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<document id="2B9A0584C8A9E8F7A83178D9F8CD8E78" ID-DOI="10.1111/j.1096-3642.2008.00415.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5446700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630461491835" checkinUser="felipe" docAuthor="Wu, Shu-Ping, Hwang, Chung-Chi &amp; Lin, Yao-Sung" docDate="2008" docId="E45287EFFFD4433AFCCEFF32867C5269" docLanguage="en" docName="j.1096-3642.2008.00415.x.pdf" docOrigin="Zoological Journal of the Linnean Society 154 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00415.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Satsuma auratibasis Wu &amp; Hwang &amp; Lin 2008, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="471" masterDocId="186BFF97FFF54318FF92FFF482155506" masterDocTitle="Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan" masterLastPageNumber="493" masterPageNumber="437" pageNumber="470" updateTime="1699279363994" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="CE8B001E54476B91925B5D7A70B53D4A">Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan</mods:title>
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<mods:namePart id="8463079B2216DD03D4D14871E35ABD65">Wu, Shu-Ping</mods:namePart>
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<mods:namePart id="D1931D8BE56DAE5DC48A31765EC286C2">Hwang, Chung-Chi</mods:namePart>
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<mods:namePart id="438CD704472B9CD63E35D715AE186974">Lin, Yao-Sung</mods:namePart>
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<mods:title id="4FEF69238586BFB24FC51A26E0DFB289">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="A25200A51B21F709737E1D2F377FA2B3">2008</mods:date>
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<mods:number id="B4101F64B45F9736CE0F3FC5FE5CFF90">2008-10-31</mods:number>
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<treatment id="E45287EFFFD4433AFCCEFF32867C5269" LSID="urn:lsid:plazi:treatment:E45287EFFFD4433AFCCEFF32867C5269" httpUri="http://treatment.plazi.org/id/E45287EFFFD4433AFCCEFF32867C5269" lastPageId="34" lastPageNumber="471" pageId="33" pageNumber="470">
<subSubSection id="24E16572FFD44339FCCEFF32879555DA" box="[860,1408,198,220]" pageId="33" pageNumber="470" type="nomenclature">
<paragraph id="6C4436F9FFD44339FCCEFF32879555DA" blockId="33.[860,1408,198,220]" box="[860,1408,198,220]" pageId="33" pageNumber="470">
<heading id="370C8195FFD44339FCCEFF32879555DA" box="[860,1408,198,220]" centered="true" fontSize="9" level="2" pageId="33" pageNumber="470" reason="2">
<taxonomicName id="ABFB4D7AFFD44339FCCEFF32866555DC" ID-CoL="6Y2P9" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[860,1136,198,219]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="470" phylum="Mollusca" rank="species" species="auratibasis" status="sp. nov.">
<emphasis id="5E8FEAEBFFD44339FCCEFF32866555DC" bold="true" box="[860,1136,198,219]" italics="true" pageId="33" pageNumber="470">SATSUMA AURATIBASIS</emphasis>
</taxonomicName>
<emphasis id="5E8FEAEBFFD44339FBE8FF3D86CA55DD" bold="true" box="[1146,1247,198,219]" pageId="33" pageNumber="470">
<taxonomicNameLabel id="45BC5790FFD44339FBE8FF3D86CA55DD" box="[1146,1247,198,219]" pageId="33" pageNumber="470" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4C02A7CFFD44339FB63FF32875955DA" box="[1265,1356,198,220]" captionStart="Figure 25" captionStartId="34.[144,223,1230,1249]" captionTargetBox="[370,1199,199,1200]" captionTargetId="figure-287@34.[367,1199,196,1200]" captionTargetPageId="34" captionText="Figure 25. Satsuma auratibasis sp. nov. A–D, holotype (TMMT 0609, shell height = 20.1 mm). E, F, paratype (TMMT 0636, shell height = 20.8 mm). G, H, paratype (TMMT 0637, shell height = 18.4 mm). I, living specimen." figureDoi="http://doi.org/10.5281/zenodo.5446754" httpUri="https://zenodo.org/record/5446754/files/figure.png" pageId="33" pageNumber="470">FIGS 25</figureCitation>
,
<figureCitation id="F4C02A7CFFD44339FAC9FF32876D55DA" box="[1371,1400,198,220]" captionStart="Figure 26" captionStartId="35.[164,243,1760,1779]" captionTargetBox="[167,1442,198,1729]" captionTargetId="figure-0@35.[164,1444,195,1730]" captionTargetPageId="35" captionText="Figure 26. Reproductive system of Satsuma auratibasis sp. nov. (paratype, TMMT 0658). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446756" httpUri="https://zenodo.org/record/5446756/files/figure.png" pageId="33" pageNumber="470">26</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="24E16572FFD4433AFCA8FF18867C5269" lastPageId="34" lastPageNumber="471" pageId="33" pageNumber="470" type="materials_examined">
<paragraph id="6C4436F9FFD44339FCA8FF1886075407" blockId="33.[826,1441,236,473]" box="[826,1042,236,257]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8FF1886075407" box="[826,1042,236,257]" italics="true" pageId="33" pageNumber="470">Material examined</emphasis>
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FEF8866454DF" blockId="33.[826,1441,236,473]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8FEF881F95427" box="[826,1004,268,289]" italics="true" pageId="33" pageNumber="470">Type specimen:</emphasis>
<materialsCitation id="DC933CA4FFD44339FC6FFEFF866454DF" collectionCode="TMMT, NMNS, BMNH, ANSP, SMF" pageId="33" pageNumber="470" specimenCode="TMMT 0609, TMMT 0658, TMMT 0636-0637, 0655-0657, NMNS 005405-3, BMNH 20060760-20060762, ANSP 413679-413681, SMF 329388-329389" specimenCount="2" typeStatus="holotype">
<typeStatus id="B340885BFFD44339FC6FFEFF86725427" box="[1021,1127,267,289]" pageId="33" pageNumber="470" type="holotype">Holotype</typeStatus>
:
<specimenCode id="3C5D9E82FFD44339FBE9FEFF87045427" box="[1147,1297,267,289]" collectionCode="TMMT" pageId="33" pageNumber="470">TMMT 0609</specimenCode>
(from type locality, dry shell, tissue in alcohol). Fifteen
<typeStatus id="B340885BFFD44339FABBFEDE87885439" box="[1321,1437,298,319]" pageId="33" pageNumber="470" type="paratype">paratypes</typeStatus>
: all from type locality,
<specimenCode id="3C5D9E82FFD44339FBD2FEBD86C75458" box="[1088,1234,328,350]" collectionCode="TMMT" pageId="33" pageNumber="470">TMMT 0658</specimenCode>
(dry shell, tissue in alcohol, dissected);
<specimenCode id="3C5D9E82FFD44339FBADFE938702547B" box="[1087,1303,359,381]" collectionCode="TMMT" pageId="33" pageNumber="470">TMMT 0636–0637</specimenCode>
,
<specimenCode id="3C5D9E82FFD44339FAB4FE9387B4547B" box="[1318,1441,359,381]" collectionCode="TMMT" pageId="33" pageNumber="470">0655–0657</specimenCode>
(dry shell, tissue in alcohol);
<specimenCode id="3C5D9E82FFD44339FB11FE728751549D" box="[1155,1348,389,411]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="33" pageNumber="470">NMNS 005405-3</specimenCode>
,
<specimenCode id="3C5D9E82FFD44339FADDFE72863354BC" collectionCode="BMNH" pageId="33" pageNumber="470">BMNH 20060760–20060762</specimenCode>
,
<specimenCode id="3C5D9E82FFD44339FBADFE50875A54BC" box="[1087,1359,420,442]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="33" pageNumber="470">ANSP 413679–413681</specimenCode>
,
<specimenCode id="3C5D9E82FFD44339FAF8FE5081F954DE" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="33" pageNumber="470">SMF 329388–329389</specimenCode>
(dry shell).
</materialsCitation>
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FDE381DE572A" blockId="33.[826,1441,535,618]" box="[826,971,535,556]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8FDE381DE572A" box="[826,971,535,556]" italics="true" pageId="33" pageNumber="470">
<typeStatus id="B340885BFFD44339FCA8FDE3817B572A" box="[826,878,535,556]" pageId="33" pageNumber="470">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FDC18189576C" blockId="33.[826,1441,535,618]" pageId="33" pageNumber="470">
<materialsCitation id="DC933CA4FFD44339FCA8FDC18182576C" country="Taiwan" location="Pingtung County" pageId="33" pageNumber="470" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
Mudan,
<collectingRegion id="AE3FF81BFFD44339FC32FDC1867F574C" box="[928,1130,565,587]" country="Taiwan" name="Pingtung" pageId="33" pageNumber="470">Pingtung County</collectingRegion>
, southern
<collectingCountry id="14EC7669FFD44339FB7DFDC18756574D" box="[1263,1347,565,587]" name="Taiwan" pageId="33" pageNumber="470">Taiwan</collectingCountry>
(
<figureCitation id="F4C02A7CFFD44339FACBFDC1878F574C" box="[1369,1434,565,587]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="33" pageNumber="470">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFD44339FCA8FDA08199576C" box="[826,908,596,618]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="33" pageNumber="470">Table 1</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FD5D81BF57B8" blockId="33.[826,1441,681,855]" box="[826,938,681,702]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8FD5D81BF57B8" box="[826,938,681,702]" italics="true" pageId="33" pageNumber="470">Diagnosis</emphasis>
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FD3386565651" blockId="33.[826,1441,681,855]" pageId="33" pageNumber="470">Spires and soft body yellowish without pedal stripe; base and sometimes the entire last whorl pale gold; number of whorls more than six; bursa stalk shorter than spermoviduct; base of bursa stalk expanded; principal pilaster long.</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FC6181A456AC" blockId="33.[826,1441,917,1000]" box="[826,945,917,938]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8FC6181A456AC" box="[826,945,917,938]" italics="true" pageId="33" pageNumber="470">Etymology</emphasis>
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FC4081DC56EE" blockId="33.[826,1441,917,1000]" pageId="33" pageNumber="470">
L.
<emphasis id="5E8FEAEBFFD44339FCCEFC40818E56CF" box="[860,923,948,969]" italics="true" pageId="33" pageNumber="470">aurat</emphasis>
: golden; L.
<emphasis id="5E8FEAEBFFD44339FBB8FC40867156CF" box="[1066,1124,948,969]" italics="true" pageId="33" pageNumber="470">basis</emphasis>
: base, referring to its pale golden base.
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FBD381AE513A" blockId="33.[826,1442,1063,1543]" box="[826,955,1063,1084]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8FBD381AE513A" box="[826,955,1063,1084]" italics="true" pageId="33" pageNumber="470">Description</emphasis>
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8FBB1868D5301" blockId="33.[826,1442,1063,1543]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8FBB18160515C" box="[826,885,1093,1114]" italics="true" pageId="33" pageNumber="470">Shell</emphasis>
(
<figureCitation id="F4C02A7CFFD44339FC18FBB181CC515D" box="[906,985,1093,1115]" captionStart="Figure 25" captionStartId="34.[144,223,1230,1249]" captionTargetBox="[370,1199,199,1200]" captionTargetId="figure-287@34.[367,1199,196,1200]" captionTargetPageId="34" captionText="Figure 25. Satsuma auratibasis sp. nov. A–D, holotype (TMMT 0609, shell height = 20.1 mm). E, F, paratype (TMMT 0636, shell height = 20.8 mm). G, H, paratype (TMMT 0637, shell height = 18.4 mm). I, living specimen." figureDoi="http://doi.org/10.5281/zenodo.5446754" httpUri="https://zenodo.org/record/5446754/files/figure.png" pageId="33" pageNumber="470">Fig. 25</figureCitation>
): Dextral, conical, thin, fragile, semitranslucent, medium sized. Apex obtuse. Whorls inflated to flat. Periphery roundly curved. Base inflated. Pre-apertural constriction behind peristome absent. Shell colour on upper whorls light yellow; base and sometimes the entire last whorl pale gold, turning white when periostracum peeled away. Shell surface smooth, with fine axial and spiral striae. Aperture diagonal, ovate-lunate to trapezoid. Inferior lip curved; outer lip almost straight. Junction between outer lip and inferior lip curved. Peristome thin, expanded. Columellar lip vertical to subvertical, reflected covering umbilicus. Umbilicus mostly covered by columellar lip, crevice-like. Junction between columellar lip and inferior lip roundly angulated.
</paragraph>
<paragraph id="6C4436F9FFD44339FCA8F9B1862B5269" blockId="33.[826,1441,1605,1903]" pageId="33" pageNumber="470">
<emphasis id="5E8FEAEBFFD44339FCA8F9B181CB535D" box="[826,990,1605,1627]" italics="true" pageId="33" pageNumber="470">Band or stain:</emphasis>
This taxon shows polymorphism. Some individuals do not exhibit any colour band or stain whilst others have a red-brown umbilicus and columellar lip; otherwise, the apex, inferior lip and outer lip lack such coloration. Some individuals have a short and thin band immediately behind the insertion of the outer lip. A rare colour pattern exists with the peripheral band and the apical and umbilicus spot. Band formula = 0 0 0 0 0 0 0 0; 1 0 0 0 0 0 7 0; 1 0 0 4 0 0 7 0 (rare).
</paragraph>
<caption id="38846671FFD7433AFF02FB3A86F951F9" ID-DOI="http://doi.org/10.5281/zenodo.5446754" ID-Zenodo-Dep="5446754" httpUri="https://zenodo.org/record/5446754/files/figure.png" pageId="34" pageNumber="471" startId="34.[144,223,1230,1249]" targetBox="[370,1199,199,1200]" targetPageId="34">
<paragraph id="6C4436F9FFD7433AFF02FB3A86F951F9" blockId="34.[144,1421,1230,1279]" pageId="34" pageNumber="471">
<emphasis id="5E8FEAEBFFD7433AFF02FB3A807151E4" bold="true" box="[144,612,1230,1250]" pageId="34" pageNumber="471">
Figure 25.
<taxonomicName id="ABFB4D7AFFD7433AFE83FB3B801351E7" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[273,518,1230,1250]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis" status="sp. nov.">
<emphasis id="5E8FEAEBFFD7433AFE83FB3B801351E7" bold="true" box="[273,518,1230,1250]" italics="true" pageId="34" pageNumber="471">Satsuma auratibasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFD7433AFD9FFB3B807151E4" box="[525,612,1231,1250]" pageId="34" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–D, holotype (TMMT 0609, shell height = 20.1 mm). E, F, paratype (TMMT 0636, shell height = 20.8 mm). G, H, paratype (TMMT 0637, shell height = 18.4 mm). I, living specimen.
</paragraph>
</caption>
<paragraph id="6C4436F9FFD7433AFF02FAC680C05318" blockId="34.[144,759,1330,1566]" pageId="34" pageNumber="471">
<emphasis id="5E8FEAEBFFD7433AFF02FAC68363504E" box="[144,374,1330,1352]" italics="true" pageId="34" pageNumber="471">Reproductive system</emphasis>
(
<figureCitation id="F4C02A7CFFD7433AFE17FAC683C1504E" box="[389,468,1330,1352]" captionStart="Figure 26" captionStartId="35.[164,243,1760,1779]" captionTargetBox="[167,1442,198,1729]" captionTargetId="figure-0@35.[164,1444,195,1730]" captionTargetPageId="35" captionText="Figure 26. Reproductive system of Satsuma auratibasis sp. nov. (paratype, TMMT 0658). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446756" httpUri="https://zenodo.org/record/5446756/files/figure.png" pageId="34" pageNumber="471">Fig. 26</figureCitation>
): The genital morphology is highly similar to that of
<taxonomicName id="ABFB4D7AFFD7433AFE48FAA5807D5060" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1909" box="[474,616,1361,1382]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="mollicula">
<emphasis id="5E8FEAEBFFD7433AFE48FAA5807D5060" box="[474,616,1361,1382]" italics="true" pageId="34" pageNumber="471">S. mollicula</emphasis>
</taxonomicName>
and can be distinguished by (1) the longer bursa stalk of
<taxonomicName id="ABFB4D7AFFD7433AFD4DFA84830450A5" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis">
<emphasis id="5E8FEAEBFFD7433AFD4DFA84830450A5" italics="true" pageId="34" pageNumber="471">S. auratibasis</emphasis>
</taxonomicName>
, which is three-fifths the length of the spermoviduct, than that of
<taxonomicName id="ABFB4D7AFFD7433AFE47FA59807750C7" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1909" box="[469,610,1452,1474]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="mollicula">
<emphasis id="5E8FEAEBFFD7433AFE47FA59807750C7" box="[469,610,1452,1474]" italics="true" pageId="34" pageNumber="471">S. mollicula</emphasis>
</taxonomicName>
; and (2) the longer epiphallus of
<taxonomicName id="ABFB4D7AFFD7433AFEE7FA3F800150E6" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[373,532,1483,1504]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis">
<emphasis id="5E8FEAEBFFD7433AFEE7FA3F800150E6" box="[373,532,1483,1504]" italics="true" pageId="34" pageNumber="471">S. auratibasis</emphasis>
</taxonomicName>
, which has a length almost equivalent to the penis. The length of epiphallus in
<taxonomicName id="ABFB4D7AFFD7433AFF4BF9FD8376531B" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1909" box="[217,355,1544,1566]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="mollicula">
<emphasis id="5E8FEAEBFFD7433AFF4BF9FD8376531B" box="[217,355,1544,1566]" italics="true" pageId="34" pageNumber="471">S. mollicula</emphasis>
</taxonomicName>
is three-fifths that of the penis.
</paragraph>
<paragraph id="6C4436F9FFD7433AFF02F9B1830E535C" blockId="34.[144,759,1605,1750]" box="[144,283,1605,1626]" pageId="34" pageNumber="471">
<emphasis id="5E8FEAEBFFD7433AFF02F9B1830E535C" box="[144,283,1605,1626]" italics="true" pageId="34" pageNumber="471">Distribution</emphasis>
</paragraph>
<paragraph id="6C4436F9FFD7433AFF02F99080F353D0" blockId="34.[144,759,1605,1750]" pageId="34" pageNumber="471">
<materialsCitation id="DC933CA4FFD7433AFF02F99080F553D0" country="Taiwan" county="Shouka" elevation="400" location="Hengchun" municipality="Shuangliu" pageId="34" pageNumber="471" specimenCount="1" stateProvince="Pingtung">
This species is distributed in lowland (&lt;
<quantity id="AB039B1CFFD7433AFDF7F99080B9537F" box="[613,684,1636,1657]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="471" unit="m" value="400.0">
<elevation id="E7D6D1CAFFD7433AFDF7F99080B9537F" box="[613,684,1636,1657]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="34" pageNumber="471" unit="m" value="400.0">400 m</elevation>
</quantity>
) broadleaf forest habitats around the north end of
<location id="69246022FFD7433AFD13F97780E3539F" LSID="urn:lsid:plazi:treatment:E45287EFFFD4433AFCCEFF32867C5269:69246022FFD7433AFD13F97780E3539F" box="[641,758,1667,1689]" country="Taiwan" county="Shouka" municipality="Shuangliu" name="Hengchun" pageId="34" pageNumber="471" stateProvince="Pingtung">Hengchun</location>
peninsula,
<collectingRegion id="AE3FF81BFFD7433AFE9FF95583C553B0" box="[269,464,1697,1719]" country="Taiwan" name="Pingtung" pageId="34" pageNumber="471">Pingtung County</collectingRegion>
, southern
<collectingCountry id="14EC7669FFD7433AFDD4F955808853B1" box="[582,669,1697,1719]" name="Taiwan" pageId="34" pageNumber="471">Taiwan</collectingCountry>
, including
<location id="69246022FFD7433AFF2EF934830553D0" LSID="urn:lsid:plazi:treatment:E45287EFFFD4433AFCCEFF32867C5269:69246022FFD7433AFF2EF934830553D0" box="[188,272,1728,1750]" country="Taiwan" county="Shouka" municipality="Shuangliu" name="Mudan" pageId="34" pageNumber="471" stateProvince="Pingtung">Mudan</location>
,
<collectingMunicipality id="8C20AC83FFD7433AFE8EF934838753D0" box="[284,402,1728,1750]" pageId="34" pageNumber="471">Shuangliu</collectingMunicipality>
and
<collectingCounty id="85254E75FFD7433AFE5FF934803653D0" box="[461,547,1728,1750]" pageId="34" pageNumber="471">Shouka</collectingCounty>
(
<figureCitation id="F4C02A7CFFD7433AFDA6F934806053D3" box="[564,629,1728,1750]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="34" pageNumber="471">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFD7433AFD16F93480C353D0" box="[644,726,1728,1750]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="34" pageNumber="471">Table 1</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="6C4436F9FFD7433AFF02F90982E05214" blockId="34.[144,759,1789,1903]" box="[144,245,1789,1810]" pageId="34" pageNumber="471">
<emphasis id="5E8FEAEBFFD7433AFF02F90982E05214" box="[144,245,1789,1810]" italics="true" pageId="34" pageNumber="471">Remarks</emphasis>
</paragraph>
<paragraph id="6C4436F9FFD7433AFF02F8E880735269" blockId="34.[144,759,1789,1903]" pageId="34" pageNumber="471">
The animals were found from
<quantity id="AB039B1CFFD7433AFE77F8E8801A5237" box="[485,527,1820,1841]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="34" pageNumber="471" unit="m" value="2.0">2 m</quantity>
above ground to the tree canopy. Adults were found in spring and usually perched higher than juveniles (
<figureCitation id="F4C02A7CFFD7433AFE6DF8AD804F5269" box="[511,602,1881,1903]" captionStart="Figure 25" captionStartId="34.[144,223,1230,1249]" captionTargetBox="[370,1199,199,1200]" captionTargetId="figure-287@34.[367,1199,196,1200]" captionTargetPageId="34" captionText="Figure 25. Satsuma auratibasis sp. nov. A–D, holotype (TMMT 0609, shell height = 20.1 mm). E, F, paratype (TMMT 0636, shell height = 20.8 mm). G, H, paratype (TMMT 0637, shell height = 18.4 mm). I, living specimen." figureDoi="http://doi.org/10.5281/zenodo.5446754" httpUri="https://zenodo.org/record/5446754/files/figure.png" pageId="34" pageNumber="471">Fig. 25I</figureCitation>
).
</paragraph>
<paragraph id="6C4436F9FFD7433AFCADFAC6867C5269" blockId="34.[806,1422,1330,1903]" pageId="34" pageNumber="471">
<taxonomicName id="ABFB4D7AFFD7433AFCADFAC6862C5041" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[831,1081,1330,1351]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis" status="sp. nov.">
<emphasis id="5E8FEAEBFFD7433AFCADFAC6862C5041" box="[831,1081,1330,1351]" italics="true" pageId="34" pageNumber="471">Satsuma auratibasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFD7433AFBDEFAC686BB5041" box="[1100,1198,1330,1351]" pageId="34" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="ABFB4D7AFFD7433AFAE4FAC681845060" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1909" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="mollicula">
<emphasis id="5E8FEAEBFFD7433AFAE4FAC681845060" italics="true" pageId="34" pageNumber="471">S. mollicula</emphasis>
</taxonomicName>
based on the morphological characters of shell and genitalia. Both were found distributed on Hengchun peninsula, southern
<collectingCountry id="14EC7669FFD7433AFB66FA7A875E50A2" box="[1268,1355,1422,1444]" name="Taiwan" pageId="34" pageNumber="471">Taiwan</collectingCountry>
, but their distributions do not overlap. The former species is distributed in the northern part of the area, the latter in the south (
<figureCitation id="F4C02A7CFFD7433AFAAFFA1E87685306" box="[1341,1405,1514,1536]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="34" pageNumber="471">Fig. 1</figureCitation>
). Furthermore, molecular phylogeny indicates
<taxonomicName id="ABFB4D7AFFD7433AFAE4F9FD81B2533A" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis">
<emphasis id="5E8FEAEBFFD7433AFAE4F9FD81B2533A" italics="true" pageId="34" pageNumber="471">S. auratibasis</emphasis>
</taxonomicName>
is close to
<taxonomicName id="ABFB4D7AFFD7433AFBDDF9D386D7533A" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[1103,1218,1575,1596]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="huberi">
<emphasis id="5E8FEAEBFFD7433AFBDDF9D386D7533A" box="[1103,1218,1575,1596]" italics="true" pageId="34" pageNumber="471">S. huberi</emphasis>
</taxonomicName>
rather than
<taxonomicName id="ABFB4D7AFFD7433AFAE4F9D38184535C" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1909" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="mollicula">
<emphasis id="5E8FEAEBFFD7433AFAE4F9D38184535C" italics="true" pageId="34" pageNumber="471">S. mollicula</emphasis>
</taxonomicName>
despite the morphological similarities.
<taxonomicName id="ABFB4D7AFFD7433AFCB4F990860B537F" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[806,1054,1636,1657]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis">
<emphasis id="5E8FEAEBFFD7433AFCB4F990860B537F" box="[806,1054,1636,1657]" italics="true" pageId="34" pageNumber="471">Satsuma auratibasis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="ABFB4D7AFFD7433AFB42F9908771537F" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1909" box="[1232,1380,1636,1657]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="mollicula">
<emphasis id="5E8FEAEBFFD7433AFB42F9908771537F" box="[1232,1380,1636,1657]" italics="true" pageId="34" pageNumber="471">S. mollicula</emphasis>
</taxonomicName>
in that its shell is larger, angulated at the periphery that is never keeled, pale gold in colour, never greenish gold as seen in
<emphasis id="5E8FEAEBFFD7433AFBDDF93486F653D3" box="[1103,1251,1728,1749]" italics="true" pageId="34" pageNumber="471">
<taxonomicName id="ABFB4D7AFFD7433AFBDDF93486CA53D3" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1909" box="[1103,1247,1728,1749]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="mollicula">S. mollicula</taxonomicName>
.
</emphasis>
Moreover, the golden tint is confined to the base or the last whorl only of
<taxonomicName id="ABFB4D7AFFD7433AFCECF90986355214" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[894,1056,1789,1810]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis">
<emphasis id="5E8FEAEBFFD7433AFCECF909819B5214" box="[894,910,1789,1810]" italics="true" pageId="34" pageNumber="471">S</emphasis>
.
<emphasis id="5E8FEAEBFFD7433AFC0DF90986355214" box="[927,1056,1789,1810]" italics="true" pageId="34" pageNumber="471">auratibasis</emphasis>
</taxonomicName>
.
<taxonomicName id="ABFB4D7AFFD7433AFBA3F90986C65214" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[1073,1235,1789,1810]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="auratibasis">
<emphasis id="5E8FEAEBFFD7433AFBA3F90986545214" box="[1073,1089,1789,1810]" italics="true" pageId="34" pageNumber="471">S</emphasis>
.
<emphasis id="5E8FEAEBFFD7433AFBC0F90986C65214" box="[1106,1235,1789,1810]" italics="true" pageId="34" pageNumber="471">auratibasis</emphasis>
</taxonomicName>
differs morphologically from
<taxonomicName id="ABFB4D7AFFD7433AFC46F8E886575237" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[980,1090,1820,1841]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="471" phylum="Mollusca" rank="species" species="huberi">
<emphasis id="5E8FEAEBFFD7433AFC46F8E886575237" box="[980,1090,1820,1841]" italics="true" pageId="34" pageNumber="471">S. huberi</emphasis>
</taxonomicName>
in having gold-tinted shell base, conical base of bursa stalk, long bursa stalk and long principal pilaster.
</paragraph>
</subSubSection>
</treatment>
</document>
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