treatments-xml/data/42/63/87/426387C8B47AFFE5FDCDFA1498C2FC04.xml

<document id="1CCF41749FFBCD93627BBBA284460614" ID-CLB-Dataset="26998" ID-DOI="10.5852/ejt.2019.517" ID-GBIF-Dataset="d05987e2-6008-4deb-9773-1d988013beba" ID-Zenodo-Dep="2647891" ID-ZooBank="CAC6F8AF-ED37-4989-9672-68316920750B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555772398596" checkinUser="plazi" docAuthor="Fernandes, Maurício Romulo &amp; Pimenta, Alexandre Dias" docDate="2019" docId="426387C8B47AFFE5FDCDFA1498C2FC04" docLanguage="en" docName="ejt-517_fernandes_pimenta.pdf" docOrigin="European Journal of Taxonomy 517" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Metaxia excelsa Faber &amp; Moolenbeek 1991" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="BE5AFFB0B47CFFECFFDEFFCA9B24FFA1" masterDocTitle="Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil" masterLastPageNumber="60" masterPageNumber="1" pageNumber="7" updateTime="1698723356892" updateUser="ExternalLinkService">
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<mods:title id="0FF27761F2C5BA435DE5FEA66C691E15">Basic anatomy of species of Triphoridae (Gastropoda, Triphoroidea) from Brazil</mods:title>
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<mods:namePart id="09EC9D80C0B49CDCEF06097C4AAA0A1F">Fernandes, Maurício Romulo</mods:namePart>
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<mods:namePart id="26770F9F881FAF7DBFF246992A618C2F">Pimenta, Alexandre Dias</mods:namePart>
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<paragraph id="CA7536DEB47AFFEAFDCDFA149F05FA58" blockId="6.[531,1057,1502,1564]" box="[531,1057,1502,1529]" pageId="6" pageNumber="7">
<heading id="913D81B2B47AFFEAFDCDFA149F05FA58" box="[531,1057,1502,1529]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="0DCA4D5DB47AFFEAFDCDFA149F05FA58" ID-CoL="6RDQT" authority="Faber &amp; Moolenbeek, 1991" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[531,1057,1502,1529]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB47AFFEAFDCDFA1499F1FA58" bold="true" box="[531,725,1502,1529]" italics="true" pageId="6" pageNumber="7">Metaxia excelsa</emphasis>
Faber &amp; Moolenbeek, 1991
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<subSubSection id="82D06555B47AFFEAFD02F9C8987CF9BD" box="[732,856,1538,1564]" pageId="6" pageNumber="7" type="description">
<paragraph id="CA7536DEB47AFFEAFD02F9C8987CF9BD" blockId="6.[531,1057,1502,1564]" box="[732,856,1538,1564]" pageId="6" pageNumber="7">
<figureCitation id="52F12A5BB47AFFEAFD02F9C89818F9BD" box="[732,828,1538,1564]" captionStart="Fig" captionStartId="7.[189,232,1707,1733]" captionTargetBox="[189,1398,742,1666]" captionTargetId="figure@7.[189,1399,742,1666]" captionTargetPageId="7" captionText="Fig. 2. Specimens studied. A. Metaxia excelsa Faber &amp; Moolenbeek, 1991 (MZSP 122353). B. Metaxia rugulosa (C.B. Adams, 1850) (BMSM 56034). C. Cosmotriphora melanura (C.B. Adams, 1850) (MNRJ 33980). D. Iniforis pseudothomae Rolán &amp; Fernández-Garcés, 1993 (MZSP 100957). E. Latitriphora albida (A. Adams, 1854) (BMSM 55442). F. Monophorus olivaceus (Dall, 1889) (MNRJ 34615). G. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989) (MZSP 85022). H. Nototriphora decorata (C.B. Adams, 1850) (MNRJ 28941). I. Sagenotriphora osclausum (Rolán &amp; Fernández-Garcés, 1995) (MNRJ 35061). J. Similiphora intermedia (C.B. Adams, 1850) (MNRJ 29763). K. Strobiligera gaesona (Dall, 1927) (MZSP 53685). L. “Inella” harryleei Rolán &amp; Fernández-Garcés, 2008 (FLMNH 450495). Scale bars: 1 mm." httpUri="https://zenodo.org/record/2647895/files/figure.png" pageId="6" pageNumber="7">Figs 2A</figureCitation>
, 
<figureCitation id="52F12A5BB47AFFEAFC97F9C8987CF9BD" box="[841,856,1538,1564]" captionStart="Fig" captionStartId="8.[189,232,1923,1949]" captionTargetBox="[206,1381,265,1883]" captionTargetId="figure@8.[206,1381,265,1883]" captionTargetPageId="8" captionText="Fig. 3. Metaxia excelsa Faber &amp; Moolenbeek, 1991. A–G. MZSP 122353. A. External morphology. B–C. Jaw, outer side. D–G. Radula. Scale bars: A = 1 mm; B = 50 µm; C = 2 µm; D–F = 10 µm; G = 5 µm." httpUri="https://zenodo.org/record/2647897/files/figure.png" pageId="6" pageNumber="7">3</figureCitation>
</paragraph>
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<subSubSection id="82D06555B47AFFEAFF63F9899E34F911" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="CA7536DEB47AFFEAFF63F9899A8EF9FC" blockId="6.[189,426,1603,1629]" box="[189,426,1603,1629]" pageId="6" pageNumber="7">
<heading id="913D81B2B47AFFEAFF63F9899A8EF9FC" bold="true" box="[189,426,1603,1629]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="F8BEEACCB47AFFEAFF63F9899A8EF9FC" bold="true" box="[189,426,1603,1629]" pageId="6" pageNumber="7">Material examined</emphasis>
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<paragraph id="CA7536DEB47AFFEAFF63F9B89E34F911" blockId="6.[189,1398,1649,1712]" pageId="6" pageNumber="7">
<materialsCitation id="7AA23C83B47AFFEAFF63F9B89E28F911" ID-GBIF-Occurrence="2236664089" collectingDate="2012-08-10" collectingDateMax="53-12-23" collectingDateMin="2012-08-10" collectorName="G. H. Pereira Filho" country="Brazil" county="Rata" elevation="12" latitude="-3.8152778" location="Fernando de Noronha Archipelago" longLatPrecision="21" longitude="-32.391945" municipality="Ressureta" pageId="6" pageNumber="7" specimenCount="1">
<collectingCountry id="B2DD764EB47AFFEAFF63F9B89A02F92D" box="[189,294,1650,1676]" name="Brazil" pageId="6" pageNumber="7">BRAZIL</collectingCountry>
• [2, 1 d] specs; 
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, canal between islands 
<collectingMunicipality id="2A11ACA4B47AFFEAFB51F9B89E25F92D" box="[1167,1281,1650,1676]" pageId="6" pageNumber="7">Ressureta</collectingMunicipality>
and 
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; 
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, 
<geoCoordinate id="AFFE5019B47AFFEAFE84F9599AD1F90E" box="[346,501,1683,1711]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="-32.391945">32°23′31″ W</geoCoordinate>
; 
<elevation id="41E7D1EDB47AFFEAFDDFF95F9919F90E" box="[513,573,1685,1711]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="m" value="12.0">
<quantity id="0D329B3BB47AFFEAFDDFF95F9919F90E" box="[513,573,1685,1711]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="m" value="12.0">12 m</quantity>
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depth; 
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<date id="BE74101EB47AFFEAFD4AF95F9813F90E" box="[660,823,1685,1711]" pageId="6" pageNumber="7" value="2012-08-10">10 Aug. 2012</date>
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; 
<collectorName id="673F5308B47AFFEAFC9DF95F9F3FF90E" box="[835,1051,1685,1711]" pageId="6" pageNumber="7">G.H. Pereira Filho</collectorName>
leg.; MZSP 
<collectingDate id="AE30E9F6B47AFFEAFB6EF95F9E28F911" box="[1200,1292,1685,1712]" pageId="6" pageNumber="7" value="53-12-23">
<date id="BE74101EB47AFFEAFB6EF95F9E28F911" box="[1200,1292,1685,1712]" pageId="6" pageNumber="7" value="53-12-23">122353</date>
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.
</paragraph>
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<subSubSection id="82D06555B47AFFE5FF63F91C99BFFECD" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" type="description">
<paragraph id="CA7536DEB47AFFEAFF63F91C990DF951" blockId="6.[189,553,1750,1776]" box="[189,553,1750,1776]" pageId="6" pageNumber="7">
<heading id="913D81B2B47AFFEAFF63F91C990DF951" bold="true" box="[189,553,1750,1776]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="F8BEEACCB47AFFEAFF63F91C990DF951" bold="true" box="[189,553,1750,1776]" pageId="6" pageNumber="7">Description of basic anatomy</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB47AFFEAFF63F8CF9F56F8E3" blockId="6.[189,1398,1797,1858]" pageId="6" pageNumber="7">OPERCULUM. Elliptical, thin, semi-transparent, membranous, nucleus and number of whorls not discernible; diameter of operculum exceeds diameter of opercular pouch in 17%.</paragraph>
<paragraph id="CA7536DEB47AFFEAFF63F8A09E53F86B" blockId="6.[189,1399,1897,1995]" pageId="6" pageNumber="7">
JAW. Wing-shaped; outer side with scales usually rectangular/squared, sometimes rhombus-shaped or even oblong-lanceolate; some scales covered by micro-pores up to 
<quantity id="0D329B3BB47AFFEAFC2EF8479F68F806" box="[1008,1100,1933,1959]" metricMagnitude="-7" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="7" unit="nm" value="270.0">270 nm</quantity>
in diameter; rectangular scales 
<quantity id="0D329B3BB47AFFEAFED2F87A9A96F86B" box="[268,434,1968,1995]" metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.54" metricValueMin="1.05" pageId="6" pageNumber="7" unit="mm" value="12.95" valueMax="15.4" valueMin="10.5">10.5–15.4 µm</quantity>
long, 
<quantity id="0D329B3BB47AFFEAFE20F87A99A3F86B" box="[510,647,1968,1994]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="7.7" metricValueMin="4.8" pageId="6" pageNumber="7" unit="mm" value="6.25" valueMax="7.7" valueMin="4.8">4.8–7.7 µm</quantity>
wide, ratio length/width 1.8–2.9, rhombus-shaped scales 17.9–
</paragraph>
<paragraph id="CA7536DEB47BFFEBFF63FEC1982AFEE9" blockId="7.[189,1399,267,329]" pageId="7" pageNumber="8">
24.5 µm long, 5.9–10.0 µm wide, ratio length/width 2.4–3.1, oblong-lanceolate scales 
<quantity id="0D329B3BB47BFFEBFB0EFEC19E53FE87" box="[1232,1399,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.555" metricValueMax="1.65" metricValueMin="1.46" pageId="7" pageNumber="8" unit="mm" value="15.55" valueMax="16.5" valueMin="14.6">14.6–16.5 µm</quantity>
long, 4.0–5.0 µm wide, ratio length/width 2.9–3.6.
</paragraph>
<paragraph id="CA7536DEB47BFFE5FF63FEBA99BFFECD" blockId="7.[189,1399,368,676]" lastBlockId="9.[189,1399,267,364]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
RADULA. Formula 4-1-1-1-4; central tooth with four to five elongated and claw-like cusps, median cusp present or not but always thinner, with the basal plate of tooth assuming a concave format (i.e., outer cusps in an upper position than median one), outer cusps usually distinctly oriented outwards and reaching slightly larger dimensions (equal or up to 1.2× more elongated) than inner cusps; lateral teeth with four elongated claw-like cusps, basal plate of tooth slightly concave (not as much as central tooth), all cusps approximately with the same length, but cusp 1 considerably broader and sometimes slightly shorter than remaining cusps; marginal teeth gradually diminishing in size outwards; M1 and M2 with four cusps very similar to lateral teeth, with all cusps similar in length or width; M3 with three similar, moderately elongated, pointed and claw-like cusps; M4 small, with three curved and pointed cusps, median one 1.2 to 1.6 times more elongated than remaining cusps; central tooth 
<quantity id="0D329B3BB475FFE5FF63FEE49A6DFEE8" box="[189,329,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="9.2" metricValueMin="6.4" pageId="9" pageNumber="10" unit="mm" value="7.8" valueMax="9.2" valueMin="6.4">6.4–9.2 µm</quantity>
wide, lateral teeth 
<quantity id="0D329B3BB475FFE5FDE2FEE599ECFEE8" box="[572,712,303,329]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.9" metricValueMin="7.1" pageId="9" pageNumber="10" unit="mm" value="8.5" valueMax="9.9" valueMin="7.1">7.1–9.9 µm</quantity>
wide, M1 
<quantity id="0D329B3BB475FFE5FC8DFEE498FBFEE8" box="[851,991,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.7" metricValueMin="5.8" pageId="9" pageNumber="10" unit="mm" value="7.25" valueMax="8.7" valueMin="5.8">5.8–8.7 µm</quantity>
wide, M2 
<quantity id="0D329B3BB475FFE5FBB4FEE49FD2FEE8" box="[1130,1270,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="6.7" metricValueMin="4.1" pageId="9" pageNumber="10" unit="mm" value="5.4" valueMax="6.7" valueMin="4.1">4.1–6.7 µm</quantity>
wide, M3 
<quantity id="0D329B3BB475FFE5FF63FE989A62FECD" box="[189,326,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="9" pageNumber="10" unit="mm" value="3.8" valueMax="4.4" valueMin="3.2">3.2–4.4 µm</quantity>
wide, M4 
<quantity id="0D329B3BB475FFE5FE16FE989975FECD" box="[456,593,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.1" metricValueMin="2.3" pageId="9" pageNumber="10" unit="mm" value="2.7" valueMax="3.1" valueMin="2.3">2.3–3.1 µm</quantity>
wide.
</paragraph>
</subSubSection>
<caption id="9EB56656B47BFFEBFF63F9619A6DF87E" httpUri="https://zenodo.org/record/2647895/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1707,1733]" targetBox="[189,1398,742,1666]" targetPageId="7">
<paragraph id="CA7536DEB47BFFEBFF63F9619A6DF87E" blockId="7.[189,1399,1707,2015]" pageId="7" pageNumber="8">
<emphasis id="F8BEEACCB47BFFEBFF63F9619A2EF964" bold="true" box="[189,266,1707,1733]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
Specimens studied. 
<emphasis id="F8BEEACCB47BFFEBFE27F961992AF964" bold="true" box="[505,526,1707,1733]" pageId="7" pageNumber="8">A</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFDC2F9669F07F964" authority="Faber &amp; Moolenbeek, 1991" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[540,1059,1707,1733]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB47BFFEBFDC2F96699FDF964" box="[540,729,1707,1733]" italics="true" pageId="7" pageNumber="8">Metaxia excelsa</emphasis>
Faber &amp; Moolenbeek, 1991
</taxonomicName>
(MZSP 122353). 
<emphasis id="F8BEEACCB47BFFEBFB2BF9669E2DF967" bold="true" box="[1269,1289,1708,1734]" pageId="7" pageNumber="8">B</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFAC8F9669937F948" authority="(C. B. Adams, 1850)" authorityName="C. B. Adams" authorityYear="1850" baseAuthorityName="C. B. Adams" baseAuthorityYear="1850" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="rugulosa">
<emphasis id="F8BEEACCB47BFFEBFAC8F9669A01F948" italics="true" pageId="7" pageNumber="8">Metaxia rugulosa</emphasis>
(C.B. Adams, 1850)
</taxonomicName>
(BMSM 56034). 
<emphasis id="F8BEEACCB47BFFEBFD3EF90499D1F949" bold="true" box="[736,757,1742,1768]" pageId="7" pageNumber="8">C</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFCDCF9059E3DF948" authority="(C. B. Adams, 1850)" authorityName="C. B. Adams" authorityYear="1850" baseAuthorityName="C. B. Adams" baseAuthorityYear="1850" box="[770,1305,1742,1769]" class="Gastropoda" family="Triphoridae" genus="Cosmotriphora" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="melanura">
<emphasis id="F8BEEACCB47BFFEBFCDCF9059F0FF948" box="[770,1067,1743,1769]" italics="true" pageId="7" pageNumber="8">Cosmotriphora melanura</emphasis>
(C.B. Adams, 1850)
</taxonomicName>
(MNRJ 33980). 
<emphasis id="F8BEEACCB47BFFEBFEFFF9389A12F8AD" bold="true" box="[289,310,1778,1804]" pageId="7" pageNumber="8">D</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFE98F93898CCF8AD" authority="Rolan &amp; Fernandez-Garces, 1993" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="1993" box="[326,1000,1778,1804]" class="Gastropoda" family="Triphoridae" genus="Iniforis" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="pseudothomae">
<emphasis id="F8BEEACCB47BFFEBFE98F938996BF8AD" box="[326,591,1778,1804]" italics="true" pageId="7" pageNumber="8">Iniforis pseudothomae</emphasis>
Rolán &amp; Fernández-Garcés, 1993
</taxonomicName>
(MZSP 100957). 
<emphasis id="F8BEEACCB47BFFEBFB1FF9389FF1F8AD" bold="true" box="[1217,1237,1778,1804]" pageId="7" pageNumber="8">E</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFB3BF9389AD5F891" authority="(A. Adams, 1854)" authorityName=", Rolan &amp; Fernandez-Garces" authorityYear="1995" baseAuthorityName="A. Adams" baseAuthorityYear="1854" class="Gastropoda" family="Triphoridae" genus="Latitriphora" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="albida">
<emphasis id="F8BEEACCB47BFFEBFB3BF9389A2CF88E" italics="true" pageId="7" pageNumber="8">Latitriphora albida</emphasis>
(A. Adams, 1854)
</taxonomicName>
(BMSM 55442). 
<emphasis id="F8BEEACCB47BFFEBFD0DF8DF99C1F88E" bold="true" box="[723,741,1813,1839]" pageId="7" pageNumber="8">F</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFD27F8DF9F8FF88E" authority="(Dall, 1889)" authorityName="Dall" authorityYear="1889" baseAuthorityName="Dall" baseAuthorityYear="1889" box="[761,1195,1813,1839]" class="Gastropoda" family="Triphoridae" genus="Monophorus" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="olivaceus">
<emphasis id="F8BEEACCB47BFFEBFD27F8DF9F2EF88E" box="[761,1034,1813,1839]" italics="true" pageId="7" pageNumber="8">Monophorus olivaceus</emphasis>
(Dall, 1889)
</taxonomicName>
(MNRJ 34615). 
<emphasis id="F8BEEACCB47BFFEBFF63F8F29BF0F8F3" bold="true" box="[189,212,1848,1874]" pageId="7" pageNumber="8">G</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFF3CF8F29818F8F3" authority="(Moolenbeek &amp; Faber, 1989)" authorityName="" authorityYear="1989" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[226,828,1848,1874]" class="Gastropoda" family="Triphoridae" genus="Nanaphora" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="verbernei">
<emphasis id="F8BEEACCB47BFFEBFF3CF8F29AFBF8F3" box="[226,479,1848,1874]" italics="true" pageId="7" pageNumber="8">Nanaphora verbernei</emphasis>
(Moolenbeek &amp; Faber, 1989)
</taxonomicName>
(MZSP 85022). 
<emphasis id="F8BEEACCB47BFFEBFBDFF8F29F3CF8F3" bold="true" box="[1025,1048,1848,1874]" pageId="7" pageNumber="8">H</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFBF8F8F29A46F8D7" authority="(C. B. Adams, 1850)" authorityName="C. B. Adams" authorityYear="1850" baseAuthorityName="C. B. Adams" baseAuthorityYear="1850" class="Gastropoda" family="Triphoridae" genus="Nototriphora" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="F8BEEACCB47BFFEBFBF8F8F29E14F8F3" box="[1062,1328,1848,1874]" italics="true" pageId="7" pageNumber="8">Nototriphora decorata</emphasis>
(C.B. Adams, 1850)
</taxonomicName>
(MNRJ 28941). 
<emphasis id="F8BEEACCB47BFFEBFDF8F8969915F8D7" bold="true" box="[550,561,1884,1910]" pageId="7" pageNumber="8">I</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFDE1F8919E3CF8D7" authority="(Rolan &amp; Fernandez-Garces, 1995)" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="1995" baseAuthorityName="Rolan &amp; Fernandez-Garces" baseAuthorityYear="1995" box="[575,1304,1883,1910]" class="Gastropoda" family="Triphoridae" genus="Sagenotriphora" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="osclausum">
<emphasis id="F8BEEACCB47BFFEBFDE1F891985DF8D4" box="[575,889,1883,1909]" italics="true" pageId="7" pageNumber="8">Sagenotriphora osclausum</emphasis>
(Rolán &amp; Fernández-Garcés, 1995)
</taxonomicName>
(MNRJ 35061). 
<emphasis id="F8BEEACCB47BFFEBFEC1F8B59A0AF838" bold="true" box="[287,302,1919,1945]" pageId="7" pageNumber="8">J</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFEE3F8B59860F838" authority="(C. B. Adams, 1850)" authorityName="C. B. Adams" authorityYear="1850" baseAuthorityName="C. B. Adams" baseAuthorityYear="1850" box="[317,836,1918,1945]" class="Gastropoda" family="Triphoridae" genus="Similiphora" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="intermedia">
<emphasis id="F8BEEACCB47BFFEBFEE3F8B5996BF838" box="[317,591,1919,1945]" italics="true" pageId="7" pageNumber="8">Similiphora intermedia</emphasis>
(C.B. Adams, 1850)
</taxonomicName>
(MNRJ 29763). 
<emphasis id="F8BEEACCB47BFFEBFBD0F8B59F01F838" bold="true" box="[1038,1061,1919,1945]" pageId="7" pageNumber="8">K</emphasis>
. 
<taxonomicName id="0DCA4D5DB47BFFEBFBEAF8B59A25F81D" authority="(Dall, 1927)" authorityName="Dall" authorityYear="1927" baseAuthorityName="Dall" baseAuthorityYear="1927" class="Gastropoda" family="Triphoridae" genus="Strobiligera" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="gaesona">
<emphasis id="F8BEEACCB47BFFEBFBEAF8B59E0FF839" box="[1076,1323,1919,1945]" italics="true" pageId="7" pageNumber="8">Strobiligera gaesona</emphasis>
(Dall, 1927)
</taxonomicName>
(MZSP 53685). 
<emphasis id="F8BEEACCB47BFFEBFE18F8689AFEF81D" bold="true" box="[454,474,1954,1980]" pageId="7" pageNumber="8">L</emphasis>
. “ 
<taxonomicName id="0DCA4D5DB47BFFEBFE2BF868991DF81D" authorityName="Fernandes &amp; Pimenta" authorityYear="2019" box="[501,569,1954,1980]" class="Gastropoda" family="Triphoridae" genus="Inella" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB47BFFEBFE2BF868991DF81D" box="[501,569,1954,1980]" italics="true" pageId="7" pageNumber="8">Inella</emphasis>
</taxonomicName>
” 
<emphasis id="F8BEEACCB47BFFEBFD93F868999DF81D" box="[589,697,1954,1980]" italics="true" pageId="7" pageNumber="8">harryleei</emphasis>
Rolán &amp; Fernández-Garcés, 2008 (FLMNH 450495). Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="9EB56656B474FFE4FF63F8499A26F845" httpUri="https://zenodo.org/record/2647897/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1923,1949]" targetBox="[206,1381,265,1883]" targetPageId="8">
<paragraph id="CA7536DEB474FFE4FF63F8499A26F845" blockId="8.[189,1399,1923,2020]" pageId="8" pageNumber="9">
<emphasis id="F8BEEACCB474FFE4FF63F8499A29F83C" bold="true" box="[189,269,1923,1949]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
<taxonomicName id="0DCA4D5DB474FFE4FEC6F84E9817F83C" authority="Faber &amp; Moolenbeek, 1991" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[280,819,1923,1949]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB474FFE4FEC6F84E9AFCF83C" box="[280,472,1923,1949]" italics="true" pageId="8" pageNumber="9">Metaxia excelsa</emphasis>
Faber &amp; Moolenbeek, 1991
</taxonomicName>
. 
<emphasis id="F8BEEACCB474FFE4FC9FF8499858F83C" bold="true" box="[833,892,1923,1949]" pageId="8" pageNumber="9">A–G</emphasis>
. MZSP 122353. 
<emphasis id="F8BEEACCB474FFE4FB93F8499F46F83C" bold="true" box="[1101,1122,1923,1949]" pageId="8" pageNumber="9">A</emphasis>
. External morphology. 
<emphasis id="F8BEEACCB474FFE4FF63F86C9BD0F861" bold="true" box="[189,244,1958,1984]" pageId="8" pageNumber="9">B–C</emphasis>
. Jaw, outer side. 
<emphasis id="F8BEEACCB474FFE4FE1DF86C9ADAF861" bold="true" box="[451,510,1958,1984]" pageId="8" pageNumber="9">D–G</emphasis>
. Radula. Scale bars: A = 1 mm; B = 50 µm; C = 2 µm; D–F = 10 µm; G = 5 µm.
</paragraph>
</caption>
<subSubSection id="82D06555B475FFE5FF63FE5998C2FC04" pageId="9" pageNumber="10" type="discussion">
<paragraph id="CA7536DEB475FFE5FF63FE599A0BFE0C" blockId="9.[189,303,403,429]" box="[189,303,403,429]" pageId="9" pageNumber="10">
<heading id="913D81B2B475FFE5FF63FE599A0BFE0C" bold="true" box="[189,303,403,429]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="F8BEEACCB475FFE5FF63FE599A0BFE0C" bold="true" box="[189,303,403,429]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="CA7536DEB475FFE5FF63FE0898C2FC04" blockId="9.[189,1399,450,934]" pageId="9" pageNumber="10">
The radula of 
<taxonomicName id="0DCA4D5DB475FFE5FE80FE099AFEFE7D" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[350,474,450,476]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FE80FE099A52FE7D" box="[350,374,451,476]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FE5AFE089AFEFE7D" box="[388,474,450,476]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
shares several features with that of 
<taxonomicName id="0DCA4D5DB475FFE5FCA8FE099E56FE7D" authority="(Powell, 1930) (Marshall 1977)" authorityName="Marshall" authorityYear="1977" baseAuthorityName="Powell" baseAuthorityYear="1930" box="[886,1394,450,476]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="exaltata">
<emphasis id="F8BEEACCB475FFE5FCA8FE0998AAFE7D" box="[886,910,451,476]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FC45FE0898DEFE7D" box="[923,1018,450,476]" italics="true" pageId="9" pageNumber="10">exaltata</emphasis>
(Powell, 1930) (
<bibRefCitation id="AE5B4B2FB475FFE5FB65FE089E4DFE7D" author="Marshall B. A." box="[1211,1385,450,476]" journalOrPublisher="New Zealand Journal of Zoology" pageId="9" pageNumber="10" pagination="111 - 117" part="4" refId="ref21206" refString="Marshall B. A. 1977. The dextral triforid genus Metaxia (Mollusca: Gastropoda) in the south-west Pacific. New Zealand Journal of Zoology 4: 111 - 117." title="The dextral triforid genus Metaxia (Mollusca: Gastropoda) in the south-west Pacific" type="journal article" year="1977">Marshall 1977</bibRefCitation>
)
</taxonomicName>
, such as the general tooth morphology and the gradual decrease of size towards the outermost marginal teeth, confirming their affinity at genus-level. The distinction lies in the number of marginal teeth (only four in 
<taxonomicName id="0DCA4D5DB475FFE5FECCFDE69AABFDE4" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[274,399,555,582]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FECCFDE69A0EFDE4" box="[274,298,556,581]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FEE7FDE19AABFDE4" box="[313,399,555,581]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
, but nine or ten in 
<taxonomicName id="0DCA4D5DB475FFE5FDB4FDE699CAFDE4" authorityName="Marshall" authorityYear="1977" baseAuthorityName="Powell" baseAuthorityYear="1930" box="[618,750,555,582]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="exaltata">
<emphasis id="F8BEEACCB475FFE5FDB4FDE699A6FDE4" box="[618,642,556,581]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FD4EFDE199CAFDE4" box="[656,750,555,581]" italics="true" pageId="9" pageNumber="10">exaltata</emphasis>
</taxonomicName>
), the median cusp of the central tooth (apparently more reduced or vestigial in 
<taxonomicName id="0DCA4D5DB475FFE5FE11FD859972FDC8" authorityName="Marshall" authorityYear="1977" baseAuthorityName="Powell" baseAuthorityYear="1930" box="[463,598,591,617]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="exaltata">
<emphasis id="F8BEEACCB475FFE5FE11FD859AC3FDC9" box="[463,487,591,616]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FE26FD859972FDC8" box="[504,598,591,617]" italics="true" pageId="9" pageNumber="10">exaltata</emphasis>
</taxonomicName>
) and the number of cusps in the outermost marginal teeth (three in M4 of 
<taxonomicName id="0DCA4D5DB475FFE5FED1FDB99AAAFD2D" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[271,398,626,652]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FED1FDB99A03FD2D" box="[271,295,627,652]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FEE6FDB89AAAFD2D" box="[312,398,626,652]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
, two in M9 or M10 of 
<taxonomicName id="0DCA4D5DB475FFE5FD78FDB99809FD2D" authorityName="Marshall" authorityYear="1977" baseAuthorityName="Powell" baseAuthorityYear="1930" box="[678,813,626,652]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="exaltata">
<emphasis id="F8BEEACCB475FFE5FD78FDB9999AFD2D" box="[678,702,627,652]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FD11FDB89809FD2D" box="[719,813,626,652]" italics="true" pageId="9" pageNumber="10">exaltata</emphasis>
</taxonomicName>
). The radula of 
<taxonomicName id="0DCA4D5DB475FFE5FC2EFDB99F49FD2D" baseAuthorityName="Delle Chiaje" baseAuthorityYear="1828" box="[1008,1133,626,652]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="metaxa">
<emphasis id="F8BEEACCB475FFE5FC2EFDB99F2CFD2D" box="[1008,1032,627,652]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FBC6FDB99F49FD2D" box="[1048,1133,627,652]" italics="true" pageId="9" pageNumber="10">metaxa</emphasis>
</taxonomicName>
described by 
<bibRefCitation id="AE5B4B2FB475FFE5FACBFDB89A28FD11" author="Bouchet P." journalOrPublisher="Lavori della Societa Italiana di Malacologia" pageId="9" pageNumber="10" pagination="5 - 58" part="21" refId="ref20065" refString="Bouchet P. 1985. Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda). Lavori della Societa Italiana di Malacologia 21: 5 - 58." title="Les Triphoridae de Mediterranee et du proche Atlantique (Mollusca, Gastropoda)" type="journal article" year="1985">Bouchet (1985)</bibRefCitation>
shows slight differences compared to 
<taxonomicName id="0DCA4D5DB475FFE5FD05FD5C987EFD0E" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[731,858,661,687]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FD05FD5C99D7FD0E" box="[731,755,662,687]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FCDAFD5F987EFD0E" box="[772,858,661,687]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
, like a central tooth with four cusps (instead of five in 
<taxonomicName id="0DCA4D5DB475FFE5FEF1FD739A88FD73" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[303,428,696,723]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FEF1FD739A63FD73" box="[303,327,697,722]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FE88FD729A88FD73" box="[342,428,696,722]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
), lateral teeth and M1-M2 with five cusps (four in 
<taxonomicName id="0DCA4D5DB475FFE5FBDAFD739FA5FD73" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[1028,1153,696,723]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FBDAFD739F38FD73" box="[1028,1052,697,722]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FBF5FD729FA5FD73" box="[1067,1153,696,722]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
) and the presence of five marginal teeth (four in 
<taxonomicName id="0DCA4D5DB475FFE5FE27FD169950FD54" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[505,628,731,758]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FE27FD169935FD54" box="[505,529,732,757]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FDC0FD119950FD54" box="[542,628,731,757]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
). Radulae of both species are similar in general tooth morphology, with claw-like cusps, and in the number of cusps of the outermost marginal teeth (i.e., three cusps). The tooth morphology of 
<taxonomicName id="0DCA4D5DB475FFE5FE68FCE99917FC9D" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[438,563,802,828]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FE68FCE99AEAFC9D" box="[438,462,803,828]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FE03FCE89917FC9D" box="[477,563,802,828]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
is also quite similar to the two unnamed species from the southwestern Pacific studied by 
<bibRefCitation id="AE5B4B2FB475FFE5FE41FC8F996DFCFE" author="Nutzel A." box="[415,585,837,863]" journalOrPublisher="Berliner Geowissenschaftliche Abhandlungen" pageId="9" pageNumber="10" pagination="1 - 229" part="26" refId="ref21310" refString="Nutzel A. 1998. Ueber die Stammesgeschichte der Ptenoglossa (Gastropoda). Berliner Geowissenschaftliche Abhandlungen, ser. E (Palaeobiologie) 26: 1 - 229." title="Ueber die Stammesgeschichte der Ptenoglossa (Gastropoda)" type="journal article" year="1998">Nützel (1998)</bibRefCitation>
, especially to 
<taxonomicName id="0DCA4D5DB475FFE5FD25FC8C987FFCFE" authorityName="Monterosato" authorityYear="1884" box="[763,859,838,863]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB475FFE5FD25FC8C987FFCFE" box="[763,859,838,863]" italics="true" pageId="9" pageNumber="10">Metaxia</emphasis>
</taxonomicName>
sp. 1, albeit these two species have six and five marginal teeth (but four in 
<taxonomicName id="0DCA4D5DB475FFE5FD99FCA399ECFC23" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[583,712,872,899]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FD99FCA3997BFC23" box="[583,607,873,898]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FDACFCA299ECFC23" box="[626,712,872,898]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
). The predominance of rectangular scales in the jaw of 
<taxonomicName id="0DCA4D5DB475FFE5FF63FC469A39FC04" authorityName="Monterosato" authorityYear="1884" box="[189,285,908,933]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="F8BEEACCB475FFE5FF63FC469A39FC04" box="[189,285,908,933]" italics="true" pageId="9" pageNumber="10">Metaxia</emphasis>
</taxonomicName>
sp. 1 looks identical to the observed in 
<taxonomicName id="0DCA4D5DB475FFE5FD31FC469848FC04" authorityName="Faber &amp; Moolenbeek" authorityYear="1991" box="[751,876,907,934]" class="Gastropoda" family="Triphoridae" genus="Metaxia" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="excelsa">
<emphasis id="F8BEEACCB475FFE5FD31FC469823FC04" box="[751,775,908,933]" italics="true" pageId="9" pageNumber="10">M</emphasis>
. 
<emphasis id="F8BEEACCB475FFE5FCC8FC419848FC04" box="[790,876,907,933]" italics="true" pageId="9" pageNumber="10">excelsa</emphasis>
</taxonomicName>
(
<figureCitation id="52F12A5BB475FFE5FCA2FC4198FCFC04" box="[892,984,907,933]" captionStart="Fig" captionStartId="8.[189,232,1923,1949]" captionTargetBox="[206,1381,265,1883]" captionTargetId="figure@8.[206,1381,265,1883]" captionTargetPageId="8" captionText="Fig. 3. Metaxia excelsa Faber &amp; Moolenbeek, 1991. A–G. MZSP 122353. A. External morphology. B–C. Jaw, outer side. D–G. Radula. Scale bars: A = 1 mm; B = 50 µm; C = 2 µm; D–F = 10 µm; G = 5 µm." httpUri="https://zenodo.org/record/2647897/files/figure.png" pageId="9" pageNumber="10">Fig. 3B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>