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<document id="89A4334DDAB8DFE2AC8FDCD156C90A3A" ID-CLB-Dataset="36895" ID-DOI="10.11646/zootaxa.3768.1.2" ID-GBIF-Dataset="6e8d46da-92c6-4fb4-9009-ef22cfed4d10" ID-ISSN="1175-5326" ID-Zenodo-Dep="285601" ID-ZooBank="D18F9ED3-D883-4413-B509-F1BB9E12FB96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460387894348" checkinUser="plazi" docAuthor="Richards, William J. &amp; Yato, Takuji" docDate="2014" docId="03FF87EBF673000EFF478ADB4179012E" docLanguage="en" docName="zt03768p042.pdf" docOrigin="Zootaxa 3768 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterygotrigla (Parapterygotrigla) multiocellata Matsubara 1937" docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FFC6FF93F6780000FFD08A4C4426032D" masterDocTitle="Revision of the subgenus Parapterygotrigla (Pisces: Triglidae: Pterygotrigla)" masterLastPageNumber="42" masterPageNumber="23" pageNumber="34" updateTime="1698297842744" updateUser="ExternalLinkService">
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<mods:title id="E5E811B7956BD478777F9AFE9009780C">Revision of the subgenus Parapterygotrigla (Pisces: Triglidae: Pterygotrigla)</mods:title>
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<mods:namePart id="24ABA4291E2F93E39CF54D9434F9CFE4">Richards, William J.</mods:namePart>
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<mods:namePart id="C96A5220FDC47D41302F143BC574AA84">Yato, Takuji</mods:namePart>
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<mods:typeOfResource id="4C4905303CA0138B19455D2F8009F46D">text</mods:typeOfResource>
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<mods:title id="AE1C44C25149AA65CEFA1D4079137FFF">Zootaxa</mods:title>
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<mods:date id="614CD0F89BF0D7F01602131CE5F21B07">2014</mods:date>
<mods:detail id="758348D0BBBA2A4825253542B1ADE2CA" type="volume">
<mods:number id="29D0C7450A5967E5834A6928ECDCA484">3768</mods:number>
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<mods:identifier id="A8A0A19705DD265C93622FE733CB3319" type="DOI">10.11646/zootaxa.3768.1.2</mods:identifier>
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<treatment id="03FF87EBF673000EFF478ADB4179012E" ID-DOI="http://doi.org/10.5281/zenodo.5614684" ID-GBIF-Taxon="127667104" ID-Zenodo-Dep="5614684" LSID="urn:lsid:plazi:treatment:03FF87EBF673000EFF478ADB4179012E" httpUri="http://treatment.plazi.org/id/03FF87EBF673000EFF478ADB4179012E" lastPageId="14" lastPageNumber="37" pageId="11" pageNumber="34">
<subSubSection id="C34C6576F673000BFF478ADB45C103D5" pageId="11" pageNumber="34" type="nomenclature">
<paragraph id="8BE936FDF673000BFF478ADB475D039D" blockId="11.[151,891,151,248]" box="[151,891,151,176]" pageId="11" pageNumber="34">
<heading id="D0A18191F673000BFF478ADB475D039D" bold="true" box="[151,891,151,176]" fontSize="10" level="2" pageId="11" pageNumber="34" reason="2">
<taxonomicName id="4C564D7EF673000BFF478ADB475D039D" ID-CoL="78LJS" authority="Matsubara 1937" authorityName="Matsubara" authorityYear="1937" box="[151,891,151,176]" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="11" pageNumber="34" phylum="Chordata" rank="species" species="multiocellata" subGenus="Parapterygotrigla">
<emphasis id="B922EAEFF673000BFF478ADB475D039D" bold="true" box="[151,891,151,176]" pageId="11" pageNumber="34">
<emphasis id="B922EAEFF673000BFF478ADB4508039D" bold="true" box="[151,302,151,176]" italics="true" pageId="11" pageNumber="34">Pterygotrigla</emphasis>
(
<emphasis id="B922EAEFF673000BFEEE8ADB462C039D" bold="true" box="[318,522,151,176]" italics="true" pageId="11" pageNumber="34">Parapterygotrigla</emphasis>
)
<emphasis id="B922EAEFF673000BFDCA8ADB4697039D" bold="true" box="[538,689,151,176]" italics="true" pageId="11" pageNumber="34">multiocellata</emphasis>
<bibRefCitation id="EFC74B0CF673000BFD688ADB475D039D" author="Matsubara" box="[696,891,151,176]" pageId="11" pageNumber="34" refString="Matsubara, K. (1937) Studies on the deep-sea fishes of Japan. V. Diagnosis of a new mail-cheeked fish, Parapterygotrigla multiocellata n. g., n. sp., belonging to Triglidae. Dobutsugaku Zasshi [Zoological Magazine Tokyo], 49 (7), 266 - 267." type="journal article" year="1937">Matsubara 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE936FDF673000BFF478AF0457603F8" blockId="11.[151,891,151,248]" box="[151,336,188,213]" pageId="11" pageNumber="34">Spotted Gurnard</paragraph>
<paragraph id="8BE936FDF673000BFF478A9345C103D5" blockId="11.[151,891,151,248]" box="[151,487,223,248]" pageId="11" pageNumber="34">
(
<figureCitation id="136D2A78F673000BFF4E8A93450203D5" box="[158,292,223,248]" captionStart="FIGURE 5 A" captionStartId="12.[151,250,1917,1939]" captionTargetBox="[239,1346,192,1916]" captionTargetId="figure@12.[239,1347,177,1916]" captionTargetPageId="12" captionText="FIGURE 5 A. Pterygotrigla multiocellata, CSIRO H 1036 - 06, 125.9 mm SL, Australia. B. Dorsal view of heads of Pterygotrigla multiocellata FAKU 4655, 241 mm SL (left) and FAKU 5782, 167 mm SL (right) showing change in shape of AIO and slight beginnings of hyperostosis. C. Lateral view of snout of Pterygotrigla multiocellata FAKU 4655, 241 mm SL showing hyperostosis bulge at spine base.." httpUri="https://zenodo.org/record/285606/files/figure.png" pageId="11" pageNumber="34">Figures 5 A</figureCitation>
–F,
<tableCitation id="C6D40346F673000BFE9E8A93458D03D5" box="[334,427,223,248]" captionStart="TABLE 1" captionStartId="3.[151,239,1033,1056]" captionText="TABLE 1. Frequency distribution of fin rays in the species of the subgenus Parapterygotrigla. Holotype values in boldface." pageId="11" pageNumber="34">Tables 1</tableCitation>
&amp;
<tableCitation id="C6D40346F673000BFE1F8AAC45F803D5" box="[463,478,224,248]" captionStart="TABLE 2" captionStartId="3.[151,239,1540,1563]" captionTargetBox="[159,1422,1162,1475]" captionTargetPageId="3" captionText="TABLE 2. Frequency distribution and length of gill rakers in the species of the subgenus Parapterygotrigla. Holotype values in boldface, r indicates rudimentary gill rakers." httpUri="http://table.plazi.org/id/DF296675F67B0003FF478C4846F10513" pageId="11" pageNumber="34" tableUuid="DF296675F67B0003FF478C4846F10513">2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C6576F673000BFF478B6445400139" pageId="11" pageNumber="34" type="reference_group">
<paragraph id="8BE936FDF673000BFF478B6447710295" blockId="11.[151,1436,296,533]" pageId="11" pageNumber="34">
<treatmentCitationGroup id="AB4611D3F673000BFF478B6447710295" pageId="11" pageNumber="34">
<taxonomicName id="4C564D7EF673000BFF478B6446F00213" ID-CoL="6TQMD" authority="Matsubara 1937:266" authorityName="Matsubara" authorityPageNumber="266" authorityYear="1937" box="[151,726,296,318]" family="Triglidae" genus="Parapterygotrigla" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpaeniformes" pageId="11" pageNumber="34" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF673000BFF478B6445C00213" box="[151,486,296,318]" italics="true" pageId="11" pageNumber="34">Parapterygotrigla multiocellata</emphasis>
<treatmentCitation id="0AF710ECF673000BFE298B6446F00213" author="Matsubara" box="[505,726,296,318]" page="266" pageId="11" pageNumber="34" year="1937">
<bibRefCitation id="EFC74B0CF673000BFE298B64468C0213" author="Matsubara" box="[505,682,296,318]" pageId="11" pageNumber="34" refString="Matsubara, K. (1937) Studies on the deep-sea fishes of Japan. V. Diagnosis of a new mail-cheeked fish, Parapterygotrigla multiocellata n. g., n. sp., belonging to Triglidae. Dobutsugaku Zasshi [Zoological Magazine Tokyo], 49 (7), 266 - 267." type="journal article" year="1937">Matsubara 1937</bibRefCitation>
:266
</treatmentCitation>
</taxonomicName>
(original description,
<collectingCountry id="F341766DF673000BFC0A8B65403E0213" box="[986,1048,297,318]" name="Japan" pageId="11" pageNumber="34">Japan</collectingCountry>
);
<treatmentCitation id="0AF710ECF673000BFBE58B6541350213" author="Kuronuma" box="[1077,1299,296,318]" page="254" pageId="11" pageNumber="34" year="1939">
<bibRefCitation id="EFC74B0CF673000BFBE58B6540C10213" author="Kuronuma" box="[1077,1255,296,318]" pageId="11" pageNumber="34" refString="Kuronuma, K. (1939) A study of the Triglidae of Japan. Bulletin of the Biogeographical Society of Japan, 9, 223 - 260." type="journal article" year="1939">Kuronuma 1939</bibRefCitation>
:254
</treatmentCitation>
(synonymy, description, Kumano-Nada, SE of Kii Peninsula,
<collectingCountry id="F341766DF673000BFD698B0B46D30271" box="[697,757,327,348]" name="Japan" pageId="11" pageNumber="34">Japan</collectingCountry>
, 150 fm. Yatou
<emphasis id="B922EAEFF673000BFC448B04478E0270" box="[916,936,328,349]" italics="true" pageId="11" pageNumber="34">in</emphasis>
Okamura
<emphasis id="B922EAEFF673000BFBC38B0440630271" box="[1043,1093,326,349]" italics="true" pageId="11" pageNumber="34">et al.</emphasis>
1982:281, 398;
<treatmentCitation id="0AF710ECF673000BFB3A8B0A41510271" author="Shen" box="[1258,1399,326,348]" page="33" pageId="11" pageNumber="34" year="1984">
<bibRefCitation id="EFC74B0CF673000BFB3A8B0A417F0271" author="Shen" box="[1258,1369,326,348]" pageId="11" pageNumber="34" refString="Shen, S. C. (1984) Coastal Fishes of Taiwan. National Taiwan University, Taipei, Taiwan, 190 pp." type="book" year="1984">Shen 1984</bibRefCitation>
:33
</treatmentCitation>
, pl. 33 (color photo, description); Ochiai &amp; Yatou in
<treatmentCitation id="0AF710ECF673000BFD178B2947990256" author="Masuda" box="[711,959,357,379]" page="334" pageId="11" pageNumber="34" year="1984">
<bibRefCitation id="EFC74B0CF673000BFD178B2947B50256" author="Masuda" box="[711,915,357,379]" pageId="11" pageNumber="34" refString="Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T. (1984) The fishes of the Japanese Archipelago. Tokai University Press Fishes of the Japanese Archipelago bb Text: i - xxii + 1 - 437, Atlas: Pls. 1 - 370." type="book chapter" year="1984">
Masuda
<emphasis id="B922EAEFF673000BFCF08B2A47700256" box="[800,854,357,379]" italics="true" pageId="11" pageNumber="34">et al.</emphasis>
1984
</bibRefCitation>
:334
</treatmentCitation>
, pl. 300-I (description, color photo); Yatou in Okamura
<emphasis id="B922EAEFF673000BFEE18BC9454F02B7" box="[305,361,388,410]" italics="true" pageId="11" pageNumber="34">et al.</emphasis>
1985:581, 724;
<treatmentCitation id="0AF710ECF673000BFDCF8BC8477602B7" author="Suzuki" box="[543,848,388,410]" page="3" pageId="11" pageNumber="34" year="1994">
<bibRefCitation id="EFC74B0CF673000BFDCF8BC8476702B7" author="Suzuki" box="[543,833,388,410]" pageId="11" pageNumber="34" refString="Suzuki, T. &amp; Hosokawa. M. (1994) First recorded fish species from the Sea of Japan. I. O. P. Division of News, 5 (4) [Apr.], 2 - 6. [in Japanese; English abstract]" type="journal article" year="1994">Suzuki &amp; Hosokawa 1994</bibRefCitation>
:3
</treatmentCitation>
; del
<treatmentCitation id="0AF710ECF673000BFC5A8BC840B002B7" author="Cerro" box="[906,1174,388,410]" page="114" pageId="11" pageNumber="34" year="1997">
<bibRefCitation id="EFC74B0CF673000BFC5A8BC8404A02B7" author="Cerro" box="[906,1132,388,410]" pageId="11" pageNumber="34" refString="del Cerro, L. &amp; Lloris, D. (1997) Gurnard fishes (Scorpaeniformes, Triglidae) from off New Caledonia, with descriptions of five new species. In: Seret, B. (Ed.), Resultats des Campagnes Musorstom. Vol. 17. Memoires du Musee National d'Histoire Naturel, 174, pp. 91 - 124." type="journal article" year="1997">Cerro &amp; Lloris 1997</bibRefCitation>
:114
</treatmentCitation>
(distribution, meristics);
<treatmentCitation id="0AF710ECF673000BFF178BEE45B00295" author="Richards" box="[199,406,418,440]" page="2378" pageId="11" pageNumber="34" year="1999">
<bibRefCitation id="EFC74B0CF673000BFF178BEE457A0295" author="Richards" box="[199,348,418,440]" pageId="11" pageNumber="34" refString="Richards, W. J. (1999) Triglidae. In: Carpenter, K. E. &amp;. Niem, V. H (Eds.), FAO Species Identification Guide for Fishery Purposes. The Living marine Resources of the Western Tropical Pacific, 4 (2), pp. 2359 - 2382 [Distributed 2000]" type="journal article" year="1999">Richards 1999</bibRefCitation>
:2378
</treatmentCitation>
; Richards in Randall &amp; Lim 2000:607 (list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE936FDF673000BFF478B8D45400139" blockId="11.[151,1436,296,533]" pageId="11" pageNumber="34">
<treatmentCitationGroup id="AB4611D3F673000BFF478B8D45400139" pageId="11" pageNumber="34">
<taxonomicName id="4C564D7EF673000BFF478B8D461102FA" ID-CoL="78LJS" authority="Matsubara" authorityName="Matsubara" box="[151,567,449,471]" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="11" pageNumber="34" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF673000BFF478B8D458902FA" box="[151,431,449,471]" italics="true" pageId="11" pageNumber="34">Pterygotrigla multiocellata</emphasis>
(Matsubara)
</taxonomicName>
:
<treatmentCitation id="0AF710ECF673000BFD948B8D472C02FA" author="Kamohara" box="[580,778,449,471]" page="72" pageId="11" pageNumber="34" year="1952">
<bibRefCitation id="EFC74B0CF673000BFD948B8D46CA02FA" author="Kamohara" box="[580,748,449,471]" pageId="11" pageNumber="34" refString="Kamohara, T. (1952) Revised descriptions of the offshore bottom-fishes of Prov. Tosa, Shikoku, Japan. Reports of the Kochi Univiversity Natural Science, (3), 122 pp." type="book" year="1952">Kamohara 1952</bibRefCitation>
:72
</treatmentCitation>
(description, illustration);
<treatmentCitation id="0AF710ECF673000BFBF38B8D40CF02FA" author="Kamohara" box="[1059,1257,449,471]" page="59" pageId="11" pageNumber="34" year="1958">
<bibRefCitation id="EFC74B0CF673000BFBF38B8D40EC02FA" author="Kamohara" box="[1059,1226,449,471]" pageId="11" pageNumber="34" refString="Kamohara, T. (1958) A catalogue of fishes of Kochi Prefecture (Province Tosa), Japan. Report of the Usa Marine Biological Station, 5 (1), 1 - 76." type="journal article" year="1958">Kamohara 1958</bibRefCitation>
:59
</treatmentCitation>
(list);
<treatmentCitation id="0AF710ECF673000BFAE28B8D453F02DB" author="Kamohara" page="78" pageId="11" pageNumber="34" year="1964">
<bibRefCitation id="EFC74B0CF673000BFAE28B8D44DA02DB" author="Kamohara" pageId="11" pageNumber="34" refString="Kamohara, T. (1964) Revised catalogue of fishes of Kochi Prefecture, Japan. Report of the Usa Marine Biological Station, 11 (1), 1 - 99." type="journal article" year="1964">Kamohara 1964</bibRefCitation>
:78
</treatmentCitation>
(list);
<treatmentCitation id="0AF710ECF673000BFE8D8BAC463202DB" author="Nakabo" box="[349,532,480,502]" page="604" pageId="11" pageNumber="34" year="2000">
<bibRefCitation id="EFC74B0CF673000BFE8D8BAC45CF02DB" author="Nakabo" box="[349,489,480,502]" pageId="11" pageNumber="34" refString="Nakabo, T. (2000) (Ed.) Fishes of Japan with pictorial keys to the species. Second edition. Tokai University Press. Fishes of Japan Pictures Keys Species, 1: i - lvi + 1 - 866. [in Japanese]" type="journal article" year="2000">Nakabo 2000</bibRefCitation>
:604
</treatmentCitation>
;
<treatmentCitation id="0AF710ECF673000BFDCF8BAC470102DB" author="Shinohara" box="[543,807,480,502]" page="317" pageId="11" pageNumber="34" year="2001">
<bibRefCitation id="EFC74B0CF673000BFDCF8BAC46DD02DB" author="Shinohara" box="[543,763,480,502]" pageId="11" pageNumber="34" refString="Shinohara, G., Endo, H., Matsuura, K., Machida, Y. &amp; Honda, H. (2001) Annotated Checklist of the Deepwater Fishes from Fishes from Tosa Bay, Japan. National Science Museum Monographs, [20], 283 - 343." type="journal article" year="2001">
Shinohara
<emphasis id="B922EAEFF673000BFD5C8BAD469902DB" box="[652,703,480,502]" italics="true" pageId="11" pageNumber="34">et al.</emphasis>
2001
</bibRefCitation>
:317
</treatmentCitation>
;
<treatmentCitation id="0AF710ECF673000BFCE18BAC47CF02DB" author="Nakabo" box="[817,1001,480,502]" page="604" pageId="11" pageNumber="34" year="2002">
<bibRefCitation id="EFC74B0CF673000BFCE18BAC479802DB" author="Nakabo" box="[817,958,480,502]" pageId="11" pageNumber="34" refString="Nakabo, T. (2002) (Ed.) Fishes of Japan with pictorial keys to the species, English edition. Tokai University Press. Fishes of Japan Pictures Keys Species, 1: i - lxi + 1 - 866. [This edition has some nomenclatural changes from the 2000 Japanese edition.]" type="journal article" year="2002">Nakabo 2002</bibRefCitation>
:604
</treatmentCitation>
;
<treatmentCitation id="0AF710ECF673000BFC238BAD410D02DB" author="Myers" box="[1011,1323,480,502]" page="619" pageId="11" pageNumber="34" year="2003">
<bibRefCitation id="EFC74B0CF673000BFC238BAD40D902DB" author="Myers" box="[1011,1279,480,502]" pageId="11" pageNumber="34" refString="Myers, R. F. &amp; Donaldson, T. J. (2003) The fishes of the Mariana Islands. Micronesica, 35 - 36, 98 - 652." type="book chapter" year="2003">Myers &amp; Donaldson 2003</bibRefCitation>
:619
</treatmentCitation>
;
<treatmentCitation id="0AF710ECF673000BFAE58BAC45440139" author="Shinohara" page="426" pageId="11" pageNumber="34" year="2005">
<bibRefCitation id="EFC74B0CF673000BFAE58BAC45130139" author="Shinohara" pageId="11" pageNumber="34" refString="Shinohara, G., Sato, T., Aonuma, Y., Horikawa, H., Matsuura, K., Nakabo, T. &amp; Sato, K. (2005) Annotated checklist of deepsea fishes from the waters around the Ryukyu Islands, Japan. Deep-sea fauna and pollutants in the Nansei Islands. National Science Museum Monographs, (29), 385 - 452." type="journal article" year="2005">
Shinohara
<emphasis id="B922EAEFF673000BFF17884C44D50139" box="[199,243,510,533]" italics="true" pageId="11" pageNumber="34">et al</emphasis>
. 2005
</bibRefCitation>
:426
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C6576F673000BFF47880D45EE0083" pageId="11" pageNumber="34" type="materials_examined">
<paragraph id="8BE936FDF673000BFF47880D405C0153" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
<emphasis id="B922EAEFF673000BFF47880D45A30177" bold="true" box="[151,389,577,602]" pageId="11" pageNumber="34">Material examined.</emphasis>
<typeStatus id="54ED885FF673000BFE40880D45DE0177" box="[400,504,577,602]" pageId="11" pageNumber="34" type="holotype">Holotype</typeStatus>
(unique)
<collectionCode id="ED47AE38F673000BFDBE880D469E0174" box="[622,696,577,601]" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="11" pageNumber="34">FAKU</collectionCode>
4655, 1 (
<quantity id="4CAE9B18F673000BFCE0880D47B50174" box="[816,915,577,601]" metricMagnitude="-1" metricUnit="m" metricValue="2.41" pageId="11" pageNumber="34" unit="mm" value="241.0">241 mm</quantity>
SL), from
<collectingCountry id="F341766DF673000BFBCD880E40460174" box="[1053,1120,578,601]" name="Japan" pageId="11" pageNumber="34">Japan</collectingCountry>
, Kumano Nada, SE of Kii Peninsula, Pacific coast of Wakayama Prefecture; 1937, at a depth of about 150 fathoms.
</paragraph>
<paragraph id="8BE936FDF673000BFF1788C640E101C2" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
Japanese waters:
<collectionCode id="ED47AE38F673000BFE5C88C645F0018F" box="[396,470,650,674]" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="11" pageNumber="34">FAKU</collectionCode>
5782, 1 (
<quantity id="4CAE9B18F673000BFD9688C54680018C" box="[582,678,649,674]" metricMagnitude="-1" metricUnit="m" metricValue="1.67" pageId="11" pageNumber="34" unit="mm" value="167.0">167 mm</quantity>
SL), from
<collectingCountry id="F341766DF673000BFCF788C6474C018C" box="[807,874,650,673]" name="Japan" pageId="11" pageNumber="34">Japan</collectingCountry>
; Saipan waters: UF118339, 2 (
<quantity id="4CAE9B18F673000BFB1A88C64145018C" box="[1226,1379,650,674]" metricMagnitude="-1" metricUnit="m" metricValue="2.405" metricValueMax="2.41" metricValueMin="2.4" pageId="11" pageNumber="34" unit="mm" value="240.5" valueMax="241.0" valueMin="240.0">240–241 mm</quantity>
SL), FRV Townsend Cromwell Cr. 53, St. 94,
<geoCoordinate id="EE62503AF673000BFDB388FC465901E4" box="[611,639,688,713]" direction="north" orientation="latitude" pageId="11" pageNumber="34" precision="925" value="15.166667">15</geoCoordinate>
o10’N,
<geoCoordinate id="EE62503AF673000BFD1E88FC471F01E5" box="[718,825,684,713]" direction="east" orientation="longitude" pageId="11" pageNumber="34" precision="925" value="145.66667">145o40’E</geoCoordinate>
, near Saipan,
<date id="FFE8103DF673000BFC0688FC404D01E5" box="[982,1131,688,713]" pageId="11" pageNumber="34" value="1974-04-30">30 Apr. 1974</date>
, 366–
<specimenCount id="9D50FD74F673000BFB7F88FC40D901E5" box="[1199,1279,688,713]" pageId="11" pageNumber="34" type="generic">
<quantity id="4CAE9B18F673000BFB7F88FC40DD01E5" box="[1199,1275,688,713]" metricMagnitude="2" metricUnit="m" metricValue="3.79" pageId="11" pageNumber="34" unit="m" value="379.0">379 m</quantity>
.
</specimenCount>
; KU 31095, 1 (
<quantity id="4CAE9B18F673000BFF4F889A44DB01C2" box="[159,253,726,751]" metricMagnitude="-1" metricUnit="m" metricValue="3.26" pageId="11" pageNumber="34" unit="mm" value="326.0">326 mm</quantity>
SL),
<geoCoordinate id="EE62503AF673000BFEEC889A459A01C2" box="[316,444,723,751]" direction="north" orientation="latitude" pageId="11" pageNumber="34" precision="5" value="15.2008">15.2008o N</geoCoordinate>
,
<geoCoordinate id="EE62503AF673000BFE18889A466901C2" box="[456,591,723,751]" direction="east" orientation="longitude" pageId="11" pageNumber="34" precision="5" value="145.6831">145.6831o E</geoCoordinate>
, e side of Saipan, hook &amp; line, T. R.
<collectingCountry id="F341766DF673000BFC3D889A401001C2" box="[1005,1078,726,751]" name="Indonesia" pageId="11" pageNumber="34">Flores</collectingCountry>
,
<quantity id="4CAE9B18F673000BFB91889A40E501C2" box="[1089,1219,726,751]" metricMagnitude="2" metricUnit="m" metricValue="2.645" metricValueMax="2.65" metricValueMin="2.6399999999999997" pageId="11" pageNumber="34" unit="m" value="264.5" valueMax="265.0" valueMin="264.0">
264–
<specimenCount id="9D50FD74F673000BFBA9889A40E501C2" box="[1145,1219,726,751]" pageId="11" pageNumber="34" type="generic">265 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="8BE936FDF673000BFF4788B145EE0083" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
Australian waters:
<collectionCode id="ED47AE38F673000BFEBA88B2459A003B" box="[362,444,766,790]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="11" pageNumber="34">CSIRO</collectionCode>
H1183-06, 1 (
<quantity id="4CAE9B18F673000BFD8E88B2469A0038" box="[606,700,766,790]" metricMagnitude="-1" metricUnit="m" metricValue="2.32" pageId="11" pageNumber="34" unit="mm" value="232.0">232 mm</quantity>
SL), Queensland, Queensland Plateau, E of Flinders Reef,
<geoCoordinate id="EE62503AF673000BFA8388B2448E0010" direction="south" orientation="latitude" pageId="11" pageNumber="34" precision="925" value="-17.533333">17o32’ S</geoCoordinate>
,
<geoCoordinate id="EE62503AF673000BFF66896945070010" box="[182,289,801,829]" direction="east" orientation="longitude" pageId="11" pageNumber="34" precision="925" value="149.56667">149o34’E</geoCoordinate>
,
<quantity id="4CAE9B18F673000BFEFF8968455D0011" box="[303,379,804,829]" metricMagnitude="2" metricUnit="m" metricValue="4.54" pageId="11" pageNumber="34" unit="m" value="454.0">454 m</quantity>
;
<collectionCode id="ED47AE38F673000BFE58896945FD0010" box="[392,475,805,829]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="11" pageNumber="34">CSIRO</collectionCode>
H780-02, 1 (
<quantity id="4CAE9B18F673000BFDA7896946F10011" box="[631,727,805,829]" metricMagnitude="-1" metricUnit="m" metricValue="2.29" pageId="11" pageNumber="34" unit="mm" value="229.0">229 mm</quantity>
SL), Queensland, Marian Plateau,
<geoCoordinate id="EE62503AF673000BFBBA896940E30010" box="[1130,1221,801,829]" direction="south" orientation="latitude" pageId="11" pageNumber="34" precision="925" value="-19.766666">19o46’S</geoCoordinate>
,
<geoCoordinate id="EE62503AF673000BFB03896841180010" box="[1235,1342,801,829]" direction="east" orientation="longitude" pageId="11" pageNumber="34" precision="925" value="152.06667">152o04’E</geoCoordinate>
,
<quantity id="4CAE9B18F673000BFA9B896841B10011" box="[1355,1431,804,829]" metricMagnitude="2" metricUnit="m" metricValue="3.6799999999999997" pageId="11" pageNumber="34" unit="m" value="368.0">368 m</quantity>
;
<collectionCode id="ED47AE38F673000BFF47890044CC0049" box="[151,234,844,868]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="11" pageNumber="34">CSIRO</collectionCode>
H1036-06, 2 (
<quantity id="4CAE9B18F673000BFE408907460E004E" box="[400,552,843,868]" metricMagnitude="-1" metricUnit="m" metricValue="1.24" metricValueMax="1.26" metricValueMin="1.22" pageId="11" pageNumber="34" unit="mm" value="124.0" valueMax="126.0" valueMin="122.0">122–126 mm</quantity>
SL), Western
<collectingCountry id="F341766DF673000BFD1C890747100049" box="[716,822,843,868]" name="Australia" pageId="11" pageNumber="34">Australia</collectingCountry>
, north of Cape Lambert, 19o06.’S,
<geoCoordinate id="EE62503AF673000BFB138900410B0049" box="[1219,1325,839,868]" direction="east" orientation="longitude" pageId="11" pageNumber="34" precision="925" value="117.11667">117o07’E</geoCoordinate>
,
<quantity id="4CAE9B18F673000BFAEA8900449600AA" metricMagnitude="2" metricUnit="m" metricValue="1.805" metricValueMax="1.83" metricValueMin="1.78" pageId="11" pageNumber="34" unit="m" value="180.5" valueMax="183.0" valueMin="178.0">178–183 m</quantity>
; AMS I.22807-029, 6 (
<quantity id="4CAE9B18F673000BFE198923464200AA" box="[457,612,879,904]" metricMagnitude="-1" metricUnit="m" metricValue="1.755" metricValueMax="1.93" metricValueMin="1.58" pageId="11" pageNumber="34" unit="mm" value="175.5" valueMax="193.0" valueMin="158.0">158–193 mm</quantity>
SL),Western
<collectingCountry id="F341766DF673000BFCD88923475400A5" box="[776,882,879,904]" name="Australia" pageId="11" pageNumber="34">Australia</collectingCountry>
, Northwest Shelf,
<quantity id="4CAE9B18F673000BFB818923408D00A5" box="[1105,1195,879,904]" metricMagnitude="5" metricUnit="m" metricValue="1.75" pageId="11" pageNumber="34" unit="km" value="175.0">175 km</quantity>
N of Port Hedland,
<geoCoordinate id="EE62503AF673000BFF4789DB44D70082" box="[151,241,914,943]" direction="south" orientation="latitude" pageId="11" pageNumber="34" precision="925" value="-18.533333">18o32’S</geoCoordinate>
,
<geoCoordinate id="EE62503AF673000BFF2D89DB45400082" box="[253,358,914,943]" direction="east" orientation="longitude" pageId="11" pageNumber="34" precision="925" value="118.28333">118o17’E</geoCoordinate>
,
<quantity id="4CAE9B18F673000BFEA389DB45990083" box="[371,447,919,943]" metricMagnitude="2" metricUnit="m" metricValue="2.04" pageId="11" pageNumber="34" unit="m" value="204.0">204 m</quantity>
;.
</paragraph>
</subSubSection>
<subSubSection id="C34C6576F673000BFF1789F64030070E" pageId="11" pageNumber="34" type="diagnosis">
<paragraph id="8BE936FDF673000BFF1789F64030070E" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
<emphasis id="B922EAEFF673000BFF1789F6456700FE" bold="true" box="[199,321,954,979]" pageId="11" pageNumber="34">Diagnosis.</emphasis>
Antrorse spine at base of AIO spine present, AIO moderate to long (6–22% SL). P1 fin long (40– 52% SL), moderate eye (8–12% SL), pterygiophores exposed along base of D1 (large plates including one in advance of 1st D1 spine), cleithral spine long (13–20%SL), GR short (1–2% SL).
</paragraph>
</subSubSection>
<subSubSection id="C34C6576F673000BFF178E7D47DA0525" pageId="11" pageNumber="34" type="description">
<paragraph id="8BE936FDF673000BFF178E7D47FB07F2" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
<emphasis id="B922EAEFF673000BFF178E7D457F0767" bold="true" box="[199,345,1073,1098]" pageId="11" pageNumber="34">Description.</emphasis>
D
<date id="FFE8103DF673000BFEA48E7345CD0764" box="[372,491,1074,1101]" pageId="11" pageNumber="34">1 VII–VIII</date>
, D2 11(10–12), A 12(11–13), P1 long (40–52% SL) 12 (11–13) + 3, C 8 + 7,
<date id="FFE8103DF673000BFA838E7E41A20767" box="[1363,1412,1074,1098]" pageId="11" pageNumber="34" value="2011-05">v 11</date>
+ 16 = 27, 10 bucklers along base of D1, GR short (1.1–2.2 mm) with 8–9 (6–11) on ceratohyal + rudiments, llsc 60– 70, nape and breast lack scales, belly 50% covered with scales, nuchal spine long (12–15%SL), no ocular spines, opercular spine short, preopercular spine short, preopercular ridge present, cleithral spine long (13–20% SL). Pterygiophore interdigitation pattern: //1/2/1/1/1/1/1/1/1/1/1/2/1/1/1/1/1/1/.
</paragraph>
<paragraph id="8BE936FDF673000BFF178EA747DA0525" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
<emphasis id="B922EAEFF673000BFF178EA745690629" bold="true" box="[199,335,1259,1284]" pageId="11" pageNumber="34">Coloration.</emphasis>
Based on color photos (
<figureCitation id="136D2A78F673000BFD8E8EA746860629" box="[606,672,1259,1284]" captionStart="FIGURE 5 A" captionStartId="12.[151,250,1917,1939]" captionTargetBox="[239,1346,192,1916]" captionTargetId="figure@12.[239,1347,177,1916]" captionTargetPageId="12" captionText="FIGURE 5 A. Pterygotrigla multiocellata, CSIRO H 1036 - 06, 125.9 mm SL, Australia. B. Dorsal view of heads of Pterygotrigla multiocellata FAKU 4655, 241 mm SL (left) and FAKU 5782, 167 mm SL (right) showing change in shape of AIO and slight beginnings of hyperostosis. C. Lateral view of snout of Pterygotrigla multiocellata FAKU 4655, 241 mm SL showing hyperostosis bulge at spine base.." httpUri="https://zenodo.org/record/285606/files/figure.png" pageId="11" pageNumber="34">Fig. 5</figureCitation>
E-F) provided by K.A. Moots of a fresh specimen from Saipan (KU 31095): body orange with red blotches on the head around the eye and snout; yellow spots on the top of the head, nape, and on the trunk above the P1 fin base, below both dorsal fins, and on and below the lateral line; pink spines in the D1 fin with membranes distally pink; D2 fin with pink rays and distally pink membranes; C fin with pink rays, white base and medial rays white; A fin white; P1 fin with pink leading and trailing rays, whitish pink medial rays, and greenish membranes with the distal rays and membranes greenish white; and margins of fin membranes light bluish. This same hue occupied about half of the length of the lower four fin membranes with the basal area pinkish; lower flank and belly white and pectoral-fin base with a bluish tinge.
</paragraph>
</subSubSection>
<subSubSection id="C34C6576F673000BFF178C5F46F00562" pageId="11" pageNumber="34" type="distribution">
<paragraph id="8BE936FDF673000BFF178C5F46F00562" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
<emphasis id="B922EAEFF673000BFF178C5F45460501" bold="true" box="[199,352,1555,1580]" pageId="11" pageNumber="34">Distribution.</emphasis>
Western Pacific and southeastern Indian Ocean. Known from off
<collectingCountry id="F341766DF673000BFB958C5840AE0506" box="[1093,1160,1556,1579]" name="Japan" pageId="11" pageNumber="34">Japan</collectingCountry>
, Saipan, and eastern and western tropical
<collectingCountry id="F341766DF673000BFE818C7B459D057D" box="[337,443,1591,1616]" name="Australia" pageId="11" pageNumber="34">Australia</collectingCountry>
, at depths of
<quantity id="4CAE9B18F673000BFD808C7B46F40562" box="[592,722,1591,1615]" metricMagnitude="2" metricUnit="m" metricValue="2.785" metricValueMax="3.79" metricValueMin="1.78" pageId="11" pageNumber="34" unit="m" value="278.5" valueMax="379.0" valueMin="178.0">
178–
<specimenCount id="9D50FD74F673000BFD578C7B46F40562" box="[647,722,1591,1615]" pageId="11" pageNumber="34" type="generic">379 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6576F673000EFF178C174179012E" lastPageId="14" lastPageNumber="37" pageId="11" pageNumber="34" type="discussion">
<paragraph id="8BE936FDF673000BFF178C17413005B5" blockId="11.[151,1436,577,2024]" pageId="11" pageNumber="34">
<emphasis id="B922EAEFF673000BFF178C17456A0559" bold="true" box="[199,332,1627,1652]" pageId="11" pageNumber="34">Discussion.</emphasis>
The unique character of the presence of antrorse spine at the base of the anterior infraorbital spine is only found in this species of triglid. It may be difficult to discern by sight, but is easily felt by touch.
</paragraph>
<paragraph id="8BE936FDF673000EFF178CEF452D0296" blockId="11.[151,1436,577,2024]" lastBlockId="14.[151,1437,151,515]" lastPageId="14" lastPageNumber="37" pageId="11" pageNumber="34">
The two specimens from Saipan have shorter anterior infraorbital (AIO) spines than the Japanese specimens, and show no indication of spots on the trunk below the first dorsal fin that are visible in the illustration of the
<typeStatus id="54ED885FF673000BFF478CA744DB0429" box="[151,253,1771,1796]" pageId="11" pageNumber="34" type="holotype">holotype</typeStatus>
. Similarly, the Australian specimens appear to lack dorsal spots also, but preservation vagaries may account for their absence. A color image by Yatou (in Okamura
<emphasis id="B922EAEFF673000BFC5D8D5C47EC0405" box="[909,970,1807,1832]" italics="true" pageId="11" pageNumber="34">et al.</emphasis>
1985: 580) shows: AIO spines reddish; dorsum of head and trunk reddish; D1 fin with reddish spines and pale membranes; D2 fin pale with anterior and posterior rays reddish; base of C fin reddish, remainder pale; P1 with white rays and black membranes, slightly reddish at base; P2 fins reddish at base with slightly reddish membranes; A fin pale; belly, gills and lower jaw white. A color image on Plate 300-I by Ochiai &amp; Yatou (in
<bibRefCitation id="EFC74B0CF673000BFCFB8DEB402404ED" author="Masuda" box="[811,1026,1959,1984]" pageId="11" pageNumber="34" refString="Masuda, H., Amaoka, K., Araga, C., Uyeno, T. &amp; Yoshino, T. (1984) The fishes of the Japanese Archipelago. Tokai University Press Fishes of the Japanese Archipelago bb Text: i - xxii + 1 - 437, Atlas: Pls. 1 - 370." type="book chapter" year="1984">
Masuda
<emphasis id="B922EAEFF673000BFC5A8DE547E404ED" box="[906,962,1959,1984]" italics="true" pageId="11" pageNumber="34">et al.</emphasis>
1984
</bibRefCitation>
) depicts: a pinkish color (rather than reddish) for the AIO spines; pinkish D1 fin; pale D2 fin with light pink rays; red C fin; upper P1 fin red with medial rays white with black membranes, and the lower rays pink; P2 pink; and the A fin and belly white.
<bibRefCitation id="EFC74B0CF676000EFB398ADB44F903FA" author="Shen" pageId="14" pageNumber="37" refString="Shen, S. C. (1984) Coastal Fishes of Taiwan. National Taiwan University, Taipei, Taiwan, 190 pp." type="book" year="1984">Shen (1984: 33, pl. 33)</bibRefCitation>
includes a dorsal view with the P1 fins spread — rays pale with bluish gray membranes; about 3 of the P2 fin rays visible are similar in colour. The dorsum is mottled pinkish red, D1 fin with black membranes distally and pinkish rays, D2 fin pale with pinkish rays, C fin has some pink but is mostly white. There appears to be some black spotting on the dorsum, appearing as scattered spots below the D1 &amp; D2 fins. The tips of the AIO spines are white. Springer (1982) suggested that a number of species are conspecific around the edge of the
<collectingCountry id="F341766DF676000EFB438B164120025E" box="[1171,1286,346,371]" name="Philippines" pageId="14" pageNumber="37">Philippine</collectingCountry>
plate; hence, we provisionally consider these forms to be conspecific, but further collecting and molecular analysis may prove otherwise.
</paragraph>
<caption id="DF296675F674000CFF478D31466C04C2" httpUri="https://zenodo.org/record/285606/files/figure.png" pageId="12" pageNumber="35" targetBox="[239,1346,192,1916]" targetPageId="12">
<paragraph id="8BE936FDF674000CFF478D31466C04C2" blockId="12.[151,1436,1917,2031]" pageId="12" pageNumber="35">
<emphasis id="B922EAEFF674000CFF478D31450804BF" bold="true" box="[151,302,1917,1939]" pageId="12" pageNumber="35">FIGURE 5A.</emphasis>
<taxonomicName id="4C564D7EF674000CFEEC8D31467E04BE" box="[316,600,1917,1939]" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="35" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF674000CFEEC8D31467E04BE" box="[316,600,1917,1939]" italics="true" pageId="12" pageNumber="35">Pterygotrigla multiocellata</emphasis>
</taxonomicName>
, CSIRO H1036-06, 125.9 mm SL, Australia.
<emphasis id="B922EAEFF674000CFBB38D32405C04BE" bold="true" box="[1123,1146,1918,1939]" pageId="12" pageNumber="35">B.</emphasis>
Dorsal view of heads of
<taxonomicName id="4C564D7EF674000CFF478DD0458B049F" box="[151,429,1948,1970]" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="35" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF674000CFF478DD0458B049F" box="[151,429,1948,1970]" italics="true" pageId="12" pageNumber="35">Pterygotrigla multiocellata</emphasis>
</taxonomicName>
FAKU 4655, 241 mm SL (left) and FAKU 5782, 167 mm SL (right) showing change in shape of AIO and slight beginnings of hyperostosis.
<emphasis id="B922EAEFF674000CFD888DF7465604FC" bold="true" box="[600,624,1979,2001]" pageId="12" pageNumber="35">C.</emphasis>
Lateral view of snout of
<taxonomicName id="4C564D7EF674000CFCA98DF740A904FC" box="[889,1167,1979,2001]" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="35" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF674000CFCA98DF740A904FC" box="[889,1167,1979,2001]" italics="true" pageId="12" pageNumber="35">Pterygotrigla multiocellata</emphasis>
</taxonomicName>
FAKU 4655, 241 mm SL showing hyperostosis bulge at spine base..
</paragraph>
</caption>
<caption id="DF296675F675000DFF478DED473004D9" httpUri="https://zenodo.org/record/285607/files/figure.png" pageId="13" pageNumber="36" targetBox="[317,1275,200,1905]" targetPageId="13">
<paragraph id="8BE936FDF675000DFF478DED473004D9" blockId="13.[151,1436,1953,2036]" pageId="13" pageNumber="36">
<emphasis id="B922EAEFF675000DFF478DED4500049A" bold="true" box="[151,294,1953,1975]" pageId="13" pageNumber="36">FIGURE 5D.</emphasis>
Dorsal view of head of
<taxonomicName id="4C564D7EF675000DFDF08DED4713049A" box="[544,821,1953,1975]" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="36" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF675000DFDF08DED4713049A" box="[544,821,1953,1975]" italics="true" pageId="13" pageNumber="36">Pterygotrigla multiocellata</emphasis>
</taxonomicName>
UF 118339, 240 mm SL showing shortened AIO spines.
<emphasis id="B922EAEFF675000DFA548DEE41BD049A" bold="true" box="[1412,1435,1954,1975]" pageId="13" pageNumber="36">E.</emphasis>
Lateral view of
<taxonomicName id="4C564D7EF675000DFEEB8D8C467604FB" box="[315,592,1984,2006]" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="36" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF675000DFEEB8D8C467604FB" box="[315,592,1984,2006]" italics="true" pageId="13" pageNumber="36">Pterygotrigla multiocellata</emphasis>
</taxonomicName>
KU 31095, 326.5 mm SL, photo by K. A. Moots.
<emphasis id="B922EAEFF675000DFB898D8C404D04F8" bold="true" box="[1113,1131,1984,2005]" pageId="13" pageNumber="36">F.</emphasis>
Dorsal view of
<taxonomicName id="4C564D7EF675000DFAC48D8C453B04D9" class="Actinopterygii" family="Triglidae" genus="Pterygotrigla" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="36" phylum="Chordata" rank="species" species="multiocellata">
<emphasis id="B922EAEFF675000DFAC48D8C453B04D9" italics="true" pageId="13" pageNumber="36">Pterygotrigla multiocellata</emphasis>
</taxonomicName>
KU 31095, 326.5 mm SL, photo by K. A. Moots.
</paragraph>
</caption>
<paragraph id="8BE936FDF676000EFF178B894179012E" blockId="14.[151,1437,151,515]" pageId="14" pageNumber="37">
Hyperostosis was evident on the anterior first infraorbital spine (Figs. 13, 14) and in some dorsal fin spines.
<bibRefCitation id="EFC74B0CF676000EFF478BA64595012E" box="[151,435,490,515]" pageId="14" pageNumber="37" refString="Smith-Vaniz, W. F., Kaufman, L. S. &amp; Glowacki, J. (1995) Species-specific patterns of hyperostosis in marine teleost fishes. Marine Biology, 121, 573 - 580. http: // dx. doi. org / 10.1007 / bf 00349291" type="journal article">
Smith-Vaniz
<emphasis id="B922EAEFF676000EFEFA8BA74544012E" box="[298,354,490,515]" italics="true" pageId="14" pageNumber="37">et al.</emphasis>
(1995)
</bibRefCitation>
reviewed this condition in several families and noted that it occurs in some triglids.
</paragraph>
</subSubSection>
</treatment>
</document>
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