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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.430.8094" ID-PMC="PMC4141162" ID-Pensoft-Pub="1313-2970-430-1" ID-Pensoft-UUID="FFE6FF8CFFD43B66335AD2203774DB49" ID-PubMed="25152663" ID-Zenodo-Dep="578441" ID-ZooBank="86F760563B8B49FB9C86FAD0DB0CBE8C" ModsDocID="1313-2970-430-1" checkinTime="1451245551152" checkinUser="pensoft" docAuthor="Erwin, Terry L. & Zamorano, Laura S." docDate="2014" docId="2C9C4976BF58FCCE35067FD5B52362C8" docLanguage="en" docName="ZooKeys 430: 1-108" docOrigin="ZooKeys 430" docPubDate="2014-08-01" docSource="http://dx.doi.org/10.3897/zookeys.430.8094" docTitle="Asklepia laetitia Zamorano & Erwin 2014, sp. n." docType="treatment" docUuid="628C9676-9823-4241-AA36-3226C8561E7E" docUuidSource="ZooBank" docVersion="5" id="FFE6FF8CFFD43B66335AD2203774DB49" lastPageNumber="41" masterDocId="FFE6FF8CFFD43B66335AD2203774DB49" masterDocTitle="A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)" masterLastPageNumber="108" masterPageNumber="1" pageNumber="40" updateTime="1668159067277" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Erwin, Terry L.</mods:namePart>
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<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zamorano, Laura S.</mods:namePart>
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<mods:affiliation>Research Student, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA & Laboratorio de Zoologia Acuatica LAZOEA, Departamento de Ciencias Biologicas, Universidad de los Andes, Colombia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-08-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>430</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>108</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.430.8094</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.430.8094</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-430-1</mods:identifier>
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<mods:identifier type="ZooBank">86F760563B8B49FB9C86FAD0DB0CBE8C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFE6FF8CFFD43B66335AD2203774DB49</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578441</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152054776" LSID="urn:lsid:zoobank.org:act:628C9676-9823-4241-AA36-3226C8561E7E" httpUri="http://treatment.plazi.org/id/2C9C4976BF58FCCE35067FD5B52362C8" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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<subSubSection pageId="39" pageNumber="40" type="nomenclature">
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<paragraph pageId="39" pageNumber="40">
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<taxonomicName LSID="http://zoobank.org/628C9676-9823-4241-AA36-3226C8561E7E" authority="Zamorano & Erwin" authorityName="Zamorano & Erwin" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asklepia laetitia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="laetitia" status="sp. n.">Asklepia laetitia Zamorano & Erwin</taxonomicName>
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<taxonomicNameLabel pageId="39" pageNumber="40">sp. n.</taxonomicNameLabel>
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Colombian pattern-wing beetle
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<figureCitation captionStart="Figure 33–36" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="39" pageNumber="40">Figs 35</figureCitation>
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<figureCitation captionStart="Figure 57–62" captionStartId="F17" captionText="Figure 57 - 62. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 57 Asklepia geminata (Bates, 1871) ADP 109186, Rio Samiria, Boca Cano Ingles Camp, Peru 58 Asklepia campbellorum Zamorano & Erwin, sp. n. ADP 109196, 20 km SW Manaus, Brazil 59 Asklepia demiti Erwin & Zamorano, sp. n. ADP 132585, Rio Demiti, Brazil 60 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 052565, Rio Sucusari, Peru 61 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia. Legend: a apical area; bl basal lobe; bo basal orifice; lp left paramere; ml median lobe; om ostial membrane; oo ostial opening; rp right paramere; sh shaft; pb phallobase; ps phallobase shaft; ms medial spine; ds distal spine 62 Asklepia lebioides (Bates, 1871), comb. n., ADP 109208, Rio Demiti, Brazil. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30647" pageId="39" pageNumber="40">, 61</figureCitation>
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<figureCitation captionStart="Figure 76–77" captionStartId="F21" captionText="Figure 76 - 77. 76 Distribution map for known localities of Asklepia geminata (Bates) 77 Distribution map for known localities of Asklepia species of the hilaris group." httpUri="https://binary.pensoft.net/fig/30651" pageId="39" pageNumber="40">, 77</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="40" type="holotype">
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<paragraph pageId="39" pageNumber="40">Holotype.</paragraph>
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<paragraph pageId="39" pageNumber="40">
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<emphasis bold="true" pageId="39" pageNumber="40">Colombia</emphasis>
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, Amazonas, Leticia,
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<geoCoordinate degrees="4.1896" direction="south" orientation="latitude" precision="5" value="-4.1896">4.1896°S</geoCoordinate>
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,
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<geoCoordinate degrees="69.9711" direction="west" orientation="longitude" precision="5" value="-69.9711">69.9711°W</geoCoordinate>
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, 72m, no date, (F.M. Oliveira, P. Wygodzinsky)(AMNH: ADP109190, male).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="40" type="derivation of specific epithet">
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<paragraph pageId="39" pageNumber="40">Derivation of specific epithet.</paragraph>
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<paragraph pageId="39" pageNumber="40">
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The specific epithet,
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<emphasis italics="true" pageId="39" pageNumber="40">laetitia</emphasis>
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, from the Latin greeting
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<emphasis italics="true" pageId="39" pageNumber="40">laetitia</emphasis>
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used in the nominative case, is a singular feminine noun in apposition, based on the name of the town near which these beetles were found, and meaning happiness, joy, gladness, and delight.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="40" type="proposed english vernacular name">
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<paragraph pageId="39" pageNumber="40">Proposed english vernacular name.</paragraph>
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<paragraph pageId="39" pageNumber="40">Colombian pattern-wing beetle.</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="40" type="diagnosis">
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<paragraph pageId="39" pageNumber="40">Diagnosis.</paragraph>
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<paragraph pageId="39" pageNumber="40">
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With the attributes of the genus
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<taxonomicName authorityName="Liebke" authorityYear="1938" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="39" pageNumber="40">Asklepia</emphasis>
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</taxonomicName>
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as described by
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<bibRefCitation author="Liebke, M" journalOrPublisher="Embrik Strand" pageId="81" pageNumber="82" pagination="37 - 141" refId="B54" refString="Liebke, M, 1938. Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr. Embrik Strand 4: 37 - 141" title="Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr." volume="4" year="1938">Liebke (1938)</bibRefCitation>
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and as noted above under the generic diagnosis, and small in size for the genus (SBL = 2.494 mm). Adults with head fuscous, prothorax brunneus, elytral maculae aurantiacus; elytron brunneus with a broad rectangular-shaped aurantiacus macula crossing basal and medial lateral and proximal quadrants, macula not extended to lateral margin or sutural area and, a rounded flavous macula in the apical proximal quadrant, sutural area fulvous; metasternum, abdominal sterna III-VI, and epipleuron testaceous, abdominal sternum VII paler; legs testaceous; antennal scape, pedicel, and antennomere 3 testaceous, antennomeres 4-7 deeply infuscated, 8-11 white. Dorsal surface devoid of microsculpture, surface luster very shiny. Pronotum cordiform, narrowly explanate; anterior angles feebly produced; lateral margin beaded; hind angle angulate, moderately prominent; median line well defined; basal and apical transverse impression punctate, punctures infuscated. Elytral interneurs evident as rows of continuous coarse punctures closely spaced.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
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<paragraph pageId="39" pageNumber="40">Description.</paragraph>
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<paragraph lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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(
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<figureCitation captionStart="Figure 33–36" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="39" pageNumber="40">Fig. 35</figureCitation>
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,
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<figureCitation captionStart="Figure 57–62" captionStartId="F17" captionText="Figure 57 - 62. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 57 Asklepia geminata (Bates, 1871) ADP 109186, Rio Samiria, Boca Cano Ingles Camp, Peru 58 Asklepia campbellorum Zamorano & Erwin, sp. n. ADP 109196, 20 km SW Manaus, Brazil 59 Asklepia demiti Erwin & Zamorano, sp. n. ADP 132585, Rio Demiti, Brazil 60 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 052565, Rio Sucusari, Peru 61 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia. Legend: a apical area; bl basal lobe; bo basal orifice; lp left paramere; ml median lobe; om ostial membrane; oo ostial opening; rp right paramere; sh shaft; pb phallobase; ps phallobase shaft; ms medial spine; ds distal spine 62 Asklepia lebioides (Bates, 1871), comb. n., ADP 109208, Rio Demiti, Brazil. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30647" pageId="39" pageNumber="40">61</figureCitation>
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).
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">
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<pageBreakToken pageId="40" pageNumber="41" start="start">Habitus</pageBreakToken>
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</emphasis>
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:
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</emphasis>
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(
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<figureCitation captionStart="Figure 33–36" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="40" pageNumber="41">Fig. 35</figureCitation>
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).
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">Size</emphasis>
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:
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</emphasis>
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[See also
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<tableCitation captionStart="Table 7" captionStartId="T7" captionText="Table 7. Asklepia laetitia Zamorano & Erwin, sp. n." httpUri="http://table.plazi.org/id/FBBA69A552A4641E04E5E5A1D4966590" pageId="40" pageNumber="41" tableUuid="FBBA69A552A4641E04E5E5A1D4966590">Table 7</tableCitation>
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] Small-size for the genus; ABL = 2.99 mm, SBL = 2.494 mm, TW (total width) 1.463 mm, LP = 0.489 mm, WP = 0.706 mm, LE = 1.633 mm.
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">Color</emphasis>
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:
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</emphasis>
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See diagnosis above.
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">Luster</emphasis>
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:
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</emphasis>
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See diagnosis above.
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<emphasis bold="true" italics="true" pageId="40" pageNumber="41">Head</emphasis>
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(
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<figureCitation captionStart="Figure 33–36" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="40" pageNumber="41">Fig. 35</figureCitation>
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): as in description for genus above.
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">Prothorax</emphasis>
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.
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</emphasis>
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Pronotum (
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<figureCitation captionStart="Figure 33–36" captionStartId="F10" captionText="Figure 33 - 36. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 33 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 133151, Pithecia, Peru 34 Asklepia hilaris (Bates, 1871), comb. n., Brazil ADP 132543, Sao Paulo de Olivenca, Brazil 35 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia 36 Asklepia lebioides (Bates, 1871), comb. n., ADP 109194, Santarem, Rio Tapajos, Brazil. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30640" pageId="40" pageNumber="41">Fig. 35</figureCitation>
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) moderately broad, as wide as head across eyes (WH/WP: 1.032), longer than head (LP/LH: 1.316), wider than long (W/L: 1.444); markedly cordiform and explanate, lateral margin beaded with seta at anterior third; apical margin straight; base markedly constricted; hind angle moderately produced and setose; median line well defined, basal and apical transverse impression punctate, punctures infuscated; surface smooth throughout.
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">Pterothorax</emphasis>
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.
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</emphasis>
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Normal for genus, see description for genus above. Elytra slightly convex; at apical third markedly wider than head across eyes (WH/TW: 0.468) and twice as wide as pronotum (WP/TW: 0.504). Elytral interneurs evident as rows of continuous, coarse punctures widely spaced, coarse punctures with infuscated halo in the proximal basal and apical quadrants. Hind wings fully developed.
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">Legs</emphasis>
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.
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</emphasis>
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Overall, normal for genus, see description for genus above.
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<emphasis bold="true" pageId="40" pageNumber="41">
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<emphasis italics="true" pageId="40" pageNumber="41">Abdominal sterna</emphasis>
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.
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</emphasis>
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Overall, normal for genus, see description for genus above.
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<emphasis bold="true" italics="true" pageId="40" pageNumber="41">Male genitalia</emphasis>
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(
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<figureCitation captionStart="Figure 57–62" captionStartId="F17" captionText="Figure 57 - 62. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 57 Asklepia geminata (Bates, 1871) ADP 109186, Rio Samiria, Boca Cano Ingles Camp, Peru 58 Asklepia campbellorum Zamorano & Erwin, sp. n. ADP 109196, 20 km SW Manaus, Brazil 59 Asklepia demiti Erwin & Zamorano, sp. n. ADP 132585, Rio Demiti, Brazil 60 Asklepia grammechrysea Zamorano & Erwin, sp. n. ADP 052565, Rio Sucusari, Peru 61 Asklepia laetitia Zamorano & Erwin, sp. n. ADP 109190, Leticia, Colombia. Legend: a apical area; bl basal lobe; bo basal orifice; lp left paramere; ml median lobe; om ostial membrane; oo ostial opening; rp right paramere; sh shaft; pb phallobase; ps phallobase shaft; ms medial spine; ds distal spine 62 Asklepia lebioides (Bates, 1871), comb. n., ADP 109208, Rio Demiti, Brazil. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30647" pageId="40" pageNumber="41">Fig. 61</figureCitation>
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). Median lobe (
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<emphasis bold="true" pageId="40" pageNumber="41">ml</emphasis>
|
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|
|
) with phallobase (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">pb</emphasis>
|
|||
|
|
) of moderate length about a fourth the length of shaft (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">ps</emphasis>
|
|||
|
|
), basal opening (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">bo</emphasis>
|
|||
|
|
) large, oriented parallel to
|
|||
|
|
<normalizedToken originalValue="shaft’s">shaft's</normalizedToken>
|
|||
|
|
apical third. Shaft broad, moderately curved ventrally, dorsally sclerotized except for short ostium (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">oo</emphasis>
|
|||
|
|
,
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">om</emphasis>
|
|||
|
|
); in ventral aspect tapered toward rather rounded apex (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">a</emphasis>
|
|||
|
|
), in lateral aspect, a narrowly rounded apex. Left paramere (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">lp</emphasis>
|
|||
|
|
) very large and broad, right (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">rp</emphasis>
|
|||
|
|
) small and triangular, apex of left paramere lobate and much longer than right paramere, about half the length of shaft (measured in left lateral aspect). Endophallus with 5 medial spines (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">ms</emphasis>
|
|||
|
|
), and one very large distal spine (
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">ds</emphasis>
|
|||
|
|
).
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">
|
|||
|
|
<emphasis italics="true" pageId="40" pageNumber="41">Female genitalia</emphasis>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Not investigated, presumably similar to that of
|
|||
|
|
<taxonomicName authorityName="Erwin & Zamorano" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="demiti">
|
|||
|
|
<emphasis italics="true" pageId="40" pageNumber="41">Asklepia demiti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis bold="true" pageId="40" pageNumber="41">sp. n.</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="40" pageNumber="41" type="dispersal potential">
|
|||
|
|
<paragraph pageId="40" pageNumber="41">Dispersal potential.</paragraph>
|
|||
|
|
<paragraph pageId="40" pageNumber="41">These beetles are macropterous and probably capable of flight. They are moderately swift and agile runners.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="40" pageNumber="41" type="distribution">
|
|||
|
|
<paragraph pageId="40" pageNumber="41">Distribution.</paragraph>
|
|||
|
|
<paragraph pageId="40" pageNumber="41">
|
|||
|
|
(
|
|||
|
|
<figureCitation captionStart="Figure 76–77" captionStartId="F21" captionText="Figure 76 - 77. 76 Distribution map for known localities of Asklepia geminata (Bates) 77 Distribution map for known localities of Asklepia species of the hilaris group." httpUri="https://binary.pensoft.net/fig/30651" pageId="40" pageNumber="41">Fig. 77</figureCitation>
|
|||
|
|
). This species has been found at only one location on the white-water system of the Amazon River drainage system. But that does not at all indicate its actual distribution, as has been pointed out above, very small beetles are inadequately sampled, especially in the Neotropics.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="40" pageNumber="41" type="way of life">
|
|||
|
|
<paragraph pageId="40" pageNumber="41">Way of life.</paragraph>
|
|||
|
|
<paragraph pageId="40" pageNumber="41">
|
|||
|
|
See
|
|||
|
|
<bibRefCitation author="Erwin, TL" journalOrPublisher="Revista Peruana de Entomologia" pageId="79" pageNumber="80" pagination="1 - 85" refId="B22" refString="Erwin, TL, 1991. Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru. Revista Peruana de Entomologia 33: 1 - 85" title="Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru." volume="33" year="1991">Erwin (1991)</bibRefCitation>
|
|||
|
|
for a general description of the genus. No way of life information is available for this species other than they occur in lowland Amazonian rainforest.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="40" pageNumber="41" type="other specimens examined">
|
|||
|
|
<paragraph pageId="40" pageNumber="41">Other specimens examined.</paragraph>
|
|||
|
|
<paragraph pageId="40" pageNumber="41">None.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|