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<document id="00384C0D42C834AE6BAEAB0B77E88F65" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A0059FFDCFF182C8A16C0341F" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus pertuberculatus Perkins" docType="treatment" docVersion="6" lastPageNumber="478" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="478" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="63CB6F8785BA46CCD513194C450507C9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B61BAB91E1D5EA1CC17A480377126467">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="B1097987EAAFB3E50CCB3D81CD0D63CF">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A0DC9C3703BAA96DFD19FB7F7C57476D">text</mods:typeOfResource>
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<mods:title id="582749FC54398462787F04FBB129DC9C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AC85CDEBAED368AEAA8DFD34369B9B24">
<mods:date id="63E1C4675A0DE237B43E1E4C3DC05127">2021</mods:date>
<mods:detail id="F288186D1C02D04EA53BF283BEDF2CA8" type="pubDate">
<mods:number id="AAE90ADA99B7DF1C10EB834FCCA4640F">2021-05-21</mods:number>
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<mods:number id="AE548EE24BC12A1469209595CFF27E6B">4974</mods:number>
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<mods:number id="F3371455F1284D3D47ECA6F10B96D0FD">3</mods:number>
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<mods:classification id="447E3C2DCA60D7A11A892D4CB3A70500">journal article</mods:classification>
<mods:identifier id="675E7420EFB4F902D0CD0BB39F26061C" type="CLB-Dataset">6214</mods:identifier>
<mods:identifier id="A5FD4E22D83D2A93CED3FD5FF88F252A" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
<mods:identifier id="3D1492CC9D26073ECD25EF7C48F127CE" type="GBIF-Dataset">5b4eaeb3-5722-4021-84ad-125ccd80855f</mods:identifier>
<mods:identifier id="47B7B1F2E321F1E653A1C1255FCE36AB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8E44660BB6B197E754EA97033B26B7F3" type="Zenodo-Dep">4777839</mods:identifier>
<mods:identifier id="A165C956FED20F77834DBF4203144C95" type="ZooBank">1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8</mods:identifier>
</mods:mods>
<treatment id="4772526A0059FFDCFF182C8A16C0341F" ID-DOI="http://doi.org/10.5281/zenodo.4777934" ID-GBIF-Taxon="182642558" ID-Zenodo-Dep="4777934" LSID="urn:lsid:plazi:treatment:4772526A0059FFDCFF182C8A16C0341F" httpUri="http://treatment.plazi.org/id/4772526A0059FFDCFF182C8A16C0341F" lastPageNumber="478" pageId="19" pageNumber="478">
<subSubSection id="87C1B0F70059FFDCFF182C8A1757319C" box="[151,586,402,429]" pageId="19" pageNumber="478" type="nomenclature">
<paragraph id="CF64E37C0059FFDCFF182C8A1757319C" blockId="19.[151,586,402,465]" box="[151,586,402,429]" pageId="19" pageNumber="478">
<heading id="942C54100059FFDCFF182C8A1757319C" bold="true" box="[151,586,402,429]" fontSize="11" level="1" pageId="19" pageNumber="478" reason="1">
<taxonomicName id="08DB98FF0059FFDCFF182C8A1757319C" ID-CoL="8Y4R3" authority="Perkins" authorityName="Perkins" box="[151,586,402,429]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="pertuberculatus">
<emphasis id="FDAF3F6E0059FFDCFF182C8A1757319C" bold="true" box="[151,586,402,429]" pageId="19" pageNumber="478">
<emphasis id="FDAF3F6E0059FFDCFF182C8A14FE319D" bold="true" box="[151,483,402,429]" italics="true" pageId="19" pageNumber="478">Hydrochus pertuberculatus</emphasis>
Perkins
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70059FFDCFF182CAF141031E1" box="[151,269,438,465]" pageId="19" pageNumber="478" type="description">
<paragraph id="CF64E37C0059FFDCFF182CAF141031E1" blockId="19.[151,586,402,465]" box="[151,269,438,465]" pageId="19" pageNumber="478">
<figureCitation id="57E0FFF90059FFDCFF182CAF15FE31E1" box="[151,227,439,465]" captionStart="FIGURE 1" captionStartId="4.[151,250,1863,1887]" captionTargetBox="[186,1395,194,1808]" captionTargetId="figure-22@4.[151,1435,179,1837]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777843" httpUri="https://zenodo.org/record/4777843/files/figure.png" pageId="19" pageNumber="478">Figs. 1</figureCitation>
,
<figureCitation id="57E0FFF90059FFDCFF7E2CAE141031E1" box="[241,269,438,465]" captionStart="FIGURES 34–36" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[166,1421,181,1920]" captionTargetId="figure-15@37.[166,1421,181,1920]" captionTargetPageId="37" captionText="FIGURES 34–36. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777912" httpUri="https://zenodo.org/record/4777912/files/figure.png" pageId="19" pageNumber="478">35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70059FFDCFF182CE315EB3293" pageId="19" pageNumber="478" type="reference_group">
<paragraph id="CF64E37C0059FFDCFF182CE317613223" blockId="19.[151,636,507,531]" box="[151,636,507,531]" pageId="19" pageNumber="478">
<treatmentCitationGroup id="EFCBC4520059FFDCFF182CE317613223" box="[151,636,507,531]" pageId="19" pageNumber="478">
<treatmentCitation id="4E7AC56D0059FFDCFF182CE317653223" ID-DOI="http://doi.org/10.5281/zenodo.3883399" ID-Zenodo-Dep="3883399" author="Perkins, P. D." box="[151,632,507,531]" httpUri="http://treatment.plazi.org/id/060AF5509B223704FC2C16D9FDA7FD9D" page="297" pageId="19" pageNumber="478" year="2020">
<taxonomicName id="08DB98FF0059FFDCFF182CE317653223" ID-CoL="8Y4R3" authority="Perkins 2020 a: 297" authorityName="Perkins" authorityPageNumber="297" authorityYear="2020" box="[151,632,507,531]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="pertuberculatus">
<emphasis id="FDAF3F6E0059FFDCFF182CE314B63223" box="[151,427,507,531]" italics="true" pageId="19" pageNumber="478">Hydrochus pertuberculatus</emphasis>
<bibRefCitation id="AB4A9E8D0059FFDCFE3D2CE317653223" author="Perkins, P. D." box="[434,632,507,531]" pageId="19" pageNumber="478" pagination="291 - 301" refId="ref25459" refString="Perkins, P. D. (2020 a) Taxonomy of Guatemalan water beetles in the genus Hydrochus (Coleoptera: Hydrochidae). Acta Entomologica Musei Nationalis Pragae, 60 (1), 291 - 301. https: // doi. org / 10.37520 / aemnp. 2020.016" type="journal article" year="2020">Perkins 2020a: 297</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CF64E37C0059FFDCFF182F5915EB3293" blockId="19.[151,1437,577,1072]" pageId="19" pageNumber="478">
<materialsCitation id="7FB3E9210059FFDCFF182F5915EF3293" ID-GBIF-Occurrence="3128194318" country="Guatemala" county="Images" location="Originally" municipality="Following" pageId="19" pageNumber="478" specimenCount="1" typeStatus="holotype">
<location id="CA04B5A70059FFDCFF182F591417326B" LSID="urn:lsid:plazi:treatment:4772526A0059FFDCFF182C8A16C0341F:CA04B5A70059FFDCFF182F591417326B" box="[151,266,577,603]" country="Guatemala" county="Images" municipality="Following" name="Originally" pageId="19" pageNumber="478">Originally</location>
described from
<collectingCountry id="B7CCA3EC0059FFDCFE4D2F591722326B" box="[450,575,577,603]" name="Guatemala" pageId="19" pageNumber="478">Guatemala</collectingCountry>
.
<collectingMunicipality id="2F0079060059FFDCFDC52F5917A0326B" box="[586,701,577,603]" pageId="19" pageNumber="478">Following</collectingMunicipality>
are new locality data records for
<collectingCountry id="B7CCA3EC0059FFDCFBB82F5911AD326B" box="[1079,1200,577,603]" name="Guatemala" pageId="19" pageNumber="478">Guatemala</collectingCountry>
and
<collectingCountry id="B7CCA3EC0059FFDCFB682F59105D326B" box="[1255,1344,577,603]" name="Mexico" pageId="19" pageNumber="478">Mexico</collectingCountry>
.
<collectingCounty id="26059BF00059FFDCFAC42F5A1081326C" box="[1355,1436,578,604]" pageId="19" pageNumber="478">Images</collectingCounty>
of the habitus and male genitalia of the
<typeStatus id="10605DDE0059FFDCFDDE2F7D17AB324F" box="[593,694,613,639]" pageId="19" pageNumber="478" type="holotype">holotype</typeStatus>
, and detailed description of the male genitalia are given in
<bibRefCitation id="AB4A9E8D0059FFDCFAC72F7D15EF3293" author="Perkins, P. D." pageId="19" pageNumber="478" pagination="291 - 301" refId="ref25459" refString="Perkins, P. D. (2020 a) Taxonomy of Guatemalan water beetles in the genus Hydrochus (Coleoptera: Hydrochidae). Acta Entomologica Musei Nationalis Pragae, 60 (1), 291 - 301. https: // doi. org / 10.37520 / aemnp. 2020.016" type="journal article" year="2020">Perkins (2020a)</bibRefCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70059FFDCFF482FB514B53367" pageId="19" pageNumber="478" type="materials_examined">
<paragraph id="CF64E37C0059FFDCFF482FB514B53367" blockId="19.[151,1437,577,1072]" pageId="19" pageNumber="478">
<emphasis id="FDAF3F6E0059FFDCFF482FB514FB32F7" bold="true" box="[199,486,685,711]" pageId="19" pageNumber="478">Material examined (10)</emphasis>
:
<materialsCitation id="7FB3E9210059FFDCFE762FB517EB32DB" ID-GBIF-Occurrence="3128194364" collectingDate="2013-10-02" collectingDateMax="2013-10-03" collectingDateMin="2013-10-02" collectionCode="WSU" collectorName="R. S. Zack" country="Guatemala" latitude="16.52414" location="Parque National El Rosario" longLatPrecision="1" longitude="-90.16009" pageId="19" pageNumber="478" specimenCount="1" stateProvince="Peten">
<emphasis id="FDAF3F6E0059FFDCFE762FB5176032F7" bold="true" box="[505,637,685,711]" pageId="19" pageNumber="478">
<collectingCountry id="B7CCA3EC0059FFDCFE762FB5176032F7" box="[505,637,685,711]" name="Guatemala" pageId="19" pageNumber="478">Guatemala</collectingCountry>
</emphasis>
:
<collectingRegion id="0D1F2D9E0059FFDCFD002FB617CD32F8" box="[655,720,686,712]" country="Guatemala" name="Peten" pageId="19" pageNumber="478">Peten</collectingRegion>
,
<location id="CA04B5A70059FFDCFD502FB6113D32F7" LSID="urn:lsid:plazi:treatment:4772526A0059FFDCFF182C8A16C0341F:CA04B5A70059FFDCFD502FB6113D32F7" box="[735,1056,685,712]" country="Guatemala" latitude="16.52414" longLatPrecision="1" longitude="-90.16009" name="Parque National El Rosario" pageId="19" pageNumber="478" stateProvince="Peten">Parque National El Rosario</location>
,
<locationDeviation id="DED6FA5E0059FFDCFBA02FB611C632F7" box="[1071,1243,685,712]" location="Sayaxche" pageId="19" pageNumber="478">E of Sayaxche</locationDeviation>
, (
<geoCoordinate id="AAEF85BB0059FFDCFB7D2FB5108532F7" box="[1266,1432,685,711]" degrees="16" direction="north" minutes="31" orientation="latitude" pageId="19" pageNumber="478" precision="1" seconds="26.9" value="16.52414">16°31’26.9’’N</geoCoordinate>
,
<geoCoordinate id="AAEF85BB0059FFDCFF182FC9145E32DC" box="[151,323,721,748]" degrees="90" direction="west" minutes="9" orientation="longitude" pageId="19" pageNumber="478" precision="1" seconds="36.32" value="-90.16009">90°9’36.32’’W</geoCoordinate>
),
<date id="BB65C5BC0059FFDCFED82FC914CE32DB" box="[343,467,721,747]" pageId="19" pageNumber="478" value="2013-10-02" valueMax="2013-10-03" valueMin="2013-10-02">
<collectingDate id="AB213C540059FFDCFED82FC914CE32DB" box="[343,467,721,747]" pageId="19" pageNumber="478" value="2013-10-02" valueMax="2013-10-03" valueMin="2013-10-02">2-3.x.2013</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0059FFDCFD802FCA179B32DB" box="[527,646,721,748]" pageId="19" pageNumber="478">R. S. Zack</collectorName>
(1
<collectionCode id="A9CA7BB90059FFDCFD242FC917F332DB" box="[683,750,721,747]" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="19" pageNumber="478">WSU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7FB3E9210059FFDCFC8E2FC914703320" ID-GBIF-Occurrence="3128194538" collectingDate="1962-07-01" collectionCode="NMNH" collectorName="D. H. Janzen" country="Mexico" location="Cotaxtla Exp. Sta." municipality="Cotaxtla" pageId="19" pageNumber="478" specimenCount="1">
<emphasis id="FDAF3F6E0059FFDCFC8E2FC9164532DB" bold="true" box="[769,856,721,747]" pageId="19" pageNumber="478">
<collectingCountry id="B7CCA3EC0059FFDCFC8E2FC9164532DB" box="[769,856,721,747]" name="Mexico" pageId="19" pageNumber="478">Mexico</collectingCountry>
</emphasis>
:
<collectingMunicipality id="2F0079060059FFDCFCE82FC916D432DB" box="[871,969,721,747]" pageId="19" pageNumber="478">Cotaxtla</collectingMunicipality>
,
<location id="CA04B5A70059FFDCFC5B2FC911BE32DB" LSID="urn:lsid:plazi:treatment:4772526A0059FFDCFF182C8A16C0341F:CA04B5A70059FFDCFC5B2FC911BE32DB" box="[980,1187,721,748]" country="Mexico" municipality="Cotaxtla" name="Cotaxtla Exp. Sta." pageId="19" pageNumber="478">Cotaxtla Exp. Sta.</location>
,
<date id="BB65C5BC0059FFDCFB212FC9103F32DB" box="[1198,1314,721,747]" pageId="19" pageNumber="478" value="1962-07-01">
<collectingDate id="AB213C540059FFDCFB212FC9103F32DB" box="[1198,1314,721,747]" pageId="19" pageNumber="478" value="1962-07-01">1.vii.1962</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0059FFDCFAD02FCA15FE3320" pageId="19" pageNumber="478">D. H. Janzen</collectorName>
(1
<collectionCode id="A9CA7BB90059FFDCFE852FEE14783320" box="[266,357,758,784]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="478" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210059FFDCFEF72FED11CA3320" ID-GBIF-Occurrence="3128194347" box="[376,1239,757,784]" collectingDate="1965-08-23" collectionCode="NMNH" collectorName="Mi. S. Las Cruces &amp; Paul J. Spangler" country="Mexico" location="Chiapas" pageId="19" pageNumber="478" specimenCount="1" stateProvince="Chiapas">
<collectingRegion id="0D1F2D9E0059FFDCFEF72FED14C8333F" box="[376,469,757,783]" country="Mexico" name="Chiapas" pageId="19" pageNumber="478">Chiapas</collectingRegion>
, 5
<collectorName id="622E86AA0059FFDCFE782FED17D8333F" box="[503,709,757,784]" pageId="19" pageNumber="478">Mi. S. Las Cruces</collectorName>
,
<date id="BB65C5BC0059FFDCFD5F2FED16463320" box="[720,859,757,784]" pageId="19" pageNumber="478" value="1965-08-23">
<collectingDate id="AB213C540059FFDCFD5F2FED16463320" box="[720,859,757,784]" pageId="19" pageNumber="478" value="1965-08-23">23.viii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0059FFDCFC182FED1153333F" box="[919,1102,757,784]" pageId="19" pageNumber="478">Paul J. Spangler</collectorName>
(3
<collectionCode id="A9CA7BB90059FFDCFBFB2FEE11D23320" box="[1140,1231,758,784]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="478" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210059FFDCFB6D2FEE16BA3304" ID-GBIF-Occurrence="3128194576" collectingDate="1963-06-28" collectionCode="NMNH" collectorName="Robt. E. Woodruff" country="Mexico" location="Huatusco" pageId="19" pageNumber="478" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<collectingRegion id="0D1F2D9E0059FFDCFB6D2FEE10543320" box="[1250,1353,758,784]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="19" pageNumber="478">Veracruz</collectingRegion>
,
<locationDeviation id="DED6FA5E0059FFDCFADB2FED15F33304" location="Rt." pageId="19" pageNumber="478">
<quantity id="08234E990059FFDCFADB2FED1080333F" box="[1364,1437,757,783]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="19" pageNumber="478" unit="mi" value="20.0">20 mi.</quantity>
NW Rt.
</locationDeviation>
144
<location id="CA04B5A70059FFDCFEAA2E02148F3304" LSID="urn:lsid:plazi:treatment:4772526A0059FFDCFF182C8A16C0341F:CA04B5A70059FFDCFEAA2E02148F3304" box="[293,402,794,820]" country="Mexico" name="Huatusco" pageId="19" pageNumber="478" stateProvince="Veracruz de Ignacio de la Llave">Huatusco</location>
,
<date id="BB65C5BC0059FFDCFE132E01170A3303" box="[412,535,793,819]" pageId="19" pageNumber="478" value="1963-06-28">
<collectingDate id="AB213C540059FFDCFE132E01170A3303" box="[412,535,793,819]" pageId="19" pageNumber="478" value="1963-06-28">28.vi.1963</collectingDate>
</date>
, leg.
<collectorName id="622E86AA0059FFDCFDDD2E01163D3303" box="[594,800,793,820]" pageId="19" pageNumber="478">Robt. E. Woodruff</collectorName>
(4
<collectionCode id="A9CA7BB90059FFDCFCCA2E0216BD3304" box="[837,928,794,820]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="478" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E9210059FFDCFC3D2E0114B93367" ID-GBIF-Occurrence="3128194435" collectingDate="1952-07-01" collectionCode="SEMC" country="Mexico" location="Cordoba" pageId="19" pageNumber="478" specimenCount="1">
<location id="CA04B5A70059FFDCFC3D2E0111083303" LSID="urn:lsid:plazi:treatment:4772526A0059FFDCFF182C8A16C0341F:CA04B5A70059FFDCFC3D2E0111083303" box="[946,1045,793,819]" country="Mexico" name="Cordoba" pageId="19" pageNumber="478">Cordoba</location>
,
<quantity id="08234E990059FFDCFBAF2E0111753303" box="[1056,1128,793,819]" metricMagnitude="4" metricUnit="m" metricValue="3.0577536" pageId="19" pageNumber="478" unit="mi" value="19.0">19 mi.</quantity>
NE,
<date id="BB65C5BC0059FFDCFB2E2E0110083304" box="[1185,1301,793,820]" pageId="19" pageNumber="478" value="1952-07-01">
<collectingDate id="AB213C540059FFDCFB2E2E0110083304" box="[1185,1301,793,820]" pageId="19" pageNumber="478" value="1952-07-01">1.vii.1952</collectingDate>
</date>
, leg. collector unknown (1
<collectionCode id="A9CA7BB90059FFDCFEC42E2514813367" box="[331,412,829,855]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="19" pageNumber="478" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F70059FFDCFF482E7916C0341F" pageId="19" pageNumber="478" type="diagnosis">
<paragraph id="CF64E37C0059FFDCFF482E7916C0341F" blockId="19.[151,1437,577,1072]" pageId="19" pageNumber="478">
<emphasis id="FDAF3F6E0059FFDCFF482E7914D3334B" bold="true" box="[199,462,865,891]" pageId="19" pageNumber="478">Differential diagnosis.</emphasis>
Among Central American species, recognized by the combination of moderately large size (ca.
<quantity id="08234E990059FFDCFF752E9D147F3390" box="[250,354,901,928]" metricMagnitude="-3" metricUnit="m" metricValue="3.56" pageId="19" pageNumber="478" unit="mm" value="3.56">3.56 mm</quantity>
), the moderately low and relatively impunctate pronotal reliefs, the large tubercle of the 5
<superScript id="38AE4E340059FFDCFAC22E9D104733A3" attach="left" box="[1357,1370,901,915]" fontSize="6" pageId="19" pageNumber="478">th</superScript>
interstria, and the male genitalia. This species has a generalized
<taxonomicName id="08DB98FF0059FFDCFCBD2EB116B133F3" authorityName="Leach" authorityYear="1817" box="[818,940,937,963]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="478" phylum="Arthropoda" rank="genus">
<emphasis id="FDAF3F6E0059FFDCFCBD2EB116B133F3" box="[818,940,937,963]" italics="true" pageId="19" pageNumber="478">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males. The male genitalia are superficially somewhat similar to that of
<taxonomicName id="08DB98FF0059FFDCFBCB2ED611F233D7" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[1092,1263,974,999]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="nayaritensis" status="sp. nov.">
<emphasis id="FDAF3F6E0059FFDCFBCB2ED611F233D7" box="[1092,1263,974,999]" italics="true" pageId="19" pageNumber="478">H. nayaritensis</emphasis>
</taxonomicName>
<emphasis id="FDAF3F6E0059FFDCFB7A2ED5102933D7" bold="true" box="[1269,1332,973,999]" pageId="19" pageNumber="478">
<taxonomicNameLabel id="E69C82150059FFDCFB7A2ED5102933D7" box="[1269,1332,973,999]" pageId="19" pageNumber="478" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, in being slender and having a very narrow aedeagus; however, the parameres, especially the sagittate tips, the basal piece shape, and the adbl shape and size are very different in
<emphasis id="FDAF3F6E0059FFDCFC8C290E16C0341F" box="[771,989,1045,1071]" italics="true" pageId="19" pageNumber="478">
<taxonomicName id="08DB98FF0059FFDCFC8C290E16C4341F" box="[771,985,1045,1071]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="478" phylum="Arthropoda" rank="species" species="pertuberculatus">H. pertuberculatus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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