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<document id="AC284EB54E9EA42375F91DFC534A0AD0" ID-CLB-Dataset="198847" ID-DOI="10.11646/zootaxa.5213.5.1" ID-GBIF-Dataset="2a26dfdb-0e17-40a1-b5fd-14787882b9d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7381607" ID-ZooBank="B16F6ED4-A8C2-4F3D-A5DD-20537DE0CF01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1669810135404" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="03FFB25377687A4DDD90F9416BFC53BA" docLanguage="en" docName="zootaxa.5213.5.1.pdf" docOrigin="Zootaxa 5213 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geodromicus malcolmi Shavrin 2022, sp.n." docType="treatment" docVersion="7" lastPageNumber="483" masterDocId="FFC6CA2B77757A6DDD07FFEE68335217" masterDocTitle="The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="496" masterPageNumber="451" pageNumber="480" updateTime="1699412587891" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="615AA06E9FC2CD161F20FFCF2C30C3EE">The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<mods:namePart id="22E0ADC840D213F9DC7D521D196C157E">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="B80D1C7412660278703FEF79B0A41ECE">Zootaxa</mods:title>
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<mods:part id="33F0A4AB2193FEF9DD666DAFC7D4B18C">
<mods:date id="770EAAF8B4F760CD6E1BCAA8DBEE1EAC">2022</mods:date>
<mods:detail id="E79685B606896DA8A31A1C344914DFF9" type="pubDate">
<mods:number id="1DDADC829D0FB5DE495A383E31E18A11">2022-11-29</mods:number>
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<mods:number id="C6F64FC8A1EC8C8E97F82B8D4113D11F">5213</mods:number>
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<mods:classification id="3B58664ABCF0826D4DCEFCEAD5A91FAD">journal article</mods:classification>
<mods:identifier id="A936F62A9AEB49A4CC96C5EC3CA90904" type="CLB-Dataset">198847</mods:identifier>
<mods:identifier id="A8FD128E23D8CC9B31CB34EA50B37F36" type="DOI">10.11646/zootaxa.5213.5.1</mods:identifier>
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<mods:identifier id="66647041CE12F28A791912AA6E7F7337" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FFB25377687A4DDD90F9416BFC53BA" ID-DOI="http://doi.org/10.5281/zenodo.7383609" ID-GBIF-Taxon="205348416" ID-Zenodo-Dep="7383609" LSID="urn:lsid:plazi:treatment:03FFB25377687A4DDD90F9416BFC53BA" httpUri="http://treatment.plazi.org/id/03FFB25377687A4DDD90F9416BFC53BA" lastPageId="32" lastPageNumber="483" pageId="29" pageNumber="480">
<subSubSection id="C34C50CE77687A70DD90F9416A8A54DD" box="[151,697,1711,1738]" pageId="29" pageNumber="480" type="nomenclature">
<paragraph id="8BE9034577687A70DD90F9416A8A54DD" blockId="29.[151,697,1711,1775]" box="[151,697,1711,1738]" pageId="29" pageNumber="480">
<heading id="D0A1B42977687A70DD90F9416A8A54DD" bold="true" box="[151,697,1711,1738]" fontSize="11" level="1" pageId="29" pageNumber="480" reason="1">
<emphasis id="B922DF5777687A70DD90F9416A8A54DD" bold="true" box="[151,697,1711,1738]" pageId="29" pageNumber="480">
<taxonomicName id="4C5678C677687A70DD90F9416A4254DD" authority="Shavrin" authorityName="Shavrin" authorityYear="2022" box="[151,625,1711,1738]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="malcolmi" status="sp. nov.">
<emphasis id="B922DF5777687A70DD90F941690B54DD" bold="true" box="[151,312,1711,1738]" italics="true" pageId="29" pageNumber="480">Geodromicus</emphasis>
(s.str.)
<emphasis id="B922DF5777687A70DC93F9416A3754DD" bold="true" box="[404,516,1711,1738]" italics="true" pageId="29" pageNumber="480">malcolmi</emphasis>
Shavrin
</taxonomicName>
,
<taxonomicNameLabel id="A211622C77687A70DF79F95E6A8A54DD" box="[638,697,1712,1738]" pageId="29" pageNumber="480" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77687A70DD90F93A69B154F8" box="[151,386,1748,1775]" pageId="29" pageNumber="480" type="description">
<paragraph id="8BE9034577687A70DD90F93A69B154F8" blockId="29.[151,697,1711,1775]" box="[151,386,1748,1775]" pageId="29" pageNumber="480">
(
<figureCitation id="136D1FC077687A70DD98F93A68CE54F9" box="[159,253,1748,1774]" captionStart="FIGURE 27" captionStartId="13.[151,250,1984,2008]" captionTargetBox="[193,1395,1150,1960]" captionTargetId="figure-31@13.[193,1395,1150,1960]" captionTargetPageId="13" captionText="FIGURE 27. Distribution of Geodromicus championi (circles) and G. malcolmi (square) in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.7381624" httpUri="https://zenodo.org/record/7381624/files/figure.png" pageId="29" pageNumber="480">Figs. 27</figureCitation>
,
<figureCitation id="136D1FC077687A70DC0EF93A691454F9" box="[265,295,1748,1774]" captionStart="FIGURES 57–62" captionStartId="30.[151,264,1878,1902]" captionTargetBox="[166,1428,284,1841]" captionTargetId="figure-70@30.[151,1436,273,1855]" captionTargetPageId="30" captionText="FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381644" httpUri="https://zenodo.org/record/7381644/files/figure.png" pageId="29" pageNumber="480">60</figureCitation>
,
<figureCitation id="136D1FC077687A70DC34F93A694854F8" box="[307,379,1748,1775]" captionStart="FIGURES 64–69" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[173,1414,192,1907]" captionTargetId="figure-15@33.[159,1428,181,1919]" captionTargetPageId="33" captionText="FIGURES 64–69. Aedeagus of Geodromicus malcolmi (Figs. 64–65: holotype) and G. affinis (Fig. 66: holotype; Figs. 67–68: India, Sonamarg; Fig. 69: Pakistan, Utrar): 64, 66–67, 69—parameral view, 65, 68—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381650" httpUri="https://zenodo.org/record/7381650/files/figure.png" pageId="29" pageNumber="480">64–65</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77687A70DD90F8F26B435569" pageId="29" pageNumber="480" type="materials_examined">
<paragraph id="8BE9034577687A70DD90F8F26B435569" blockId="29.[151,1437,1820,2027]" pageId="29" pageNumber="480">
<emphasis id="B922DF5777687A70DD90F8F269FA5521" bold="true" box="[151,457,1820,1846]" pageId="29" pageNumber="480">Type material examined.</emphasis>
<typeStatus id="54EDBDE777687A70DCD3F8F26A0C5521" box="[468,575,1820,1846]" pageId="29" pageNumber="480" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D50C8CC77687A70DF57F8F36A565520" box="[592,613,1821,1847]" pageId="29" pageNumber="480" type="male">♂</specimenCount>
(
<figureCitation id="136D1FC077687A70DF7EF8F26AE25521" box="[633,721,1820,1846]" captionStart="FIGURES 57–62" captionStartId="30.[151,264,1878,1902]" captionTargetBox="[166,1428,284,1841]" captionTargetId="figure-70@30.[151,1436,273,1855]" captionTargetPageId="30" captionText="FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381644" httpUri="https://zenodo.org/record/7381644/files/figure.png" pageId="29" pageNumber="480">Fig. 60</figureCitation>
; dissected): ‘Arni Gad, | Mussoorie.’ &lt;yellow, printed&gt;, ‘Dr. Cameron | 13. IV. 22’ &lt;printed&gt;, ‘
<taxonomicName id="4C5678C677687A70DF09F8AE6A59554D" box="[526,618,1856,1882]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="genus">Lesteva</taxonomicName>
| torrentum |
<collectingRegion id="4992CDA777687A70DFF8F8AE6B0F554D" box="[767,828,1856,1882]" country="Myanmar" name="Chin" pageId="29" pageNumber="480">Cam</collectingRegion>
.’ &lt;handwritten&gt;, ‘
<typeStatus id="54EDBDE777687A70D906F8AE6CA9554D" box="[1025,1178,1856,1882]" pageId="29" pageNumber="480" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C5678C677687A70D9AAF8AE6D70554D" authorityName="Redtenbacher" authorityYear="1857" box="[1197,1347,1856,1882]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="genus">
<emphasis id="B922DF5777687A70D9AAF8AE6D70554D" box="[1197,1347,1856,1882]" italics="true" pageId="29" pageNumber="480">Geodromicus</emphasis>
</taxonomicName>
(s.str.) |
<taxonomicName id="4C5678C677687A70DD90F88A68CC5569" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,255,1892,1918]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="malcolmi" status="sp. nov.">
<emphasis id="B922DF5777687A70DD90F88A68CC5569" box="[151,255,1892,1918]" italics="true" pageId="29" pageNumber="480">malcolmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211622C77687A70DC01F88B69085568" box="[262,315,1893,1919]" pageId="29" pageNumber="480" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A.V. des. 2022’ &lt;red, printed&gt; (
<collectionCode id="ED479B8077687A70DE09F88A6B575569" box="[782,868,1892,1918]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="29" pageNumber="480">NMPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77687A4DDDC0F86669C852C2" lastPageId="32" lastPageNumber="483" pageId="29" pageNumber="480" type="description">
<paragraph id="8BE9034577687A70DDC0F86669A255FD" blockId="29.[151,1437,1820,2027]" pageId="29" pageNumber="480">
<emphasis id="B922DF5777687A70DDC0F866696A55B5" bold="true" box="[199,345,1928,1954]" pageId="29" pageNumber="480">Description.</emphasis>
Measurements (
<typeStatus id="54EDBDE777687A70DF16F8666A4955B5" box="[529,634,1928,1954]" pageId="29" pageNumber="480" type="holotype">holotype</typeStatus>
): HW: 0.97; HL: 0.60; OL: 0.25; TL: 0.15; AL: 3.30; PL: 0.85; PWmax: 1.08; PWmin: 0.95; ESL: 1.87; EW: 1.82; MTbL: 1.25; MTrL: 0.57 (MTrL 1–4: 0.32; MTrL 5: 0.25); AW: 1.65; AedL: 0.94; BL: 4.90.
</paragraph>
<paragraph id="8BE90345776B7A73DDC0FF796B7452EE" blockId="30.[151,1436,151,249]" pageId="30" pageNumber="481">
Habitus as in
<figureCitation id="136D1FC0776B7A73DC5DFF79699852A6" box="[346,427,151,177]" captionStart="FIGURES 57–62" captionStartId="30.[151,264,1878,1902]" captionTargetBox="[166,1428,284,1841]" captionTargetId="figure-70@30.[151,1436,273,1855]" captionTargetPageId="30" captionText="FIGURES 57–62. Habitus of Geodromicus: 57–59—G. macrothorax (Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—G. malcolmi (holotype), 61—G. medvedevi (holotype), 62—G. affinis (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7381644" httpUri="https://zenodo.org/record/7381644/files/figure.png" pageId="30" pageNumber="481">Fig. 60</figureCitation>
. Body dark-brown, with slightly paler abdomen; mouthparts, antennae and legs brown (tarsi slightly paler). Forebody without microsculpture except head with traces of shallow transverse microreticulation in frontal portion and moderately coarse meshes between ocelli.
</paragraph>
<caption id="DF2953CD776B7A73DD90F8B869E455A1" ID-DOI="http://doi.org/10.5281/zenodo.7381644" ID-Zenodo-Dep="7381644" httpUri="https://zenodo.org/record/7381644/files/figure.png" pageId="30" pageNumber="481" startId="30.[151,264,1878,1902]" targetBox="[166,1428,284,1841]" targetPageId="30">
<paragraph id="8BE90345776B7A73DD90F8B869E455A1" blockId="30.[151,1437,1878,1974]" pageId="30" pageNumber="481">
<emphasis id="B922DF57776B7A73DD90F8B869665579" bold="true" box="[151,341,1878,1902]" pageId="30" pageNumber="481">FIGURES 57–62.</emphasis>
Habitus of
<taxonomicName id="4C5678C6776B7A73DCCBF8B86A675579" authorityName="Redtenbacher" authorityYear="1857" box="[460,596,1878,1902]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="B922DF57776B7A73DCCBF8B86A675579" box="[460,596,1878,1902]" italics="true" pageId="30" pageNumber="481">Geodromicus</emphasis>
</taxonomicName>
: 57–59—
<taxonomicName id="4C5678C6776B7A73DFBDF8B96B685579" authorityName="Kashcheev" authorityYear="1999" box="[698,859,1878,1902]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="macrothorax">
<emphasis id="B922DF57776B7A73DFBDF8B96AFF5579" box="[698,716,1879,1902]" italics="true" pageId="30" pageNumber="481">G</emphasis>
.
<emphasis id="B922DF57776B7A73DFDEF8B86B685579" box="[729,859,1878,1902]" italics="true" pageId="30" pageNumber="481">macrothorax</emphasis>
</taxonomicName>
(Fig. 57: Kazakhstan, Chelkydysu; Fig. 58: Kazakhstan, Kokjota Mt.; Fig. 59: Kyrgyzstan, Grigoryevka), 60—
<taxonomicName id="4C5678C6776B7A73DFCDF8956B7A5585" authorityName="Shavrin" authorityYear="2022" box="[714,841,1914,1938]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="malcolmi">
<emphasis id="B922DF57776B7A73DFCDF8956B7A5585" box="[714,841,1914,1938]" italics="true" pageId="30" pageNumber="481">G. malcolmi</emphasis>
</taxonomicName>
(holotype), 61—
<taxonomicName id="4C5678C6776B7A73DEFAF8956CBA5585" authorityName="Shavrin" authorityYear="2009" box="[1021,1161,1914,1938]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="medvedevi">
<emphasis id="B922DF57776B7A73DEFAF8956CBA5585" box="[1021,1161,1914,1938]" italics="true" pageId="30" pageNumber="481">G. medvedevi</emphasis>
</taxonomicName>
(holotype), 62—
<taxonomicName id="4C5678C6776B7A73D83BF8956DAF5585" authorityName="Cameron" authorityYear="1934" box="[1340,1436,1914,1938]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B922DF57776B7A73D83BF8956D7D5585" box="[1340,1358,1915,1938]" italics="true" pageId="30" pageNumber="481">G</emphasis>
.
<emphasis id="B922DF57776B7A73D85AF8946DAF5585" box="[1373,1436,1914,1938]" italics="true" pageId="30" pageNumber="481">affinis</emphasis>
</taxonomicName>
(holotype). Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="DF2953CD776A7A72DD90FB406D1F56D1" ID-DOI="http://doi.org/10.5281/zenodo.7381648" ID-Zenodo-Dep="7381648" box="[151,1324,1198,1222]" httpUri="https://zenodo.org/record/7381648/files/figure.png" pageId="31" pageNumber="482" startId="31.[151,250,1198,1222]" targetBox="[151,1437,181,1173]" targetPageId="31">
<paragraph id="8BE90345776A7A72DD90FB406D1F56D1" blockId="31.[151,1324,1198,1222]" box="[151,1324,1198,1222]" pageId="31" pageNumber="482">
<emphasis id="B922DF57776A7A72DD90FB40691356D1" bold="true" box="[151,288,1198,1222]" pageId="31" pageNumber="482">FIGURE 63.</emphasis>
Distribution of
<taxonomicName id="4C5678C6776A7A72DCC5FB406AE156D1" authorityName="Kashcheev" authorityYear="1999" box="[450,722,1198,1222]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="macrothorax">
<emphasis id="B922DF57776A7A72DCC5FB406AE156D1" box="[450,722,1198,1222]" italics="true" pageId="31" pageNumber="482">Geodromicus macrothorax</emphasis>
</taxonomicName>
(squares) and
<taxonomicName id="4C5678C6776A7A72DE60FB406C3A56D1" authorityName="Roubal" authorityYear="1929" box="[871,1033,1198,1222]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="lebedevianus">
<emphasis id="B922DF57776A7A72DE60FB406B4A56D2" box="[871,889,1198,1221]" italics="true" pageId="31" pageNumber="482">G</emphasis>
.
<emphasis id="B922DF57776A7A72DE82FB406C3A56D1" box="[901,1033,1198,1222]" italics="true" pageId="31" pageNumber="482">lebedevianus</emphasis>
</taxonomicName>
(circles) in the Middle Asia.
</paragraph>
</caption>
<paragraph id="8BE90345776A7A72DDC0FB1A6DA957F1" blockId="31.[151,1437,1268,2014]" pageId="31" pageNumber="482">Head 1.6 times as broad as long, with distinctly convex infraorbital portions and supra-antennal elevations, with deep and wide anterio-median depression, distinctly narrowed basad toward wide and deep interocellar depression; temples convex, 1.6 times as long as longitudinal length of eyes; interocellar depression separated from infraorbital portions by deep and long anteocellar foveae, reaching level of apical third of eyes. Ocelli large, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Punctation moderately sparse and fine, finer in middle and slightly larger, deeper and sparser in infraorbital portions. Antennomere 3 distinctly longer than antennomere 2, 4 distinctly shorter than 3, 5–10 slightly longer than 4, apical antennomere 1.3 times as long as 10.</paragraph>
<paragraph id="8BE90345776A7A72DDC0FA1F6D055445" blockId="31.[151,1437,1268,2014]" pageId="31" pageNumber="482">Pronotum moderately small, convex, 1.2 times as broad as long, from widest anterior third gradually narrowed toward moderately long subparallel mediolateral portions; mediobasal portion with deep, transverse depression. Punctation dense, moderately large and deep, sparser in medial portion in front of mediobasal depression.</paragraph>
<paragraph id="8BE90345776A7A72DDC0F9B269B854A9" blockId="31.[151,1437,1268,2014]" pageId="31" pageNumber="482">Elytra slightly longer than broad, slightly broadened posteriad, slightly more than twice as long as pronotum. Punctation dense, slightly larger and deeper than that on pronotum, coarser in parascutellar and finer and sparser in medioapical portions.</paragraph>
<paragraph id="8BE90345776A7A72DDC0F9266B7654F4" blockId="31.[151,1437,1268,2014]" box="[199,837,1736,1763]" pageId="31" pageNumber="482">Metatibia slightly more than twice as long as metatarsus.</paragraph>
<paragraph id="8BE90345776A7A72DDC0F9026A865580" blockId="31.[151,1437,1268,2014]" pageId="31" pageNumber="482">
Male. Apical margin of abdominal tergite VIII and abdominal sternite VIII concave. Aedeagus with very wide basal portion, sharply narrowed toward narrow median lobe, from apical third gradually narrowed toward subacute apex; parameres wide, slightly widened apicad and slightly exceeding apex of median lobe, with four apical moderately long setae; internal sac without visible structures except short flagellum in middle portion (
<figureCitation id="136D1FC0776A7A72D868F8B668845581" captionStart="FIGURES 64–69" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[173,1414,192,1907]" captionTargetId="figure-15@33.[159,1428,181,1919]" captionTargetPageId="33" captionText="FIGURES 64–69. Aedeagus of Geodromicus malcolmi (Figs. 64–65: holotype) and G. affinis (Fig. 66: holotype; Figs. 67–68: India, Sonamarg; Fig. 69: Pakistan, Utrar): 64, 66–67, 69—parameral view, 65, 68—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381650" httpUri="https://zenodo.org/record/7381650/files/figure.png" pageId="31" pageNumber="482">Fig. 64</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="136D1FC0776A7A72DF59F8926A835580" box="[606,688,1916,1943]" captionStart="FIGURES 64–69" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[173,1414,192,1907]" captionTargetId="figure-15@33.[159,1428,181,1919]" captionTargetPageId="33" captionText="FIGURES 64–69. Aedeagus of Geodromicus malcolmi (Figs. 64–65: holotype) and G. affinis (Fig. 66: holotype; Figs. 67–68: India, Sonamarg; Fig. 69: Pakistan, Utrar): 64, 66–67, 69—parameral view, 65, 68—lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7381650" httpUri="https://zenodo.org/record/7381650/files/figure.png" pageId="31" pageNumber="482">Fig. 65</figureCitation>
.
</paragraph>
<paragraph id="8BE90345776A7A72DDC0F84E69A255AD" blockId="31.[151,1437,1268,2014]" box="[199,401,1952,1978]" pageId="31" pageNumber="482">Female unknown.</paragraph>
<paragraph id="8BE90345776A7A4DDDC0F82A69C852C2" blockId="31.[151,1437,1268,2014]" lastBlockId="32.[151,1436,151,429]" lastPageId="32" lastPageNumber="483" pageId="31" pageNumber="482">
<emphasis id="B922DF57776A7A72DDC0F82A698755C9" bold="true" box="[199,436,1988,2014]" pageId="31" pageNumber="482">Comparative notes.</emphasis>
<taxonomicName id="4C5678C6776A7A72DCB8F82A6B2B55C9" authorityName="Shavrin" authorityYear="2022" box="[447,792,1988,2014]" class="Insecta" family="Staphylinidae" genus="Geodromicus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="malcolmi">
<emphasis id="B922DF57776A7A72DCB8F82A6A6655C9" box="[447,597,1988,2014]" italics="true" pageId="31" pageNumber="482">Geodromicus</emphasis>
(s.str.)
<emphasis id="B922DF57776A7A72DFB7F82A6B2B55C9" box="[688,792,1988,2014]" italics="true" pageId="31" pageNumber="482">malcolmi</emphasis>
</taxonomicName>
differs from the remaining species of the
<taxonomicName id="4C5678C6776A7A72D80DF82A6DAF55C9" box="[1290,1436,1988,2014]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="convexicollis">
<emphasis id="B922DF57776A7A72D80DF82A6DAF55C9" box="[1290,1436,1988,2014]" italics="true" pageId="31" pageNumber="482">convexicollis</emphasis>
</taxonomicName>
group by the shape of the elytra slightly longer than broad and twice longer than pronotum, as well as by the different shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77557A4DDDC0FF306B04530A" pageId="32" pageNumber="483" type="distribution">
<paragraph id="8BE9034577557A4DDDC0FF306D1752EE" blockId="32.[151,1436,151,429]" box="[199,1316,222,249]" pageId="32" pageNumber="483">
<emphasis id="B922DF5777557A4DDDC0FF30695252EF" bold="true" box="[199,353,222,248]" pageId="32" pageNumber="483">Distribution.</emphasis>
The new species is known only from the
<typeStatus id="54EDBDE777557A4DDE34FF316B5052EE" box="[819,867,223,249]" pageId="32" pageNumber="483">type</typeStatus>
locality in
<collectingRegion id="4992CDA777557A4DDEE5FF316C5C52EE" box="[994,1135,223,249]" country="India" name="Uttarakhand" pageId="32" pageNumber="483">Uttarakhand</collectingRegion>
,
<collectingCountry id="F34143D577557A4DD97CFF316C8652EE" box="[1147,1205,223,249]" name="India" pageId="32" pageNumber="483">India</collectingCountry>
(
<figureCitation id="136D1FC077557A4DD9C3FF316D2B52EE" box="[1220,1304,223,249]" captionStart="FIGURE 27" captionStartId="13.[151,250,1984,2008]" captionTargetBox="[193,1395,1150,1960]" captionTargetId="figure-31@13.[193,1395,1150,1960]" captionTargetPageId="13" captionText="FIGURE 27. Distribution of Geodromicus championi (circles) and G. malcolmi (square) in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.7381624" httpUri="https://zenodo.org/record/7381624/files/figure.png" pageId="32" pageNumber="483">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BE9034577557A4DDDC0FEEC6B04530A" blockId="32.[151,1436,151,429]" box="[199,823,258,285]" pageId="32" pageNumber="483">
<emphasis id="B922DF5777557A4DDDC0FEEC6978530B" bold="true" box="[199,331,258,284]" pageId="32" pageNumber="483">Bionomics.</emphasis>
The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77557A4DDDC0FEC969C35372" pageId="32" pageNumber="483" type="etymology">
<paragraph id="8BE9034577557A4DDDC0FEC969C35372" blockId="32.[151,1436,151,429]" pageId="32" pageNumber="483">
<emphasis id="B922DF5777557A4DDDC0FEC9697D5356" bold="true" box="[199,334,295,321]" pageId="32" pageNumber="483">Etymology.</emphasis>
The species is named in the memory of famous coleopterologist Malcolm Cameron (1873–1954), collector of the
<typeStatus id="54EDBDE777557A4DDC4FFEA5694B5372" box="[328,376,331,357]" pageId="32" pageNumber="483">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
<subSubSection id="C34C50CE77557A4DDDC0FE816BFC53BA" pageId="32" pageNumber="483" type="discussion">
<paragraph id="8BE9034577557A4DDDC0FE816BFC53BA" blockId="32.[151,1436,151,429]" pageId="32" pageNumber="483">
<emphasis id="B922DF5777557A4DDDC0FE816908539E" bold="true" box="[199,315,367,393]" pageId="32" pageNumber="483">Remarks.</emphasis>
Specimen apparently was collected together with
<taxonomicName id="4C5678C677557A4DDE6DFE816CDB539E" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[874,1256,367,393]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="483" phylum="Arthropoda" rank="species" species="torrentum">
<emphasis id="B922DF5777557A4DDE6DFE816C06539F" box="[874,1077,367,392]" italics="true" pageId="32" pageNumber="483">Lesteva torrentum</emphasis>
<bibRefCitation id="EFC77EB477557A4DD93BFE816CDB539E" author="Cameron, M." box="[1084,1256,367,393]" pageId="32" pageNumber="483" pagination="160 - 198" refId="ref30826" refString="Cameron, M. (1924). New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
from ‘Mussorie District,
<quantity id="4CAEAEA077557A4DDDFEFE7C698D53BA" box="[249,446,402,429]" metricMagnitude="3" metricUnit="m" metricValue="2.0574" metricValueMax="2.1336" metricValueMin="1.9812" pageId="32" pageNumber="483" unit="ft" value="6750.0" valueMax="7000.0" valueMin="6500.0">6500 to 7000 feet</quantity>
above the sea’ (
<bibRefCitation id="EFC77EB477557A4DDF76FE7D6B2F53BA" author="Cameron, M." box="[625,796,403,429]" pageId="32" pageNumber="483" pagination="160 - 198" refId="ref30826" refString="Cameron, M. (1924). New species of Staphylinidae from India. Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron 1924</bibRefCitation>
;
<bibRefCitation id="EFC77EB477557A4DDE2FFE7D6BF053BA" author="Shavrin, A. V." box="[808,963,402,429]" pageId="32" pageNumber="483" pagination="528 - 538" refId="ref31958" refString="Shavrin, A. V. (2015) New data on Himalayan Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae) with description of a new species from Nepal. Zootaxa, 3973 (3), 528 - 538. http: // doi. org / 10.11646 / zootaxa. 3973.3.7" type="journal article" year="2015">Shavrin 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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