId="12" pageNumber="240">Marphysa</emphasis>
</taxonomicName>
cooccuring on the same mud flats at several locations along the 
<collectingRegion id="A105F834A92B173DF978FEE3FBC0FED6" box="[840,1041,295,320]" country="Australia" name="New South Wales" pageId="12" pageNumber="240">
New 
<collectingRegion id="A105F834A92B173DF9B5FEECFBC0FED6" box="[901,1041,295,320]" inferredBySuffix="true" pageId="12" pageNumber="240">
<collectingRegion id="A105F834A92B173DF9B5FEECFC17FED6" box="[901,966,295,320]" country="Tanzania" name="Kusini Unguja" pageId="12" pageNumber="240">South</collectingRegion>
Wales
</collectingRegion>
</collectingRegion>
coast
</materialsCitation>
, Australia.
</paragraph>
<caption id="37BE665EA92D173BFAA7F963FEECF867" httpUri="https://zenodo.org/record/573074/files/figure.png" pageId="10" pageNumber="238" targetBox="[284,1306,198,1683]" targetPageId="10">
<paragraph id="637E36D6A92D173BFAA7F963FEECF867" blockId="10.[151,1436,1704,2033]" pageId="10" pageNumber="238">
<emphasis id="51B5EAC4A92D173BFAA7F963FEC3F928" bold="true" box="[151,274,1704,1726]" pageId="10" pageNumber="238">FIGURE 4.</emphasis>
<taxonomicName id="A4C14D55A92D173BFB29F963FDDCF928" box="[281,525,1704,1726]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="10" pageNumber="238" phylum="Annelida" rank="species" species="tribranchiata" status="sp. nov.">
<emphasis id="51B5EAC4A92D173BFB29F963FDDCF928" box="[281,525,1704,1726]" italics="true" pageId="10" pageNumber="238">Marphysa tribranchiata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92D173BF824F963FD9CF928" bold="true" box="[532,589,1704,1726]" pageId="10" pageNumber="238">
<taxonomicNameLabel id="4A8657BFA92D173BF824F963FD9CF928" box="[532,589,1704,1726]" pageId="10" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and pectinate chaetae variation present in the three new species. Light photographs A. Anterior end of holotype, dorsal view. B. Branchiae at median region of holotype. C. Right median parapodium of holotype, lateral view. D. Mandible, ventral view. E. Maxillary apparatus, dorsal view. F. Pygidium of holotype, lateral view. G. 2 types of pectinate chaetae of holotype, parapodium 195: (I) symmetrical anodont pectinate chaetae with about 14 teeth; (II) symmetrical isodont pectinate chaetae with about 17 teeth. H. Schematic drawings of pectinate chaetae present in posterior parapodia of three new species. I. symmetrical anodont with parallel 12 teeth; II. symmetrical isodont with 8–12 teeth; III. symmetrical isodont with about 14–16 teeth; IV.asymmetrical anodont with about 4 teeth; V. asymmetrical anodont with about 14 teeth; VI. asymmetrical anodont with about 17 teeth; VII. asymmetrical anodont with 14–18 teeth; VIII. asymmetrical anodont with 5–7 teeth; IX. asymmetrical isodont with more than 30 teeth. A, B, C, F, G. ASIZW0000348-2. D, E. ASIZW0000348-3. H. ASIZW0000348-2, ASIZW0000345-1, AM W.49069. Scale bars: A, B.= 2 mm. C = 0.5 mm. D, E, F = 1 mm. G = 0.2 mm. H = is not to scale.
</paragraph>
</caption>
<caption id="37BE665EA92C173AFAA7F8F0FB56F87D" httpUri="https://zenodo.org/record/573075/files/figure.png" pageId="11" pageNumber="239" targetBox="[252,1317,201,1814]" targetPageId="11">
<paragraph id="637E36D6A92C173AFAA7F8F0FB56F87D" blockId="11.[151,1436,1851,2027]" pageId="11" pageNumber="239">
<emphasis id="51B5EAC4A92C173AFAA7F8F0FEC2F8C7" bold="true" box="[151,275,1851,1873]" pageId="11" pageNumber="239">FIGURE 5.</emphasis>
SEM images of 
<taxonomicName id="A4C14D55A92C173AFB8EF8F0FD73F8C7" box="[446,674,1851,1873]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="239" phylum="Annelida" rank="species" species="tripectinata" status="sp. nov.">
<emphasis id="51B5EAC4A92C173AFB8EF8F0FD73F8C7" box="[446,674,1851,1873]" italics="true" pageId="11" pageNumber="239">Marphysa tripectinata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92C173AF898F8F0FD30F8C7" bold="true" box="[680,737,1851,1873]" pageId="11" pageNumber="239">
<taxonomicNameLabel id="4A8657BFA92C173AF898F8F0FD30F8C7" box="[680,737,1851,1873]" pageId="11" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="A4C14D55A92C173AF923F8F0FBC2F8C7" box="[787,1043,1851,1873]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="239" phylum="Annelida" rank="species" species="multipectinata" status="sp. nov.">
<emphasis id="51B5EAC4A92C173AF923F8F0FBC2F8C7" box="[787,1043,1851,1873]" italics="true" pageId="11" pageNumber="239">Marphysa multipectinata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92C173AFE29F8F0FB82F8C7" bold="true" box="[1049,1107,1851,1873]" pageId="11" pageNumber="239">
<taxonomicNameLabel id="4A8657BFA92C173AFE29F8F0FB82F8C7" box="[1049,1107,1851,1873]" pageId="11" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="A4C14D55A92C173AFE69F8F0FAECF8C7" box="[1113,1341,1851,1873]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="239" phylum="Annelida" rank="species" species="tripectinata" status="sp. nov.">
<emphasis id="51B5EAC4A92C173AFE69F8F0FAECF8C7" box="[1113,1341,1851,1873]" italics="true" pageId="11" pageNumber="239">Marphysa tripectinata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92C173AFF73F8F0FAADF8C7" bold="true" box="[1347,1404,1851,1873]" pageId="11" pageNumber="239">
<taxonomicNameLabel id="4A8657BFA92C173AFF73F8F0FAADF8C7" box="[1347,1404,1851,1873]" pageId="11" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A. Compound spinigers chaetae, left parapodium 70. B. Symmetrical isodont pectinate chaeta, left parapodium 137. C. Two symmetrical anodont pectinate chaetae with parallel edge, left parapodium 204. 
<taxonomicName id="A4C14D55A92C173AF9C1F8B2FB24F819" box="[1009,1269,1913,1935]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="239" phylum="Annelida" rank="species" species="multipectinata" status="sp. nov.">
<emphasis id="51B5EAC4A92C173AF9C1F8B2FB24F819" box="[1009,1269,1913,1935]" italics="true" pageId="11" pageNumber="239">Marphysa multipectinata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92C173AFF30F8B2FAEFF819" bold="true" box="[1280,1342,1913,1935]" pageId="11" pageNumber="239">
<taxonomicNameLabel id="4A8657BFA92C173AFF30F8B2FAEFF819" box="[1280,1342,1913,1935]" pageId="11" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
D. Subacicular hook and compound spinigers chaetae, holotype, left parapodium 3. E. Two symmetrical isodont pectinate chaetae holotype, left parapodium 130. F. Symmetrical isodont pectinate chaeta with few coarse teeth, holotype, left parapodium 250. A–C AM W.49073. D–F ASIZW0000345-1. Scale bars: A, D = 50 µm. B = 5 µm. C, E, F = 10 µm.
</paragraph>
</caption>
<paragraph id="637E36D6A92B173DFAF7FE87FE66FE63" blockId="12.[151,1437,151,680]" pageId="12" pageNumber="240">
<taxonomicName id="A4C14D55A92B173DFAF7FE87FE06FEF2" box="[199,471,332,356]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="tribranchiata" status="sp. nov.">
<emphasis id="51B5EAC4A92B173DFAF7FE87FE06FEF2" box="[199,471,332,356]" italics="true" pageId="12" pageNumber="240">Marphysa tribranchiata</emphasis>
</taxonomicName>
<emphasis id="51B5EAC4A92B173DFBD1FE87FDF2FEF3" bold="true" box="[481,547,332,357]" pageId="12" pageNumber="240">
<taxonomicNameLabel id="4A8657BFA92B173DFBD1FE87FDF2FEF3" box="[481,547,332,357]" pageId="12" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from the other recorded species of 
<taxonomicName id="A4C14D55A92B173DFE88FE87FAFBFEF2" box="[1208,1322,332,356]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="genus">
<emphasis id="51B5EAC4A92B173DFE88FE87FAFBFEF2" box="[1208,1322,332,356]" italics="true" pageId="12" pageNumber="240">Marphysa</emphasis>
</taxonomicName>
from this region, including 
<taxonomicName id="A4C14D55A92B173DFB50FEBBFD97FE1E" box="[352,582,368,392]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="orientalis">
<emphasis id="51B5EAC4A92B173DFB50FEBBFD97FE1E" box="[352,582,368,392]" italics="true" pageId="12" pageNumber="240">Marphysa orientalis</emphasis>
</taxonomicName>
which lacks pectinate chaetae until chaetiger 67 although details regarding their dentition are not known and 
<taxonomicName id="A4C14D55A92B173DF82EFE5EFCEBFE3A" authority="Okuda, 1934" authorityName="Okuda" authorityYear="1934" box="[542,826,404,429]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="tamurai">
<emphasis id="51B5EAC4A92B173DF82EFE5EFD4FFE3A" box="[542,670,405,428]" italics="true" pageId="12" pageNumber="240">M. tamurai</emphasis>
Okuda, 1934
</taxonomicName>
which has pectinate chaetae from about chaetiger 20 and anterior ones with about 18 teeth, although again the numbers present along the body or 
<typeStatus id="BC7A8874A92B173DFEE1FE73FADDFE46" box="[1233,1292,440,464]" pageId="12" pageNumber="240">types</typeStatus>
of pectinate chaetae remain unknown.
</paragraph>
</subSubSection>
<subSubSection id="2BDB655DA92B173DFAF7FE34FE1AFDF6" pageId="12" pageNumber="240" type="etymology">
<paragraph id="637E36D6A92B173DFAF7FE34FF3EFDAA" blockId="12.[151,1437,151,680]" pageId="12" pageNumber="240">
<emphasis id="51B5EAC4A92B173DFAF7FE34FE9FFD8E" bold="true" box="[199,334,511,536]" pageId="12" pageNumber="240">Etymology.</emphasis>
The specific name 
<taxonomicName id="A4C14D55A92B173DF81CFDCBFD10FD8E" box="[556,705,512,536]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="240" phylum="Annelida" rank="species" species="tribranchiata">
<emphasis id="51B5EAC4A92B173DF81CFDCBFD10FD8E" box="[556,705,512,536]" italics="true" pageId="12" pageNumber="240">tribranchiata</emphasis>
</taxonomicName>
refers to the maximum number of branchial filaments which are present.
</paragraph>
<paragraph id="637E36D6A92B173DFAF7FD8CFE1AFDF6" blockId="12.[151,1437,151,680]" box="[199,459,583,608]" pageId="12" pageNumber="240">
<emphasis id="51B5EAC4A92B173DFAF7FD8CFEFBFDF6" bold="true" box="[199,298,583,608]" pageId="12" pageNumber="240">Habitat.</emphasis>
Not recorded.
</paragraph>
</subSubSection>
<subSubSection id="2BDB655DA92B173DFAF7FDA7FC30FD12" box="[199,993,620,645]" pageId="12" pageNumber="240" type="materials_examined">
<paragraph id="637E36D6A92B173DFAF7FDA7FC30FD12" blockId="12.[151,1437,151,680]" box="[199,993,620,645]" pageId="12" pageNumber="240">
<emphasis id="51B5EAC4A92B173DFAF7FDA7FEB4FD13" bold="true" box="[199,357,620,645]" pageId="12" pageNumber="240">
<typeStatus id="BC7A8874A92B173DFAF7FDA7FED3FD13" box="[199,258,620,645]" pageId="12" pageNumber="240">Type</typeStatus>
locality.
</emphasis>
Wanli, 
<collectingCountry id="1BD67646A92B173DFB8CFDA7FDDFFD13" box="[444,526,620,645]" name="Taiwan" pageId="12" pageNumber="240">Taiwan</collectingCountry>
of 
<collectingCountry id="1BD67646A92B173DF804FDA7FDABFD13" box="[564,634,620,645]" name="China" pageId="12" pageNumber="240">China</collectingCountry>
, (
<geoCoordinate id="06F55011A92B173DF8BCFDA7FD2AFD13" box="[652,763,620,645]" direction="east" orientation="longitude" pageId="12" pageNumber="240" precision="555" value="121.67">121.67°E</geoCoordinate>
; 
<geoCoordinate id="06F55011A92B173DF936FDA7FC8BFD13" box="[774,858,620,645]" direction="north" orientation="latitude" pageId="12" pageNumber="240" precision="5555" value="25.2">25.2°N</geoCoordinate>
), see 
<figureCitation id="FBFA2A53A92B173DF9A8FDA7FC0AFD12" box="[920,987,620,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1342,1364]" captionTargetBox="[151,1434,269,1318]" captionTargetId="figure@3.[151,1436,267,1318]" captionTargetPageId="3" captionText="FIGURE 1. Type localities of Marphysa tripectinata n. sp., M. multipectinata n. sp., M. tribranchiata n. sp. and M. sinensis." httpUri="https://zenodo.org/record/573071/files/figure.png" pageId="12" pageNumber="240">Fig. 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2BDB655DA92B173DFAF7FD44FCD4FD3E" box="[199,773,655,680]" pageId="12" pageNumber="240" type="distribution">
<paragraph id="637E36D6A92B173DFAF7FD44FCD4FD3E" blockId="12.[151,1437,151,680]" box="[199,773,655,680]" pageId="12" pageNumber="240">
<emphasis id="51B5EAC4A92B173DFAF7FD44FEB1FD3E" bold="true" box="[199,352,655,680]" pageId="12" pageNumber="240">Distribution.</emphasis>
Wanli and Shimen, 
<collectingCountry id="1BD67646A92B173DF873FD44FD44FD3E" box="[579,661,655,680]" name="Taiwan" pageId="12" pageNumber="240">Taiwan</collectingCountry>
of 
<collectingCountry id="1BD67646A92B173DF88BFD44FCD3FD3E" box="[699,770,655,680]" name="China" pageId="12" pageNumber="240">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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