244 lines
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244 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.732.21677" ID-GBIF-Dataset="05db50e8-b4e5-422a-b78f-9676d74ac319" ID-PMC="PMC5799741" ID-Pensoft-Pub="1313-2970-732-1" ID-PubMed="29416404" ID-ZBK="9995702B61464BA1BB5323DC9BA9650F" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-732-1" ModsDocOrigin="ZooKeys 732" ModsDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" checkinTime="1516831538731" checkinUser="pensoft" docAuthor="Mattia, Willy De, Neiber, Marco T. & Groh, Klaus" docDate="2018" docId="4CFDB3619791C6B943319DD1FFECCDE8" docLanguage="en" docName="ZooKeys 732: 1-125" docOrigin="ZooKeys 732" docSource="http://dx.doi.org/10.3897/zookeys.732.21677" docTitle="Wollastonia subcarinulata , comb. n., stat. n." docType="treatment" docVersion="5" lastPageNumber="61" masterDocId="FFA3FF812B32FFA8FFBAFFCDFF9AFF92" masterDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" masterLastPageNumber="125" masterPageNumber="1" pageNumber="59" updateTime="1668165370779" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mattia, Willy De</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neiber, Marco T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Groh, Klaus</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>732</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>125</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.732.21677</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.732.21677</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-732-1</mods:identifier>
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<mods:identifier type="ZBK">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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<mods:identifier type="ZooBank">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="140417771" LSID="urn:lsid:plazi:treatment:4CFDB3619791C6B943319DD1FFECCDE8" httpUri="http://treatment.plazi.org/id/4CFDB3619791C6B943319DD1FFECCDE8" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59">
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<subSubSection pageId="58" pageNumber="59" type="nomenclature">
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<paragraph pageId="58" pageNumber="59">
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†
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<taxonomicName authority="(Wollaston, 1878), comb. n." class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia subcarinulata" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="subcarinulata">Wollastonia subcarinulata (Wollaston, 1878), comb. n.</taxonomicName>
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<taxonomicNameLabel pageId="58" pageNumber="59">stat. n.</taxonomicNameLabel>
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Figs 196-198, 199
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</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="list of synonyms">
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<paragraph pageId="58" pageNumber="59">List of synonyms.</paragraph>
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<paragraph pageId="58" pageNumber="59">
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1878
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Helix (Hystricella) oxytropis var. subcarinulata" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="variety" species="oxytropis" subGenus="Hystricella" variety="subcarinulata">
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Helix (Hystricella) oxytropis var.
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<normalizedToken originalValue="ß">ss</normalizedToken>
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subcarinulata
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</taxonomicName>
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Wollaston: 168.
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</paragraph>
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<paragraph pageId="58" pageNumber="59">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Discula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discula (Hystricella) oxytropis" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="subSpecies" species="oxytropis" subGenus="Hystricella" subSpecies="subcarinulata">Discula (Hystricella) oxytropis subcarinulata</taxonomicName>
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-
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<normalizedToken originalValue="Waldén">Walden</normalizedToken>
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: 267.
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</paragraph>
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<paragraph pageId="58" pageNumber="59">
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2002
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Geomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geomitra oxytropis" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="subSpecies" species="oxytropis" subSpecies="subcarinulata">Geomitra oxytropis subcarinulata</taxonomicName>
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- Bank et al.: 124.
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</paragraph>
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<paragraph pageId="58" pageNumber="59">
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2006
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Discula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discula oxytropis" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="oxytropis">Discula oxytropis</taxonomicName>
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- Cameron et al.: 49 (1): 40 [partim].
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</paragraph>
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<paragraph pageId="58" pageNumber="59">
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2009
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella oxytropis" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="subSpecies" species="oxytropis" subSpecies="subcarinulata">Hystricella oxytropis subcarinulata</taxonomicName>
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- Groh et al.: 21, fig. 27.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="type material">
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<paragraph pageId="58" pageNumber="59">Type material.</paragraph>
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<paragraph pageId="58" pageNumber="59">NMWC 80.202, Acc 55.158, lectotype (herewith designated), from loc. typ., ex coll. Melvill-Tomlin, ex coll. T. V. Wollaston, Porto Santo, Madeira (Fig. 196).</paragraph>
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<caption pageId="58" pageNumber="59">
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<paragraph pageId="58" pageNumber="59">
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Figures 196-198. Shells of
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia subcarinulata" order="Stylommatophora" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="subcarinulata">Wollastonia subcarinulata</taxonomicName>
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. 196 lectotype, NMWC 80.202, Acc 55.158; 197-198 specimens from Barbinha, Quaternary aeolinites. Scale bar 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="materials_examined">
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<paragraph pageId="58" pageNumber="59">Further material examined.</paragraph>
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<paragraph pageId="58" pageNumber="59">
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All from Porto Santo, CKG/2, Barbinha, Quaternary aeolinites, layer 3,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.067776">33°04'04"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.296944">16°17'49"W</geoCoordinate>
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, 8 m, leg. K. & C. Groh, Jul. 4 1983 and Jul. 5 1983; CKG/3, Barbinha, Quaternary aeolinites, layer 2,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.067776">33°04'04"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.296944">16°17'49"W</geoCoordinate>
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, 8 m, leg. K. & C. Groh, Jul. 4 1983 and Jul. 5 1983; CKG/8, Barbinha, Quaternary aeolinites, layer 4,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.067776">33°04'04"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.296944">16°17'49"W</geoCoordinate>
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, 8 m, leg. K. & C. Groh, Jul. 4 1983 and Jul. 5 1983; CKG/2, south coast, ditch at the in 1985 new built harbour E Vila Baleira, lens of Quaternary calcareous aeolinite within a marine sandy beach terrace (nowadays beyond the quay of the harbour),
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.06389">33°03'50"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.315556">16°18'56"W</geoCoordinate>
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, 0-1.5 m, leg. J. Gerber, K. Groh & J. Hemmen, Aug. 1985; CWDM/8, Porto Santo, excavated mud walls behind the cart speedway E of the new harbour of Porto Santo,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.06333">33°03'48"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.30611">16°18'22"W</geoCoordinate>
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, 30 m, leg. W. De Mattia & J. Macor, May 24 2015; CWDM/9, E of Vila Baleira, S slope of the hill above Vale do Touro, 50 m W of the oil tanks, excavated Quaternary mixed gravel,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.063053">33°03'47"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.32389">16°19'26"W</geoCoordinate>
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, 24 m, leg. W. De Mattia & J. Macor, May 24 2015; CWDM/20, Pico do Baixo, E entrance of the tunnel, Quaternary mud deposit,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.06222">33°03'44"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.295834">16°17'45"W</geoCoordinate>
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, 20 m, leg. W. De Mattia & J. Macor, May 24. 2015.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="locus typicus">
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<paragraph pageId="58" pageNumber="59">Locus typicus.</paragraph>
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<paragraph pageId="58" pageNumber="59">Porto Santo, Madeira.</paragraph>
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</subSubSection>
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<subSubSection pageId="58" pageNumber="59" type="description">
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<paragraph pageId="58" pageNumber="59">Original description.</paragraph>
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<paragraph pageId="58" pageNumber="59">
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From
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<bibRefCitation author="Wollaston, TV" journalOrPublisher="Reeve, London" pageId="77" pageNumber="78" title="Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena." year="1878">Wollaston 1878</bibRefCitation>
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: var.
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<normalizedToken originalValue="β">β</normalizedToken>
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. subcarinulata - Major, spira magis elevata, ad apicem paulo magis acuta, anfractibus in medio obsolete subcarinulato. - Long. axis
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<normalizedToken originalValue="2½">21/2</normalizedToken>
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lin.; diam.
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<normalizedToken originalValue="3½">31/2</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="redescription">
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<paragraph pageId="58" pageNumber="59">Redescription of shell.</paragraph>
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<paragraph lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
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Shell large for the genus, with 5.9 regularly increasing whorls, the protoconch with 2.5 whorls. The form of the shell is flat conical, the slightly convex teleoconch whorls separated by a rather shallow but distinct, simple suture. The body whorl shows a distinct, rounded keel, which is enforced by a concave impression below the periphery. The last whorl measures 70%, the penultimate whorl 10% of the total shell height. The lower part of the body whorl is beneath the significantly marked periphery in frontal view first significantly concave, later moderately convex. The keel of the body whorl is located in the upper part of its total height. The aperture, which is inclined to the vertical axis of the shell in an angle of 69° and is descending in its last 5% in an angle of 36° to the horizontal axis, is horizontally elliptic; towards the umbilical region a bit flattened. Its width measures 45% of the total shell width and its height 44% of the total shell height. It has a peristome that is completely detached
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<pageBreakToken pageId="59" pageNumber="60" start="start">from</pageBreakToken>
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the body whorl. The lip is distinctly reflected and very wide at its basal and columellar sides. The eccentric umbilicus, which measures 13% of the total shell width, is shaped like a pinhole in the penultimate whorl and is completely closed in the earlier whorls. The protoconch is smooth, the teleoconch shows a number of oblique radial ribs, 10 in the penultimate quadrant of the body-whorl and is additionally covered by numerous small, round tubercles. The number of tubercles in the standard-quadrate of the base is 108. There is a yellowish hue of colouration in the teleoconch; the keel usually distinctly lighter in colour. See Figs 197-198.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="59" pageNumber="60" type="description">
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<paragraph pageId="59" pageNumber="60">Measurements.</paragraph>
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<paragraph pageId="59" pageNumber="60">
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D 8.3
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.3 mm (range 8.0-8.8 mm); H 6.1
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.3 mm (range 5.8-6.4 mm); FW 4.1
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2 mm (range 3.9-4.2 mm); PA 48.2
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<normalizedToken originalValue="±">+/-</normalizedToken>
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5.4° (range 42.6-54.4°); DU 0.6
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0,06 mm (range 0.5-0.7 mm); NT> 100; NW 5.6
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.1 (range 5.5-5.7) (n = 25). Ratio D/H 1.4; ratio FW/H 0.5.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="59" pageNumber="60" type="distribution">
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<paragraph pageId="59" pageNumber="60">Distribution.</paragraph>
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<paragraph pageId="59" pageNumber="60">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia subcarinulata" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="subcarinulata">Wollastonia subcarinulata</taxonomicName>
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is known from the Quaternary mud and slope deposits along the southern slopes of the eastern part of Porto Santo, from Vale do Touro to the area around the tunnel of Ponta da
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<normalizedToken originalValue="Galé">Gale</normalizedToken>
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beneath Pico do Baixo. See map in Fig. 199.
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</paragraph>
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<caption pageId="59" pageNumber="60">
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<paragraph pageId="59" pageNumber="60">
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Figure 199. Map of the distribution of
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia subcarinulata" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="subcarinulata">Wollastonia subcarinulata</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="comparisons">
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<paragraph pageId="59" pageNumber="60">Comparison and comments.</paragraph>
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<paragraph lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia subcarinulata" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="subcarinulata">Wollastonia subcarinulata</taxonomicName>
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can be confused with other, rather large-sized species of the genera
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">Hystricella</taxonomicName>
|
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and
|
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">Wollastonia</taxonomicName>
|
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|
like
|
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<taxonomicName lsidName="H. echinoderma" pageId="59" pageNumber="60" rank="species" species="echinoderma">H. echinoderma</taxonomicName>
|
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|
|
,
|
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|
|
<taxonomicName lsidName="W. vermetiformis" pageId="59" pageNumber="60" rank="species" species="vermetiformis">W. vermetiformis</taxonomicName>
|
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|
|
and
|
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|
|
<taxonomicName lsidName="W. falknerorum" pageId="59" pageNumber="60" rank="species" species="falknerorum">W. falknerorum</taxonomicName>
|
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|
|
sp. n., from which it can be distinguished, however, by the lack of a second keel, the rounded shape of the keel and a much finer sculpture as
|
|||
|
|
<pageBreakToken pageId="60" pageNumber="61" start="start">well</pageBreakToken>
|
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|
|
as a more depressed conical form. From similar sized
|
|||
|
|
<taxonomicName lsidName="W. inexpectata" pageId="60" pageNumber="61" rank="species" species="inexpectata">W. inexpectata</taxonomicName>
|
|||
|
|
it is distinguishable by the wider umbilicus, a flatter form, the keeled instead of angled periphery and a coarser granulation. From the recent
|
|||
|
|
<taxonomicName lsidName="W. oxytropis" pageId="60" pageNumber="61" rank="species" species="oxytropis">W. oxytropis</taxonomicName>
|
|||
|
|
it can be distinguished by the larger size, the wider umbilicus, the more convex teleoconch whorls that are separated by a more distinctly marked suture, the finer granulation, and the wider aperture.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="60" pageNumber="61" type="taxonomic remarks">
|
|||
|
|
<paragraph pageId="60" pageNumber="61">Taxonomic remarks.</paragraph>
|
|||
|
|
<paragraph pageId="60" pageNumber="61">
|
|||
|
|
The species is included in the genus
|
|||
|
|
<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia" order="Stylommatophora" pageId="60" pageNumber="61" phylum="Mollusca" rank="genus">Wollastonia</taxonomicName>
|
|||
|
|
because of its size and surface sculpture that is similar to
|
|||
|
|
<taxonomicName lsidName="W. oxytropis" pageId="60" pageNumber="61" rank="species" species="oxytropis">W. oxytropis</taxonomicName>
|
|||
|
|
, as a variety of which it was described by
|
|||
|
|
<bibRefCitation author="Wollaston, TV" journalOrPublisher="Reeve, London" pageId="77" pageNumber="78" title="Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena." year="1878">Wollaston (1878)</bibRefCitation>
|
|||
|
|
. Subsequent authors, except
|
|||
|
|
<bibRefCitation author="Cameron, RAD" journalOrPublisher="Malacologia" pageId="74" pageNumber="75" pagination="25 - 59" title="Fossil land snail faunas from Porto Santo, Madeiran Archipelago: change and stasis in Pleistocene and recent times." url="https://doi.org/10.4002/1543-8120-49.1.25" volume="49" year="2006">Cameron et al. (2006)</bibRefCitation>
|
|||
|
|
, recognised the taxon as a subspecies of
|
|||
|
|
<taxonomicName lsidName="W. oxytropis" pageId="60" pageNumber="61" rank="species" species="oxytropis">W. oxytropis</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="60" pageNumber="61" type="status and conservation">
|
|||
|
|
<paragraph pageId="60" pageNumber="61">Status and conservation.</paragraph>
|
|||
|
|
<paragraph pageId="60" pageNumber="61">
|
|||
|
|
Extinct before the
|
|||
|
|
<normalizedToken originalValue="islands’">islands'</normalizedToken>
|
|||
|
|
scientific exploration in the 19th century, possibly already before human settlement.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|