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<document ID-DOI="10.11646/zootaxa.5239.2.6" ID-GBIF-Dataset="3f730d5b-0eac-441c-a968-a13dd754ccc3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7624372" ID-ZooBank="0AE5096C-B891-4C51-AEBD-B18EC489E41A" checkinTime="1675933891016" checkinUser="plazi" docAuthor="Gao, Hao-Ran &amp; Li, Yong-He" docDate="2023" docId="2E54045D816DFFE019A0FCAC66CEFF39" docLanguage="en" docName="zootaxa.5239.2.6.pdf" docOrigin="Zootaxa 5239 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spinomarmessoidea damingenis Gao &amp; Li 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="285" masterDocId="D26D7C25816FFFE51937FF8B6457FFEC" masterDocTitle="First report of the genus Spinomarmessoidea (Phasmatodea, Lonchodidae Necrosciinae) from China, with the description of a new species" masterLastPageNumber="288" masterPageNumber="280" pageNumber="282" updateTime="1675934553992" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>First report of the genus Spinomarmessoidea (Phasmatodea, Lonchodidae Necrosciinae) from China, with the description of a new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gao, Hao-Ran</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5093-0474</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, Yunnan Agricultural University, Kunming 650201, China. &amp; gaohaoran 909 @ qq. com; https: // orcid. org / 0000 - 0001 - 5093 - 0474</mods:affiliation>
<mods:nameIdentifier type="email">gaohaoran909@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Yong-He</mods:namePart>
<mods:affiliation>College of Plant Protection, Yunnan Agricultural University, Kunming 650201, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5239</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>280</mods:start>
<mods:end>288</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5239.2.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5239.2.6</mods:identifier>
<mods:identifier type="GBIF-Dataset">3f730d5b-0eac-441c-a968-a13dd754ccc3</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7624372</mods:identifier>
<mods:identifier type="ZooBank">0AE5096C-B891-4C51-AEBD-B18EC489E41A</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7624370" ID-Zenodo-Dep="7624370" LSID="urn:lsid:plazi:treatment:2E54045D816DFFE019A0FCAC66CEFF39" httpUri="http://treatment.plazi.org/id/2E54045D816DFFE019A0FCAC66CEFF39" lastPageId="5" lastPageNumber="285" pageId="2" pageNumber="282">
<subSubSection box="[151,642,807,834]" pageId="2" pageNumber="282" type="nomenclature">
<paragraph blockId="2.[151,642,807,871]" box="[151,642,807,834]" pageId="2" pageNumber="282">
<heading bold="true" box="[151,642,807,834]" fontSize="11" level="1" pageId="2" pageNumber="282" reason="1">
<emphasis bold="true" box="[151,642,807,834]" pageId="2" pageNumber="282">
<taxonomicName authority="Gao &amp; Li, 2023" authorityName="Gao &amp; Li" authorityYear="2023" box="[151,541,807,834]" class="Insecta" family="Lonchodidae" genus="Spinomarmessoidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="282" phylum="Arthropoda" rank="species" species="damingenis" status="sp. nov.">
<emphasis bold="true" box="[151,541,807,834]" italics="true" pageId="2" pageNumber="282">Spinomarmessoidea damingenis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[549,642,808,834]" pageId="2" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,322,844,871]" pageId="2" pageNumber="282" type="description">
<paragraph blockId="2.[151,642,807,871]" box="[151,322,844,871]" pageId="2" pageNumber="282">
(
<figureCitation box="[161,268,844,870]" captionStart-0="FIGURES 2" captionStart-1="FIGURES 3" captionStart-2="FIGURES 4" captionStartId-0="3.[151,264,1975,1999]" captionStartId-1="4.[151,264,1256,1280]" captionStartId-2="5.[151,264,811,835]" captionTargetBox-0="[161,1421,787,1941]" captionTargetBox-1="[213,1372,188,1223]" captionTargetBox-2="[210,1378,249,778]" captionTargetId-0="figure-273@3.[151,1436,777,1952]" captionTargetId-1="figure-17@4.[201,1387,181,1232]" captionTargetId-2="figure-51@5.[199,1389,237,787]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 2. Spinomarmessoidea damingensis sp. nov., female, type materials. A. holotype habitus, dorsal view; B. holotype habitus, lateral view; C. holotype habitus, ventral view; D. paratype habitus, dorsal view." captionText-1="FIGURES 3. Spinomarmessoidea damingensis sp. nov., female, holotype. A. tegmen; B. hind wing; C. terminalia, dorsal view; D. terminalia, ventral view; E. terminalia, lateral view." captionText-2="FIGURES 4. Spinomarmessoidea damingensis sp. nov., egg. A. dorsal; B. ventral." figureDoi-0="http://doi.org/10.5281/zenodo.7624377" figureDoi-1="http://doi.org/10.5281/zenodo.7624379" figureDoi-2="http://doi.org/10.5281/zenodo.7624381" httpUri-0="https://zenodo.org/record/7624377/files/figure.png" httpUri-1="https://zenodo.org/record/7624379/files/figure.png" httpUri-2="https://zenodo.org/record/7624381/files/figure.png" pageId="2" pageNumber="282">Figs. 2–4</figureCitation>
,
<figureCitation box="[279,314,844,871]" captionStart="FIGURES 5" captionStartId="5.[151,264,1971,1995]" captionTargetBox="[214,1374,859,1947]" captionTargetId="figure-66@5.[214,1374,859,1947]" captionTargetPageId="5" captionText="FIGURES 5. Spinomarmessoidea damingensis sp. nov.. A. Living photo (in the lab); B. habitat, Damingshan Nature Reserve, Guangxi." figureDoi="http://doi.org/10.5281/zenodo.7624383" httpUri="https://zenodo.org/record/7624383/files/figure.png" pageId="2" pageNumber="282">5A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="282" type="materials_examined">
<paragraph blockId="2.[151,1437,916,2023]" pageId="2" pageNumber="282">
<emphasis bold="true" box="[151,273,916,942]" pageId="2" pageNumber="282">Materials.</emphasis>
<materialsCitation collectingDate="2021-10-04" collectionCode="YNAU" country="China" county="ZhuangAutonomous Region" latitude="23.469168" location="Wuming District" longLatPrecision="21" longitude="108.43611" municipality="Nanning City" pageId="2" pageNumber="282" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis bold="true" box="[277,393,916,942]" pageId="2" pageNumber="282">
<typeStatus box="[277,388,916,942]" pageId="2" pageNumber="282">Holotype</typeStatus>
.
</emphasis>
<specimenCount box="[396,431,916,942]" pageId="2" pageNumber="282" type="female">1♀</specimenCount>
,
<collectingCountry box="[441,511,916,942]" name="China" pageId="2" pageNumber="282">China</collectingCountry>
,
<collectingRegion box="[518,614,916,942]" country="China" name="Guangxi" pageId="2" pageNumber="282">Guangxi</collectingRegion>
<collectingCounty box="[618,937,916,942]" pageId="2" pageNumber="282">ZhuangAutonomous Region</collectingCounty>
,
<collectingMunicipality box="[944,1093,916,942]" pageId="2" pageNumber="282">Nanning City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2E54045D816DFFE019A0FCAC66CEFF39:A322E390816DFFE71D7BFC1F6150FC42" box="[1100,1287,916,942]" country="China" county="ZhuangAutonomous Region" latitude="23.469168" longLatPrecision="21" longitude="108.43611" municipality="Nanning City" name="Wuming District" pageId="2" pageNumber="282" stateProvince="Guangxi">Wuming District</location>
,
<location LSID="urn:lsid:plazi:treatment:2E54045D816DFFE019A0FCAC66CEFF39:A322E390816DFFE71C38FC1F651DFC3F" country="China" county="ZhuangAutonomous Region" latitude="23.469168" longLatPrecision="21" longitude="108.43611" municipality="Nanning City" name="Damingshan Nature Reserve" pageId="2" pageNumber="282" stateProvince="Guangxi">Damingshan Nature Reserve</location>
[
<emphasis bold="true" box="[352,576,953,976]" pageId="2" pageNumber="282">kffiƜÞṂṘṁffl</emphasis>
],
<geoCoordinate box="[604,741,953,980]" degrees="23" direction="north" minutes="28" orientation="latitude" pageId="2" pageNumber="282" precision="15" seconds="09" value="23.469168">23°28′09″N</geoCoordinate>
,
<geoCoordinate box="[757,904,953,980]" degrees="108" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="282" precision="15" seconds="10" value="108.43611">108°26′10″E</geoCoordinate>
,
<date box="[921,1027,952,978]" pageId="2" pageNumber="282" value="2021-10-04">
<collectingDate box="[921,1027,952,978]" pageId="2" pageNumber="282" value="2021-10-04">4.X.2021</collectingDate>
</date>
,
<location LSID="urn:lsid:plazi:treatment:2E54045D816DFFE019A0FCAC66CEFF39:A322E390816DFFE71D23FC326039FC3F" box="[1044,1134,953,979]" country="China" county="ZhuangAutonomous Region" latitude="23.469168" longLatPrecision="21" longitude="108.43611" municipality="Nanning City" name="Hao Lu" pageId="2" pageNumber="282" stateProvince="Guangxi">Hao Lu</location>
(
<collectionCode box="[1155,1242,952,978]" pageId="2" pageNumber="282">YNAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="2021-08-23" collectionCode="MT, YNAU" collectorName="Shang-Qia Fu &amp; Ning Huang" country="China" county="Autonomous Region" location="Wuming District" municipality="Nanning City" pageId="2" pageNumber="282" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="paratype">
<emphasis bold="true" box="[1266,1382,952,978]" pageId="2" pageNumber="282">
<typeStatus box="[1266,1377,952,978]" pageId="2" pageNumber="282">Paratype</typeStatus>
.
</emphasis>
<specimenCount box="[1394,1429,952,978]" pageId="2" pageNumber="282" type="female">1♀</specimenCount>
,
<collectingCountry box="[151,221,988,1014]" name="China" pageId="2" pageNumber="282">China</collectingCountry>
,
<collectingRegion box="[232,422,988,1014]" country="China" name="Guangxi" pageId="2" pageNumber="282">Guangxi Zhuang</collectingRegion>
<collectingCounty box="[427,663,988,1014]" pageId="2" pageNumber="282">Autonomous Region</collectingCounty>
,
<collectingMunicipality box="[675,827,988,1014]" pageId="2" pageNumber="282">Nanning City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2E54045D816DFFE019A0FCAC66CEFF39:A322E390816DFFE71A70FC576051FC1A" box="[839,1030,988,1014]" country="China" county="Autonomous Region" municipality="Nanning City" name="Wuming District" pageId="2" pageNumber="282" stateProvince="Guangxi">Wuming District</location>
,
<location LSID="urn:lsid:plazi:treatment:2E54045D816DFFE019A0FCAC66CEFF39:A322E390816DFFE71D26FC576104FC1B" box="[1041,1363,988,1015]" country="China" county="Autonomous Region" municipality="Nanning City" name="Damingshan Nature Reserve" pageId="2" pageNumber="282" stateProvince="Guangxi">Damingshan Nature Reserve</location>
[
<emphasis bold="true" pageId="2" pageNumber="282">
kffi
<collectionCode box="[151,204,1025,1048]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="2" pageNumber="282">ƜÞ</collectionCode>
ṂṘṁffl
</emphasis>
],
<date box="[343,492,1024,1050]" pageId="2" pageNumber="282" value="2021-08-23">
<collectingDate box="[343,492,1024,1050]" pageId="2" pageNumber="282" value="2021-08-23">23.VIII.2021</collectingDate>
</date>
,
<collectorName box="[503,659,1024,1051]" pageId="2" pageNumber="282">Shang-Qia Fu</collectorName>
&amp;
<collectorName box="[695,833,1024,1051]" pageId="2" pageNumber="282">Ning Huang</collectorName>
(
<collectionCode box="[847,934,1024,1050]" pageId="2" pageNumber="282">YNAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[151,1437,916,2023]" box="[199,807,1060,1087]" pageId="2" pageNumber="282">
<emphasis bold="true" box="[199,396,1060,1086]" pageId="2" pageNumber="282">
<typeStatus box="[199,391,1060,1086]" isOtherTypeStatus="true" pageId="2" pageNumber="282">Other materials</typeStatus>
.
</emphasis>
<specimenCount box="[403,478,1060,1087]" pageId="2" pageNumber="282" type="egg">2 eggs</specimenCount>
, dissected from the
<typeStatus box="[702,804,1060,1086]" pageId="2" pageNumber="282">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="282" type="diagnosis">
<paragraph blockId="2.[151,1437,916,2023]" pageId="2" pageNumber="282">
<emphasis bold="true" box="[199,322,1096,1122]" pageId="2" pageNumber="282">Diagnosis.</emphasis>
This species can first be related to the genera
<taxonomicName authorityName="Seow-Choen" authorityYear="2021" box="[830,1056,1096,1122]" class="Insecta" family="Lonchodidae" genus="Spinomarmessoidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="282" phylum="Arthropoda" rank="genus">
<emphasis box="[830,1056,1096,1122]" italics="true" pageId="2" pageNumber="282">Spinomarmessoidea</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Brunner" authorityYear="1893" box="[1108,1273,1096,1122]" class="Insecta" family="Diapheromeridae" genus="Marmessoidea" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="282" phylum="Arthropoda" rank="genus">
<emphasis box="[1108,1273,1096,1122]" italics="true" pageId="2" pageNumber="282">Marmessoidea</emphasis>
</taxonomicName>
by the general egg morphology while being separated from other genera of the subfamily. This new species can be distinguished from
<taxonomicName authorityName="Brunner" authorityYear="1893" box="[214,379,1168,1194]" class="Insecta" family="Diapheromeridae" genus="Marmessoidea" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="282" phylum="Arthropoda" rank="genus">
<emphasis box="[214,379,1168,1194]" italics="true" pageId="2" pageNumber="282">Marmessoidea</emphasis>
</taxonomicName>
by both sides of the mesonotum median line with aligned spines. Although the spines on the mesonotum in the new species are not as distinct and variable as in the other two species, and the two type-localities are far apart in distance, this feature alone is not sufficient to divide the new species into a new genus.
</paragraph>
<paragraph blockId="2.[151,1437,916,2023]" pageId="2" pageNumber="282">
The female of the new species is very similar to
<taxonomicName baseAuthorityName="Seow-Choen" baseAuthorityYear="2021" box="[749,824,1276,1302]" class="Insecta" family="Lonchodidae" genus="Spinomarmessoidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="282" phylum="Arthropoda" rank="species" species="raja">
<emphasis box="[749,824,1276,1302]" italics="true" pageId="2" pageNumber="282">S. raja</emphasis>
</taxonomicName>
. Neither of them has distinct and large spines on their heads. But they can still be distinguished by the difference in the spines on the mesothorax supination and profemora slightly longer than metafemora. The spines on the mesonotum of
<taxonomicName authorityName="Gao &amp; Li" authorityYear="2023" box="[916,1086,1348,1375]" class="Insecta" family="Lonchodidae" genus="Spinomarmessoidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="282" phylum="Arthropoda" rank="species" species="damingensis">
<emphasis box="[916,930,1348,1374]" italics="true" pageId="2" pageNumber="282">S</emphasis>
.
<emphasis box="[946,1086,1348,1374]" italics="true" pageId="2" pageNumber="282">damingensis</emphasis>
</taxonomicName>
are not swollen on a common base.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="282" type="etymology">
<paragraph blockId="2.[151,1437,916,2023]" pageId="2" pageNumber="282">
<emphasis bold="true" box="[199,334,1420,1446]" pageId="2" pageNumber="282">Etymology.</emphasis>
The new species is named after the type-locality, Mt. Daming, Wuming District,
<collectingRegion box="[1247,1437,1420,1446]" country="China" name="Guangxi" pageId="2" pageNumber="282">Guangxi Zhuang</collectingRegion>
Autonomous Region,
<collectingCountry box="[398,469,1456,1482]" name="China" pageId="2" pageNumber="282">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="284" pageId="2" pageNumber="282" type="description">
<paragraph blockId="2.[151,1437,916,2023]" box="[199,1384,1492,1518]" pageId="2" pageNumber="282">
<emphasis bold="true" box="[199,446,1492,1518]" pageId="2" pageNumber="282">Description. Female.</emphasis>
Small-sized. Robust. General coloration variegated with black and brown. Winged.
</paragraph>
<paragraph blockId="2.[151,1437,916,2023]" pageId="2" pageNumber="282">
<emphasis box="[199,267,1528,1554]" italics="true" pageId="2" pageNumber="282">Head.</emphasis>
Longer than wide. Oval and flat, with many tubercles generally in 4 rows. Ocelli absent. The midline appears as a black longitudinal stripe and is divided into two at the middle of the compound eyes. There are two black longitudinal stripes behind the compound eyes, the upper one parallel to the middle stripe and darker than the lower one. Compound eyes variegated in life and brown in death. Antennae filiform, longer than the apex of the forelegs, generally dark with some light bans; scapus flat; pedicellus cylindrical, slightly thicker than the rest of the segments.
</paragraph>
<paragraph blockId="2.[151,1437,916,2023]" pageId="2" pageNumber="282">
<emphasis box="[199,288,1744,1770]" italics="true" pageId="2" pageNumber="282">Thorax.</emphasis>
Pronotum general dark coloration, sparsely covered with many tiny granules; transverse and longitudinal sulci crossing at mid-area and distinct. There are many short spines on the mesonotum generally in 4 longitudinal rows; 8 spines that belong to the two rows near the middle largest; the spines gradually become smaller from the anterior to the posterior; the posterior spines gradually become small granules (
<figureCitation box="[1123,1214,1852,1878]" captionStart="FIGURES 2" captionStartId="3.[151,264,1975,1999]" captionTargetBox="[161,1421,787,1941]" captionTargetId="figure-273@3.[151,1436,777,1952]" captionTargetPageId="3" captionText="FIGURES 2. Spinomarmessoidea damingensis sp. nov., female, type materials. A. holotype habitus, dorsal view; B. holotype habitus, lateral view; C. holotype habitus, ventral view; D. paratype habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7624377" httpUri="https://zenodo.org/record/7624377/files/figure.png" pageId="2" pageNumber="282">Fig. 2C</figureCitation>
). The larger spines apex dark. Mesopleurae dark and relatively smooth, without distinct tubercles. Mesosternum covered with small tubercles. Metanotum smooth. Metapleurae and Metasternum light brown and sparsely covered with tubercles.
</paragraph>
<paragraph blockId="2.[151,1437,916,2023]" lastBlockId="3.[151,1437,151,753]" lastPageId="3" lastPageNumber="283" pageId="2" pageNumber="282">
<emphasis box="[199,310,1960,1986]" italics="true" pageId="2" pageNumber="282">Abdomen.</emphasis>
Dark and cylindrical. Tergites II–V almost equal in length. Tergite VI shorter and suddenly narrower than the preceding tergites. Sternite II with a distinct median carina. Anal segment triangular in dorsal view, with a distinct median carina. Praeopercular organ tongue-shaped, flat and indistinct (
<figureCitation box="[1021,1109,151,177]" captionStart="FIGURES 3" captionStartId="4.[151,264,1256,1280]" captionTargetBox="[213,1372,188,1223]" captionTargetId="figure-17@4.[201,1387,181,1232]" captionTargetPageId="4" captionText="FIGURES 3. Spinomarmessoidea damingensis sp. nov., female, holotype. A. tegmen; B. hind wing; C. terminalia, dorsal view; D. terminalia, ventral view; E. terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7624379" httpUri="https://zenodo.org/record/7624379/files/figure.png" pageId="3" pageNumber="283">Fig. 3D</figureCitation>
). Operculum dark and scoopshaped, apex blunt (
<figureCitation box="[372,459,187,213]" captionStart="FIGURES 3" captionStartId="4.[151,264,1256,1280]" captionTargetBox="[213,1372,188,1223]" captionTargetId="figure-17@4.[201,1387,181,1232]" captionTargetPageId="4" captionText="FIGURES 3. Spinomarmessoidea damingensis sp. nov., female, holotype. A. tegmen; B. hind wing; C. terminalia, dorsal view; D. terminalia, ventral view; E. terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7624379" httpUri="https://zenodo.org/record/7624379/files/figure.png" pageId="3" pageNumber="283">Fig. 3D</figureCitation>
). Cercus short, reaching the end of anal segment, cylindrical, setose, with rounded apex.
</paragraph>
<paragraph blockId="3.[151,1437,151,753]" pageId="3" pageNumber="283">
<emphasis box="[199,260,223,248]" italics="true" pageId="3" pageNumber="283">Legs.</emphasis>
Unarmed, dark with light banded. Profemora incurved basally. Tarsus is composed of 5 tarsomeres (the 4th tarsomere shortest): the basiprotarsomere is about the length of the next 4 tarsomeres combined; the basimesotarsomere and basimetatarsomere shorter than basiprotarsomere, and shorter than each next 4 tarsomeres combined.
</paragraph>
<paragraph blockId="3.[151,1437,151,753]" pageId="3" pageNumber="283">
<emphasis box="[199,267,367,392]" italics="true" pageId="3" pageNumber="283">Wings</emphasis>
. Tegmen dark brown with black veins. The gibbus rounded and raised. The costal area of hind wings generally dark and variegated with some yellow. The anal area of hind wings pale brown (
<figureCitation box="[1178,1266,403,429]" captionStart="FIGURES 3" captionStartId="4.[151,264,1256,1280]" captionTargetBox="[213,1372,188,1223]" captionTargetId="figure-17@4.[201,1387,181,1232]" captionTargetPageId="4" captionText="FIGURES 3. Spinomarmessoidea damingensis sp. nov., female, holotype. A. tegmen; B. hind wing; C. terminalia, dorsal view; D. terminalia, ventral view; E. terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7624379" httpUri="https://zenodo.org/record/7624379/files/figure.png" pageId="3" pageNumber="283">Fig. 3B</figureCitation>
). Apex of hind wings reaches the tergite IX.
</paragraph>
<paragraph blockId="3.[151,1437,151,753]" box="[199,392,475,501]" pageId="3" pageNumber="283">
<emphasis bold="true" box="[199,267,475,501]" pageId="3" pageNumber="283">Male.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="3.[151,1437,151,753]" pageId="3" pageNumber="283">
<emphasis bold="true" box="[199,253,511,537]" pageId="3" pageNumber="283">Egg.</emphasis>
The description of the eggs is based on
<specimenCount box="[723,825,511,537]" pageId="3" pageNumber="283" type="egg">two eggs</specimenCount>
dissected from the
<typeStatus box="[1053,1154,511,537]" pageId="3" pageNumber="283">holotype</typeStatus>
, which may not be fully formed.
</paragraph>
<paragraph blockId="3.[151,1437,151,753]" pageId="3" pageNumber="283">
Measurements [mm]: Length: 1.7; width: 1.0; height: 0.9. Capsule dark yellow, covered with tiny brown granules, rice-shaped, posterior pole rounded (
<figureCitation box="[686,756,619,645]" captionStart="FIGURES 4" captionStartId="5.[151,264,811,835]" captionTargetBox="[210,1378,249,778]" captionTargetId="figure-51@5.[199,1389,237,787]" captionTargetPageId="5" captionText="FIGURES 4. Spinomarmessoidea damingensis sp. nov., egg. A. dorsal; B. ventral." figureDoi="http://doi.org/10.5281/zenodo.7624381" httpUri="https://zenodo.org/record/7624381/files/figure.png" pageId="3" pageNumber="283">Fig. 4</figureCitation>
). Operculum rounded and brown, covered with tiny brown granules that are denser than other parts; a relatively larger elevation on the top of the center. Micropylar plate expands to the left and right sides with white edges, the middle expands up and down, and the two sides that expand out expand up and down again (
<figureCitation box="[510,600,727,753]" captionStart="FIGURES 4" captionStartId="5.[151,264,811,835]" captionTargetBox="[210,1378,249,778]" captionTargetId="figure-51@5.[199,1389,237,787]" captionTargetPageId="5" captionText="FIGURES 4. Spinomarmessoidea damingensis sp. nov., egg. A. dorsal; B. ventral." figureDoi="http://doi.org/10.5281/zenodo.7624381" httpUri="https://zenodo.org/record/7624381/files/figure.png" pageId="3" pageNumber="283">Fig. 4A</figureCitation>
). Plate gap white. Median line indistinct.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7624377" ID-Zenodo-Dep="7624377" httpUri="https://zenodo.org/record/7624377/files/figure.png" pageId="3" pageNumber="283" startId="3.[151,264,1975,1999]" targetBox="[161,1421,787,1941]" targetPageId="3">
<paragraph blockId="3.[151,1437,1975,2036]" pageId="3" pageNumber="283">
<emphasis bold="true" box="[151,289,1975,1999]" pageId="3" pageNumber="283">FIGURES 2.</emphasis>
<taxonomicName authority="Gao &amp; Li, 2023" authorityName="Gao &amp; Li" authorityYear="2023" box="[296,634,1975,1999]" class="Insecta" family="Lonchodidae" genus="Spinomarmessoidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="283" phylum="Arthropoda" rank="species" species="damingensis" status="sp. nov.">
<emphasis box="[296,634,1975,1999]" italics="true" pageId="3" pageNumber="283">Spinomarmessoidea damingensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[640,721,1975,1999]" pageId="3" pageNumber="283">
<taxonomicNameLabel box="[640,721,1975,1999]" pageId="3" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, type materials.
<emphasis bold="true" box="[973,998,1975,1999]" pageId="3" pageNumber="283">A.</emphasis>
holotype habitus, dorsal view;
<emphasis bold="true" box="[1318,1341,1975,1999]" pageId="3" pageNumber="283">B.</emphasis>
holotype habitus, lateral view;
<emphasis bold="true" box="[369,394,2011,2035]" pageId="3" pageNumber="283">C.</emphasis>
holotype habitus, ventral view;
<emphasis bold="true" box="[720,745,2011,2035]" pageId="3" pageNumber="283">D.</emphasis>
paratype habitus, dorsal view.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7624379" ID-Zenodo-Dep="7624379" httpUri="https://zenodo.org/record/7624379/files/figure.png" pageId="4" pageNumber="284" startId="4.[151,264,1256,1280]" targetBox="[213,1372,188,1223]" targetPageId="4">
<paragraph blockId="4.[151,1436,1256,1316]" pageId="4" pageNumber="284">
<emphasis bold="true" box="[151,288,1256,1280]" pageId="4" pageNumber="284">FIGURES 3.</emphasis>
<taxonomicName authority="Gao &amp; Li, 2023" authorityName="Gao &amp; Li" authorityYear="2023" box="[294,630,1256,1280]" class="Insecta" family="Lonchodidae" genus="Spinomarmessoidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="4" pageNumber="284" phylum="Arthropoda" rank="species" species="damingensis" status="sp. nov.">
<emphasis box="[294,630,1256,1280]" italics="true" pageId="4" pageNumber="284">Spinomarmessoidea damingensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[635,715,1256,1280]" pageId="4" pageNumber="284">
<taxonomicNameLabel box="[635,715,1256,1280]" pageId="4" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype.
<emphasis bold="true" box="[908,933,1256,1280]" pageId="4" pageNumber="284">A.</emphasis>
tegmen;
<emphasis bold="true" box="[1026,1049,1256,1280]" pageId="4" pageNumber="284">B.</emphasis>
hind wing;
<emphasis bold="true" box="[1168,1192,1256,1280]" pageId="4" pageNumber="284">C.</emphasis>
terminalia, dorsal view;
<emphasis bold="true" box="[151,176,1292,1316]" pageId="4" pageNumber="284">D.</emphasis>
terminalia, ventral view;
<emphasis bold="true" box="[437,460,1292,1316]" pageId="4" pageNumber="284">E.</emphasis>
terminalia, lateral view.
</paragraph>
</caption>
<paragraph blockId="4.[199,530,1362,1388]" box="[199,530,1362,1388]" pageId="4" pageNumber="284">
<emphasis bold="true" box="[199,382,1362,1388]" pageId="4" pageNumber="284">Measurements.</emphasis>
See
<emphasis bold="true" box="[436,530,1362,1388]" pageId="4" pageNumber="284">
<tableCitation box="[436,523,1362,1388]" captionStart="TABLE 1" captionStartId="4.[152,244,1436,1460]" captionText="TABLE 1. Measurements of S. damingensis sp. nov. Length (mm) Holotype (♀) Body 60.6" pageId="4" pageNumber="284">Table 1</tableCitation>
.
</emphasis>
</paragraph>
</subSubSection>
<caption pageId="4" pageNumber="284" startId="4.[152,244,1436,1460]">
<paragraph blockId="4.[152,721,1436,2023]" box="[152,721,1436,1460]" pageId="4" pageNumber="284">
<emphasis bold="true" box="[152,271,1436,1460]" pageId="4" pageNumber="284">TABLE 1.</emphasis>
Measurements of
<taxonomicName authority="Gao &amp; Li, 2023" authorityName="Gao &amp; Li" authorityYear="2023" box="[469,628,1436,1460]" class="Insecta" family="Lonchodidae" genus="Spinomarmessoidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="4" pageNumber="284" phylum="Arthropoda" rank="species" species="damingensis" status="sp. nov.">
<emphasis box="[469,628,1436,1460]" italics="true" pageId="4" pageNumber="284">S. damingensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[635,721,1436,1460]" pageId="4" pageNumber="284">
<taxonomicNameLabel box="[635,721,1436,1460]" pageId="4" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<table box="[166,1100,1476,2023]" gridcols="3" gridrows="14" pageId="4" pageNumber="284">
<tr box="[166,1100,1476,1501]" gridrow="0" pageId="4" pageNumber="284" rowspan-2="1">
<paragraph blockId="4.[152,721,1436,2023]" box="[166,664,1476,1500]" pageId="4" pageNumber="284">
<th box="[166,300,1476,1501]" gridcol="0" gridrow="0" pageId="4" pageNumber="284">Length (mm)</th>
<th box="[527,664,1476,1501]" gridcol="1" gridrow="0" pageId="4" pageNumber="284">Holotype (♀)</th>
</paragraph>
<paragraph blockId="4.[152,721,1436,2023]" box="[166,571,1516,1540]" pageId="4" pageNumber="284">
<tr box="[166,1100,1516,1540]" gridrow="1" pageId="4" pageNumber="284" rowspan-2="1">
<th box="[166,300,1516,1540]" gridcol="0" gridrow="1" pageId="4" pageNumber="284">Body</th>
<td box="[527,664,1516,1540]" gridcol="1" gridrow="1" pageId="4" pageNumber="284">60.6</td>
</tr>
</paragraph>
</tr>
</table>
</caption>
<subSubSection pageId="5" pageNumber="285" type="distribution">
<paragraph blockId="5.[199,1254,150,213]" box="[199,1254,150,177]" pageId="5" pageNumber="285">
<emphasis bold="true" box="[199,353,150,176]" pageId="5" pageNumber="285">Distribution.</emphasis>
<collectingCountry box="[359,426,151,177]" name="China" pageId="5" pageNumber="285">China</collectingCountry>
[
<collectingRegion box="[440,543,151,177]" country="China" name="Guangxi" pageId="5" pageNumber="285">Guangxi</collectingRegion>
: Damingshan (
<figureCitation box="[712,800,150,177]" captionStart="FIGURES 5" captionStartId="5.[151,264,1971,1995]" captionTargetBox="[214,1374,859,1947]" captionTargetId="figure-66@5.[214,1374,859,1947]" captionTargetPageId="5" captionText="FIGURES 5. Spinomarmessoidea damingensis sp. nov.. A. Living photo (in the lab); B. habitat, Damingshan Nature Reserve, Guangxi." figureDoi="http://doi.org/10.5281/zenodo.7624383" httpUri="https://zenodo.org/record/7624383/files/figure.png" pageId="5" pageNumber="285">Fig. 5B</figureCitation>
)]. Only known from type-locality so far.
</paragraph>
<paragraph blockId="5.[199,1254,150,213]" box="[199,665,186,213]" pageId="5" pageNumber="285">
<emphasis bold="true" box="[199,658,186,213]" pageId="5" pageNumber="285">Chinese common name. kffiḦṃHdz</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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