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<document id="EF8E2E7BCAA13DF0AFFBF48790763FAB" ID-DOI="10.11646/zootaxa.3895.3.3" ID-GBIF-Dataset="7a17da49-b377-4023-b253-701f3ca9c093" ID-ISSN="1175-5326" ID-Zenodo-Dep="252840" ID-ZooBank="7F3E9CC1-E61F-4FE3-9CD5-B8A6C91E7562" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461000873347" checkinUser="plazi" docAuthor="Kaya, Sarp, Chobanov, Dragan &amp; Çiplak, Battal" docDate="2014" docId="03D82877FFC3FFA1FF20807077D8F041" docLanguage="en" docName="zt03895p400.pdf" docOrigin="Zootaxa 3895 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psorodonotus rize Kaya &amp; Ciplak, sp. n." docType="treatment" docVersion="9" lastPageNumber="397" masterDocId="FFE1500FFFDDFFBFFFB7811F7735F76B" masterDocTitle="Review of Psorodonotus Specularis Group (Orthoptera, Tettigoniidae, Tettigoniinae): two new species from North-east Anatolia" masterLastPageNumber="400" masterPageNumber="367" pageNumber="397" updateTime="1698564590624" updateUser="plazi">
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<mods:title id="A26251FFECFFD11FA2F5EFB4702676CC">Review of Psorodonotus Specularis Group (Orthoptera, Tettigoniidae, Tettigoniinae): two new species from North-east Anatolia</mods:title>
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<mods:namePart id="E07EB6AB7178580BEC9AE7F79D9BCBF1">Kaya, Sarp</mods:namePart>
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<mods:namePart id="32510822E4CD21B4A730908412F2E26F">Chobanov, Dragan</mods:namePart>
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<mods:namePart id="F8A6C33CD850AB48B5C71E46812F9215">Çiplak, Battal</mods:namePart>
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<mods:title id="9EE8C9C2FB12A50E1362095545167BCE">Zootaxa</mods:title>
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<mods:date id="9DB622F520B11DFACBD1331692F52CEE">2014</mods:date>
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<mods:number id="934DC84D871C9011E3B9AEF72E029076">3895</mods:number>
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<mods:identifier id="77ADBF627598348C8829146308FD5F46" type="DOI">10.11646/zootaxa.3895.3.3</mods:identifier>
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<treatment id="03D82877FFC3FFA1FF20807077D8F041" ID-DOI="http://doi.org/10.5281/zenodo.5615884" ID-GBIF-Taxon="119598051" ID-Zenodo-Dep="5615884" LSID="urn:lsid:plazi:treatment:03D82877FFC3FFA1FF20807077D8F041" httpUri="http://treatment.plazi.org/id/03D82877FFC3FFA1FF20807077D8F041" lastPageNumber="397" pageId="30" pageNumber="397">
<subSubSection id="C36BCAEAFFC3FFA1FF20807074EAF6C0" pageId="30" pageNumber="397" type="nomenclature">
<paragraph id="8BCE9961FFC3FFA1FF20807075B5F6E1" blockId="30.[151,991,367,427]" box="[151,640,367,394]" pageId="30" pageNumber="397">
<heading id="D0862E0DFFC3FFA1FF20807075B5F6E1" bold="true" box="[151,640,367,394]" fontSize="11" level="1" pageId="30" pageNumber="397" reason="1">
<emphasis id="B9054573FFC3FFA1FF20807075B5F6E1" bold="true" box="[151,640,367,394]" pageId="30" pageNumber="397">
<taxonomicName id="4C71E2E2FFC3FFA1FF2080707503F6E1" ID-CoL="8PJ84" authority="Kaya &amp; Ciplak" authorityName="Kaya &amp; Ciplak" box="[151,566,367,394]" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="rize" status="sp. nov.">
<emphasis id="B9054573FFC3FFA1FF2080707645F6E2" bold="true" box="[151,368,367,393]" italics="true" pageId="30" pageNumber="397">Psorodonotus rize</emphasis>
Kaya &amp; Çıplak
</taxonomicName>
<taxonomicNameLabel id="A236F808FFC3FFA1FD8A806F75B5F6E1" box="[573,640,368,394]" pageId="30" pageNumber="397" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE9961FFC3FFA1FF20808D74EAF6C0" blockId="30.[151,991,367,427]" box="[151,991,402,427]" pageId="30" pageNumber="397">
(
<figureCitation id="134A85E4FFC3FFA1FF28808D77D0F6C0" box="[159,229,402,427]" captionStart="FIGURE 1" captionStartId="1.[151,250,1980,2002]" captionTargetBox="[160,1427,1207,1953]" captionTargetId="figure@1.[151,1436,1199,1957]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of P. s pe c u l ar i s group and populations studied (open symbols indicate to localities in literature for the respective species)." httpUri="https://zenodo.org/record/252841/files/figure.png" pageId="30" pageNumber="397">Figs 1</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FF44808C7636F6C0" box="[243,259,403,427]" captionStart="FIGURES 6 – 19" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[227,1364,202,1920]" captionTargetId="figure@8.[216,1371,193,1932]" captionTargetPageId="8" captionText="FIGURES 6 – 19. Male pronotum (A dorsal view, B lateral view, scale: 5 mm). 6 — RizeOvit-A, 7 — RizeCimil-A, 8 — RizeElevit, 9 — RizeOvit-B, 10 — RizeCimil-B, 11 — BayburtSoğanlı, 12 — GiresunTamdere, 13 — ArtvinSavsat, 14 — ArdahanHanak, 15 — ArdahanCıldır, 16 — LesserCaucasus, 17 — ArmeniaSyunik, 18 — GeorgiaMestia 19 — GeorgiaKitochi" httpUri="https://zenodo.org/record/252843/files/figure.png" pageId="30" pageNumber="397">8</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FEB8808C7618F6C0" box="[271,301,403,427]" captionStart="FIGURES 20 – 33" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[226,1361,210,1917]" captionTargetId="figure@9.[207,1379,193,1931]" captionTargetPageId="9" captionText="FIGURES 20 – 33. Female pronotum (A dorsal view, B lateral view, scale: 5 mm). 20 — RizeOvit-A, 21 — RizeCimil-A, 22 — RizeElevit, 23 — RizeOvit-B, 24 — RizeCimil-B, 25 — BayburtSoğanlı, 26 — GiresunTamdere, 27 — ArtvinSavsat, 28 — ArdahanHanak, 29 - ArdahanCıldır, 30 — LesserCaucasus, 31 — ArmeniaSyunik, 32 — GeorgiaMestia, 33 — GeorgiaKitochi" httpUri="https://zenodo.org/record/252844/files/figure.png" pageId="30" pageNumber="397">22</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FE8E808D766DF6C0" box="[313,344,402,427]" captionStart="FIGURES 34 – 47" captionStartId="10.[151,264,1528,1551]" captionTargetBox="[167,1423,209,1493]" captionTargetId="figure@10.[150,1436,193,1507]" captionTargetPageId="10" captionText="FIGURES 34 – 47. Male tegmina (scale: 5 mm). 34 — RizeOvit-A, 35 — RizeCimil-A, 36 — RizeElevit, 37 — RizeOvit-B, 38 — RizeCimil-B, 39 — BayburtSoğanlı, 40 — GiresunTamdere, 41 — ArtvinSavsat, 42 — ArdahanHanak, 43 — ArdahanCıldır 44 — LesserCaucasus, 45 — ArmeniaSyunik, 46 — GeorgiaMestia 47 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252845/files/figure.png" pageId="30" pageNumber="397">36</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FED4808D76B4F6C0" box="[355,385,402,427]" captionStart="FIGURES 49 – 62" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[166,1423,184,1496]" captionTargetId="figure@12.[150,1436,175,1506]" captionTargetPageId="12" captionText="FIGURES 49 – 62. Male anal tergite (scale: 2.5 mm). 49 — RizeOvit-A, 50 — RizeCimil-A, 51 — RizeElevit, 52 — RizeOvit-B, 53 — RizeCimil-B, 54 — BayburtSoğanlı, 55 — GiresunTamdere, 56 — ArtvinSavsat, 57 — ArdahanHanak, 58 - ArdahanCıldır, 59 — LesserCaucasus, 60 — ArmeniaSyunik, 61 — GeorgiaMestia, 62 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252847/files/figure.png" pageId="30" pageNumber="397">51</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FE3A808D769EF6C0" box="[397,427,402,427]" captionStart="FIGURES 63 – 76" captionStartId="14.[151,264,1952,1975]" captionTargetBox="[279,1302,203,1923]" captionTargetId="figure@14.[268,1319,193,1932]" captionTargetPageId="14" captionText="FIGURES 63 – 76. Male cerci (scale: 2.5 mm). 63 — RizeOvit-A, 64 — RizeCimil-A, 65 — RizeElevit, 66 — RizeOvit-B, 67 — RizeCimil-B, 68 — BayburtSoğanlı, 69 — GiresunTamdere, 70 — ArtvinSavsat, 71 — ArdahanHanak, 72 - ArdahanCıldır, 73 — LesserCaucasus, 74 — ArmeniaSyunik, 75 — GeorgiaMestia, 76 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252848/files/figure.png" pageId="30" pageNumber="397">65</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FE00808C76E0F6C0" box="[439,469,403,427]" captionStart="FIGURES 77 – 90" captionStartId="15.[151,264,1407,1430]" captionTargetBox="[164,1423,206,1377]" captionTargetId="figure@15.[151,1435,193,1386]" captionTargetPageId="15" captionText="FIGURES 77 – 90. Male subgenital plate (scale: 2.5 mm). 77 — RizeOvit-A, 78 — RizeCimil-A, 79 — RizeElevit, 80 — RizeOvit- B, 81 — RizeCimil-B, 82 — BayburtSoğanlı, 83 — GiresunTamdere, 84 — ArtvinSavsat, 85 — ArdahanHanak, 86 — ArdahanCıldır, 87 — LesserCaucasus, 88 — ArmeniaSyunik, 89 — GeorgiaMestia, 90 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252849/files/figure.png" pageId="30" pageNumber="397">79</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FE56808C76CAF6C0" box="[481,511,403,427]" captionStart="FIGURES 91 – 104" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[167,1419,432,1896]" captionTargetId="figure@16.[151,1435,414,1914]" captionTargetPageId="16" captionText="FIGURES 91 – 104. Male titillator (scale: 2.5 mm). 91 — RizeOvit-A, 92 — RizeCimil-A, 93 — RizeElevit, 94 — RizeOvit-B, 95 — RizeCimil-B, 96 — BayburtSoğanlı, 97 — GiresunTamdere, 98 — ArtvinSavsat, 99 — ArdahanHanak, 100 — ArdahanCıldır, 101 — LesserCaucasus, 102 — ArmeniaSyunik, 103 — GeorgiaMestia, 104 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252850/files/figure.png" pageId="30" pageNumber="397">93</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FDBC808C7502F6C0" box="[523,567,403,427]" captionStart="FIGURES 105 – 118" captionStartId="17.[151,264,1906,1929]" captionTargetBox="[188,1386,207,1857]" captionTargetId="figure@17.[175,1399,193,1876]" captionTargetPageId="17" captionText="FIGURES 105 – 118. Ovipositor (scale: 10 mm). 105 — RizeOvit-A, 106 — RizeCimil-A, 107 — RizeElevit, 108 — RizeOvit-B, 109 — RizeCimil-B, 110 — BayburtSoğanlı, 111 — GiresunTamdere, 112 — ArtvinSavsat, 113 — ArdahanHanak, 114 — ArdahanCıldır, 115 — LesserCaucasus, 116 — ArmeniaSyunik, 117 — GeorgiaMestia, 118 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252851/files/figure.png" pageId="30" pageNumber="397">107</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FDF4808C755AF6C0" box="[579,623,403,427]" captionStart="FIGURES 119 – 132" captionStartId="18.[151,264,1912,1935]" captionTargetBox="[272,1325,186,1911]" captionTargetId="figure@18.[261,1325,177,1917]" captionTargetPageId="18" captionText="FIGURES 119 – 132. Female subgenital plate (scale: 2.5 mm). 119 — RizeOvit-A, 120 — RizeCimil-A, 121 — RizeElevit, 122 — RizeOvit-B, 123 — RizeCimil-B, 124 — BayburtSoğanlı, 125 — GiresunTamdere, 126 — ArtvinSavsat, 127 — ArdahanHanak, 128 — ArdahanCıldır, 129 — LesserCaucasus, 130 — ArmeniaSyunik, 131 — GeorgiaMestia, 132 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252852/files/figure.png" pageId="30" pageNumber="397">121</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FDCC808C7590F6C0" box="[635,677,403,427]" captionStart-0="FIGURE 133" captionStart-1="FIGURE 134" captionStart-2="FIGURE 135" captionStart-3="FIGURE 136" captionStart-4="FIGURE 137" captionStartId-0="21.[151,250,842,864]" captionStartId-1="22.[151,250,1955,1977]" captionStartId-2="23.[151,250,837,859]" captionStartId-3="23.[151,250,1549,1571]" captionStartId-4="24.[151,250,1953,1975]" captionTargetBox-0="[156,1430,199,816]" captionTargetBox-1="[207,1355,223,1883]" captionTargetBox-2="[157,1427,197,813]" captionTargetBox-3="[156,1428,913,1525]" captionTargetBox-4="[179,1407,205,1913]" captionTargetId-0="figure@21.[151,1436,195,821]" captionTargetId-1="figure@22.[165,1422,193,1934]" captionTargetId-2="figure@23.[151,1436,193,816]" captionTargetId-3="figure@23.[151,1436,909,1528]" captionTargetId-4="figure@24.[164,1422,193,1932]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="23" captionTargetPageId-4="24" captionText-0="FIGURE 133. Graph obtained from CV analysis of male pronotum." captionText-1="FIGURE 134. Plot for centroid size of pronotum obtained from the same data set (A male and B female)." captionText-2="FIGURE 135. Graph obtained from CV analysis of ovipositor." captionText-3="FIGURE 136. Graph obtained from CV analysis of cerci." captionText-4="FIGURE 137. Plot for ratio of length and width of hind femurs (A male and B female)." httpUri-0="https://zenodo.org/record/252853/files/figure.png" httpUri-1="https://zenodo.org/record/252854/files/figure.png" httpUri-2="https://zenodo.org/record/252855/files/figure.png" httpUri-3="https://zenodo.org/record/252856/files/figure.png" httpUri-4="https://zenodo.org/record/252857/files/figure.png" pageId="30" pageNumber="397">133</figureCitation>
–137,
<figureCitation id="134A85E4FFC3FFA1FD5C808D7422F6C0" box="[747,791,402,427]" captionStart="FIGURES 144 – 146" captionStartId="26.[151,264,1908,1931]" captionTargetBox="[285,1286,201,1874]" captionTargetId="figure@26.[282,1300,193,1891]" captionTargetPageId="26" captionText="FIGURES 144 – 146. Song diagrams per populations (A—a phrase series, B—elements in a phrase) 144 — RizeOvit-A, 145 — RizeCimil-A, 146 — RizeElevit." httpUri="https://zenodo.org/record/252859/files/figure.png" pageId="30" pageNumber="397">146</figureCitation>
,
<figureCitation id="134A85E4FFC3FFA1FC94808D7465F6C0" box="[803,848,402,427]" captionStart="FIGURE 157" captionStartId="31.[151,250,977,999]" captionTargetBox="[182,1398,200,944]" captionTargetId="figure@31.[175,1412,193,955]" captionTargetPageId="31" captionText="FIGURE 157. Plots for temporal song parameter (phrase)" httpUri="https://zenodo.org/record/252863/files/figure.png" pageId="30" pageNumber="397">157</figureCitation>
;
<tableCitation id="C6F3ACDAFFC3FFA1FCEB808D74E2F6C0" box="[860,983,402,427]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="2.[119,200,707,728]" captionStartId-1="3.[151,239,692,715]" captionStartId-2="7.[125,206,749,770]" captionStartId-3="13.[125,206,768,789]" captionTargetBox-0="[119,1881,782,1523]" captionTargetBox-1="[159,1348,892,1005]" captionTargetBox-3="[157,1857,881,1435]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-3="13" captionText-0="TABLE 1. Collection details (see Fig. 1) of specimens used for morphology and song studies (BC, B. Çıplak; SK, S. Kaya; EMK, E. M. Korkmaz; DC, D. Chobanov; M, male; F, female; NA, not included in statistical analyses)." captionText-1="TABLE 2. Number of specimens photographed per population and per structure used in morphometric studies (* not included in statistical analyses)." captionText-2="TABLE 3. (Continued)" captionText-3="TABLE 4. Mahalanοbis distances results amοng pοpulatiοns οf the Specularis Group fοr male cerci and P-values (P &lt;0.0001 fοr all results) frοm permutatiοn test (number οf iteratiοns ∶ 10000)." httpUri-0="http://table.plazi.org/id/DF0EC9E9FFDFFFBDFFC083DC757EF46A" httpUri-1="http://table.plazi.org/id/DF0EC9E9FFDEFFBCFF2083AB76DFF580" httpUri-3="http://table.plazi.org/id/DF0EC9E9FFD0FFB2FFCA821F761BF456" pageId="30" pageNumber="397" tableUuid-0="DF0EC9E9FFDFFFBDFFC083DC757EF46A" tableUuid-1="DF0EC9E9FFDEFFBCFF2083AB76DFF580" tableUuid-3="DF0EC9E9FFD0FFB2FFCA821F761BF456">Tables 1–4</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36BCAEAFFC3FFA1FF2080C2725FF571" pageId="30" pageNumber="397" type="materials_examined">
<paragraph id="8BCE9961FFC3FFA1FF2080C2725FF571" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF2080C2764FF69D" bold="true" box="[151,378,477,502]" pageId="30" pageNumber="397">Material examined</emphasis>
. Holoype, male;
<collectingCountry id="F366D9F1FFC3FFA1FD8A80C17584F69E" box="[573,689,478,501]" name="Turkey" pageId="30" pageNumber="397">TURKEY</collectingCountry>
:
<taxonomicName id="4C71E2E2FFC3FFA1FD0B80C275C7F69D" box="[700,754,477,502]" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="rize" status="sp. nov.">Rize</taxonomicName>
, Çamlıhemşin, Elevit,
<geoCoordinate id="EE45FFA6FFC3FFA1FC4380C173BDF69E" box="[1012,1160,473,502]" direction="north" orientation="latitude" pageId="30" pageNumber="397" precision="1" value="40.857685">40o51.461' N</geoCoordinate>
,
<geoCoordinate id="EE45FFA6FFC3FFA1FB2380C173F0F68C" box="[1172,1221,473,502]" direction="west" orientation="longitude" pageId="30" pageNumber="397" precision="925" value="-0.68333334">
0
<geoCoordinate id="EE45FFA6FFC3FFA1FB1580C173F0F68C" box="[1186,1221,473,502]" direction="east" orientation="longitude" pageId="30" pageNumber="397" precision="1" value="41.00675">41o</geoCoordinate>
</geoCoordinate>
00.405' E,
<quantity id="4C893484FFC3FFA1FA8880C272A2F69E" box="[1343,1431,477,502]" metricMagnitude="3" metricUnit="m" metricValue="1.825" pageId="30" pageNumber="397" unit="m" value="1825.0">1825 m</quantity>
, 0 2.08.2012 (leg. B. Çıplak, S. Kaya &amp; E.M. Korkmaz) (
<collectionCode id="ED6001A4FFC3FFA1FCBD831D7456F571" box="[778,867,514,538]" pageId="30" pageNumber="397">AUZM</collectionCode>
);
<typeStatus id="54CA27C3FFC3FFA1FCC1831D74D3F571" box="[886,998,514,538]" pageId="30" pageNumber="397" type="paratype">Paratypes</typeStatus>
, 19M, 24F, same data as
<typeStatus id="54CA27C3FFC3FFA1FAB7831E7253F571" box="[1280,1382,513,538]" pageId="30" pageNumber="397" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C36BCAEAFFC3FFA1FF70833A73A9F4E9" pageId="30" pageNumber="397" type="diagnosis">
<paragraph id="8BCE9961FFC3FFA1FF70833A73A9F4E9" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF70833A7674F555" bold="true" box="[199,321,549,574]" pageId="30" pageNumber="397">Diagnosis.</emphasis>
<taxonomicName id="4C71E2E2FFC3FFA1FEFE833A7523F556" box="[329,534,549,573]" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="rize" status="sp. nov.">
<emphasis id="B9054573FFC3FFA1FEFE833A7523F556" box="[329,534,549,573]" italics="true" pageId="30" pageNumber="397">Psorodonotus rize</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A236F808FFC3FFA1FDA8833A7555F555" box="[543,608,549,574]" pageId="30" pageNumber="397" rank="species">
<emphasis id="B9054573FFC3FFA1FDA8833A7555F555" bold="true" box="[543,608,549,574]" pageId="30" pageNumber="397">sp. n.</emphasis>
</taxonomicNameLabel>
is belonging to the
<emphasis id="B9054573FFC3FFA1FCF3833A732DF555" bold="true" box="[836,1048,549,574]" pageId="30" pageNumber="397">
<taxonomicName id="4C71E2E2FFC3FFA1FCF3833A74F5F555" box="[836,960,549,574]" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="392" phylum="Arthropoda" rank="species" species="specularis">Specularis</taxonomicName>
Group
</emphasis>
and shows close affinities with
<taxonomicName id="4C71E2E2FFC3FFA1FA30833977EFF50A" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="B9054573FFC3FFA1FA30833977EFF50A" italics="true" pageId="30" pageNumber="397">P. davisi</emphasis>
</taxonomicName>
by sharing several synapomorphies that are unique in the genus. Especially prominent features are the male anal tergite shape with wide shallow medial incision and two triangular processes, the short male cercus with a long tooth, male subgenital plate with long styli and the robust titillators with enlarged basal arms and more spinose apical arms. The new species differs from its sister species (
<taxonomicName id="4C71E2E2FFC3FFA1FC8B83A974AFF5A6" box="[828,922,693,717]" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="B9054573FFC3FFA1FC8B83A974AFF5A6" box="[828,922,693,717]" italics="true" pageId="30" pageNumber="397">P. davisi</emphasis>
</taxonomicName>
) by the slender cercus with a tooth located in the middle (in
<emphasis id="B9054573FFC3FFA1FEF983C5769AF59A" box="[334,431,729,753]" italics="true" pageId="30" pageNumber="397">P. da v i s i</emphasis>
cercus is more robust and with a tooth located in its first half close to the base). Additionally, males of the new species produce two different song
<typeStatus id="54CA27C3FFC3FFA1FC2383E174E6F47D" box="[916,979,766,790]" pageId="30" pageNumber="397">types</typeStatus>
; one of the songs is similar to that of
<taxonomicName id="4C71E2E2FFC3FFA1FA3083E177EFF452" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="B9054573FFC3FFA1FA3083E177EFF452" italics="true" pageId="30" pageNumber="397">P. davisi</emphasis>
</taxonomicName>
by including 4–6 elements in phrase unit while the second includes 7–8 elements in a unit. A phrase with 7–8 elements is unique for the genus. Our unpublished molecular data suggest that there is no gene exchange between these two species and they shared a common ancestor about two million years ago.
</paragraph>
</subSubSection>
<subSubSection id="C36BCAEAFFC3FFA1FF7082927486F4A1" pageId="30" pageNumber="397" type="etymology">
<paragraph id="8BCE9961FFC3FFA1FF7082927486F4A1" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF708292767BF4CD" bold="true" box="[199,334,909,934]" pageId="30" pageNumber="397">Etymology.</emphasis>
Named after its range of occurrence in the
<taxonomicName id="4C71E2E2FFC3FFA1FC9882927456F4CD" box="[815,867,909,934]" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="rize" status="sp. nov.">Rize</taxonomicName>
province, including its
<typeStatus id="54CA27C3FFC3FFA1FBDE829173ACF4CD" box="[1129,1177,910,934]" pageId="30" pageNumber="397">type</typeStatus>
locality, located on the northern slopes of Kaçkar Mountain Range in the north-east of
<collectingCountry id="F366D9F1FFC3FFA1FCE982AE7485F4A1" box="[862,944,945,970]" name="Turkey" pageId="30" pageNumber="397">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36BCAEAFFC3FFA1FF7082CA7259F189" pageId="30" pageNumber="397" type="description">
<paragraph id="8BCE9961FFC3FFA1FF7082CA7319F485" blockId="30.[151,1437,473,1834]" box="[199,1068,981,1006]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF7082CA7520F485" bold="true" box="[199,533,981,1006]" pageId="30" pageNumber="397">
Description Male (
<typeStatus id="54CA27C3FFC3FFA1FE1382CA753BF485" box="[420,526,981,1006]" pageId="30" pageNumber="397" type="holotype">holotype</typeStatus>
)
</emphasis>
. Medium sized for the genus and for the group.
</paragraph>
<paragraph id="8BCE9961FFC3FFA1FF7082E67681F3AD" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF7082E67615F379" bold="true" box="[199,288,1017,1042]" pageId="30" pageNumber="397">Thorax</emphasis>
. Pronotum (
<figureCitation id="134A85E4FFC3FFA1FE0482E676CCF379" box="[435,505,1017,1042]" captionStart="FIGURES 6 – 19" captionStartId="8.[151,264,1953,1976]" captionTargetBox="[227,1364,202,1920]" captionTargetId="figure@8.[216,1371,193,1932]" captionTargetPageId="8" captionText="FIGURES 6 – 19. Male pronotum (A dorsal view, B lateral view, scale: 5 mm). 6 — RizeOvit-A, 7 — RizeCimil-A, 8 — RizeElevit, 9 — RizeOvit-B, 10 — RizeCimil-B, 11 — BayburtSoğanlı, 12 — GiresunTamdere, 13 — ArtvinSavsat, 14 — ArdahanHanak, 15 — ArdahanCıldır, 16 — LesserCaucasus, 17 — ArmeniaSyunik, 18 — GeorgiaMestia 19 — GeorgiaKitochi" httpUri="https://zenodo.org/record/252843/files/figure.png" pageId="30" pageNumber="397">Fig. 8</figureCitation>
) long, at least one and a half longer than the fore tibia, and suddenly widened backward in metazona. Disc of pronotum flattened, distinctly depressed in the middle, with rounded lateral margins and widely rounded hind margin, smooth and shiny in prozona and indistinctly tuberculate in metazona. Paranota smooth and shiny. Tegmina reach to the sixth abdominal tergite and are covered by pronotum up to their one-fourth (
<figureCitation id="134A85E4FFC3FFA1FF28859677C1F3C9" box="[159,244,1161,1186]" captionStart="FIGURES 34 – 47" captionStartId="10.[151,264,1528,1551]" captionTargetBox="[167,1423,209,1493]" captionTargetId="figure@10.[150,1436,193,1507]" captionTargetPageId="10" captionText="FIGURES 34 – 47. Male tegmina (scale: 5 mm). 34 — RizeOvit-A, 35 — RizeCimil-A, 36 — RizeElevit, 37 — RizeOvit-B, 38 — RizeCimil-B, 39 — BayburtSoğanlı, 40 — GiresunTamdere, 41 — ArtvinSavsat, 42 — ArdahanHanak, 43 — ArdahanCıldır 44 — LesserCaucasus, 45 — ArmeniaSyunik, 46 — GeorgiaMestia 47 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252845/files/figure.png" pageId="30" pageNumber="397">Fig. 36</figureCitation>
). The stridulatory peg number varies between 90 and 138 (mean 108).
<collectingCountry id="F366D9F1FFC3FFA1FBA48596737EF3C9" box="[1043,1099,1161,1186]" name="India" pageId="30" pageNumber="397">Hind</collectingCountry>
femora extend beyond the tip of abdomen by one-third.
</paragraph>
<paragraph id="8BCE9961FFC3FFA1FF7085CE722EF2F5" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF7085CE7676F381" bold="true" box="[199,323,1233,1258]" pageId="30" pageNumber="397">Abdomen.</emphasis>
Cercus (
<figureCitation id="134A85E4FFC3FFA1FE1F85CE76CFF381" box="[424,506,1233,1258]" captionStart="FIGURES 63 – 76" captionStartId="14.[151,264,1952,1975]" captionTargetBox="[279,1302,203,1923]" captionTargetId="figure@14.[268,1319,193,1932]" captionTargetPageId="14" captionText="FIGURES 63 – 76. Male cerci (scale: 2.5 mm). 63 — RizeOvit-A, 64 — RizeCimil-A, 65 — RizeElevit, 66 — RizeOvit-B, 67 — RizeCimil-B, 68 — BayburtSoğanlı, 69 — GiresunTamdere, 70 — ArtvinSavsat, 71 — ArdahanHanak, 72 - ArdahanCıldır, 73 — LesserCaucasus, 74 — ArmeniaSyunik, 75 — GeorgiaMestia, 76 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252848/files/figure.png" pageId="30" pageNumber="397">Fig. 65</figureCitation>
) short for both the genus and the species group; with a long and apically incurved tooth located at the beginning of the distal half; almost in the same width basally and distally of the tooth. Anal tergite with a wide and round incision and two projections at the hind margin (
<figureCitation id="134A85E4FFC3FFA1FC4884067367F259" box="[1023,1106,1305,1330]" captionStart="FIGURES 49 – 62" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[166,1423,184,1496]" captionTargetId="figure@12.[150,1436,175,1506]" captionTargetPageId="12" captionText="FIGURES 49 – 62. Male anal tergite (scale: 2.5 mm). 49 — RizeOvit-A, 50 — RizeCimil-A, 51 — RizeElevit, 52 — RizeOvit-B, 53 — RizeCimil-B, 54 — BayburtSoğanlı, 55 — GiresunTamdere, 56 — ArtvinSavsat, 57 — ArdahanHanak, 58 - ArdahanCıldır, 59 — LesserCaucasus, 60 — ArmeniaSyunik, 61 — GeorgiaMestia, 62 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252847/files/figure.png" pageId="30" pageNumber="397">Fig. 51</figureCitation>
). Subgenital plate wide in the base and suddenly tapering distalward in its apical third, with a shallow triangular incision and narrow apical lobes; styli very long, half or more of the medial length of subgenital plate. Titillators (
<figureCitation id="134A85E4FFC3FFA1FB9B847E734AF211" box="[1068,1151,1377,1402]" captionStart="FIGURES 91 – 104" captionStartId="16.[151,264,1935,1958]" captionTargetBox="[167,1419,432,1896]" captionTargetId="figure@16.[151,1435,414,1914]" captionTargetPageId="16" captionText="FIGURES 91 – 104. Male titillator (scale: 2.5 mm). 91 — RizeOvit-A, 92 — RizeCimil-A, 93 — RizeElevit, 94 — RizeOvit-B, 95 — RizeCimil-B, 96 — BayburtSoğanlı, 97 — GiresunTamdere, 98 — ArtvinSavsat, 99 — ArdahanHanak, 100 — ArdahanCıldır, 101 — LesserCaucasus, 102 — ArmeniaSyunik, 103 — GeorgiaMestia, 104 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252850/files/figure.png" pageId="30" pageNumber="397">Fig. 93</figureCitation>
) are strong, with smooth basal arms and wide spinose apical arms; spinules of apical arms ordered in two or more rows dorsally.
</paragraph>
<paragraph id="8BCE9961FFC3FFA1FF7084B675B1F161" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF7084B6766DF2A9" bold="true" box="[199,344,1449,1474]" pageId="30" pageNumber="397">Colouration</emphasis>
. General colouration reddish brown. Vertex of head and disk of pronotum reddish brown or dark olive green. Paranota dark olive green. Tegmina brown with yellow veins.
<collectingCountry id="F366D9F1FFC3FFA1FC4084D2731AF28D" box="[1015,1071,1485,1510]" name="India" pageId="30" pageNumber="397">Hind</collectingCountry>
femur and tibia reddish brown, Abdomen pale reddish brown. Cerci brown.
</paragraph>
<paragraph id="8BCE9961FFC3FFA1FF70870A725BF1D5" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF70870A762BF145" bold="true" box="[199,286,1557,1582]" pageId="30" pageNumber="397">Female</emphasis>
. Pronotum similar to that of male (
<figureCitation id="134A85E4FFC3FFA1FD14870A75C3F145" box="[675,758,1557,1582]" captionStart="FIGURES 20 – 33" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[226,1361,210,1917]" captionTargetId="figure@9.[207,1379,193,1931]" captionTargetPageId="9" captionText="FIGURES 20 – 33. Female pronotum (A dorsal view, B lateral view, scale: 5 mm). 20 — RizeOvit-A, 21 — RizeCimil-A, 22 — RizeElevit, 23 — RizeOvit-B, 24 — RizeCimil-B, 25 — BayburtSoğanlı, 26 — GiresunTamdere, 27 — ArtvinSavsat, 28 — ArdahanHanak, 29 - ArdahanCıldır, 30 — LesserCaucasus, 31 — ArmeniaSyunik, 32 — GeorgiaMestia, 33 — GeorgiaKitochi" httpUri="https://zenodo.org/record/252844/files/figure.png" pageId="30" pageNumber="397">Fig. 22</figureCitation>
), but the rounded lateral margins more gradually widen from mid of prozona toward metazona and the tubercles in metazona are relatively prominent when compared to male. Tegmina fully covered by pronotum, reduced to scale-like appendages hardly overlapping dorsally.
<collectingCountry id="F366D9F1FFC3FFA1FABB87427271F11D" box="[1292,1348,1629,1654]" name="India" pageId="30" pageNumber="397">Hind</collectingCountry>
femora extend beyond end of abdomen as in male. Subgenital (
<figureCitation id="134A85E4FFC3FFA1FCA5879E7446F1F1" box="[786,883,1665,1690]" captionStart="FIGURES 119 – 132" captionStartId="18.[151,264,1912,1935]" captionTargetBox="[272,1325,186,1911]" captionTargetId="figure@18.[261,1325,177,1917]" captionTargetPageId="18" captionText="FIGURES 119 – 132. Female subgenital plate (scale: 2.5 mm). 119 — RizeOvit-A, 120 — RizeCimil-A, 121 — RizeElevit, 122 — RizeOvit-B, 123 — RizeCimil-B, 124 — BayburtSoğanlı, 125 — GiresunTamdere, 126 — ArtvinSavsat, 127 — ArdahanHanak, 128 — ArdahanCıldır, 129 — LesserCaucasus, 130 — ArmeniaSyunik, 131 — GeorgiaMestia, 132 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252852/files/figure.png" pageId="30" pageNumber="397">Fig. 121</figureCitation>
) plate short, transverse, incised medially at hind margin with rounded apical lobes. Ovipositor (
<figureCitation id="134A85E4FFC3FFA1FD1587BA7437F1D5" box="[674,770,1701,1726]" captionStart="FIGURES 105 – 118" captionStartId="17.[151,264,1906,1929]" captionTargetBox="[188,1386,207,1857]" captionTargetId="figure@17.[175,1399,193,1876]" captionTargetPageId="17" captionText="FIGURES 105 – 118. Ovipositor (scale: 10 mm). 105 — RizeOvit-A, 106 — RizeCimil-A, 107 — RizeElevit, 108 — RizeOvit-B, 109 — RizeCimil-B, 110 — BayburtSoğanlı, 111 — GiresunTamdere, 112 — ArtvinSavsat, 113 — ArdahanHanak, 114 — ArdahanCıldır, 115 — LesserCaucasus, 116 — ArmeniaSyunik, 117 — GeorgiaMestia, 118 — GeorgiaKitochi." httpUri="https://zenodo.org/record/252851/files/figure.png" pageId="30" pageNumber="397">Fig. 107</figureCitation>
) slender, long, roughly 3 times of the pronotum length.
</paragraph>
<paragraph id="8BCE9961FFC3FFA1FF7087D67259F189" blockId="30.[151,1437,473,1834]" box="[199,1388,1737,1762]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF7087D6766AF189" bold="true" box="[199,351,1737,1762]" pageId="30" pageNumber="397">Colouration.</emphasis>
Totally olive green, with black spots at the base of abdominal terga, tarsi reddish as in male.
</paragraph>
</subSubSection>
<subSubSection id="C36BCAEAFFC3FFA1FF7087F277D8F041" pageId="30" pageNumber="397" type="distribution">
<paragraph id="8BCE9961FFC3FFA1FF7087F277D8F041" blockId="30.[151,1437,473,1834]" pageId="30" pageNumber="397">
<emphasis id="B9054573FFC3FFA1FF7087F2766CF06D" bold="true" box="[199,345,1773,1798]" pageId="30" pageNumber="397">Distribution</emphasis>
. The new species occurs in the alpine zone of the Northern Kaçkar Range in the
<taxonomicName id="4C71E2E2FFC3FFA1FB5687F27221F06D" box="[1249,1300,1773,1798]" class="Insecta" family="Tettigoniidae" genus="Psorodonotus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="397" phylum="Arthropoda" rank="species" species="rize" status="sp. nov.">Rize</taxonomicName>
Province of
<collectingCountry id="F366D9F1FFC3FFA1FF20860E77DCF041" box="[151,233,1809,1834]" name="Turkey" pageId="30" pageNumber="397">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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