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<document id="0921E53567DA89FA1EBCF09DF4A5BAD6" ID-CLB-Dataset="8324" ID-DOI="10.11646/zootaxa.4920.4.3" ID-GBIF-Dataset="3f6c02fc-66e9-4ad6-a759-ef23da595c48" ID-ISSN="1175-5326" ID-Zenodo-Dep="4491181" ID-ZooBank="22E8AB54-654D-4882-9084-41F0D229D010" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1612249726991" checkinUser="plazi" docAuthor="Ruschel, Tatiana P. &amp; Sanborn, Allen F." docDate="2021" docId="03D787C8EC4EFFC9FFE05881FDE43786" docLanguage="en" docName="zootaxa.4920.4.3.pdf" docOrigin="Zootaxa 4920 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nosola paradoxa Stal 1866" docType="treatment" docVersion="10" lastPageNumber="516" masterDocId="FFEEFFB0EC4AFFCEFF775816FFDA377F" masterDocTitle="Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae)" masterLastPageNumber="527" masterPageNumber="509" pageNumber="513" updateTime="1698926442746" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8F02ED75733F457230F71C55D640B211" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D7EA9A604DAF5A14FD95AF7C2C26D4A2">
<mods:title id="1225937CDF9EB1AF1A9EB6B956B423A4">Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae)</mods:title>
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<mods:namePart id="53CF9B42FC4EDEEC76A0AC3DBA797249">Ruschel, Tatiana P.</mods:namePart>
<mods:nameIdentifier id="30C055605E262330F13803874CCC3F91" type="ORCID">0000-0002-9052-1760</mods:nameIdentifier>
<mods:affiliation id="E181BBD14826989BCEFF50A350C8404B">Plazi, Rio Grande do Sul, Porto Alegre, RS, Brazil. urn: lsid: zoobank. org: author: 9 D 376223 - 5637 - 4376 - 9 CBC- 9 F 98441455 A 6 tatiana. ruschel @ gmail. com; https: // orcid. org / 0000 - 0002 - 9052 - 1760</mods:affiliation>
<mods:nameIdentifier id="9697794799854FC6316B5E3346ED6C3E" type="email">tatiana.ruschel@gmail.com</mods:nameIdentifier>
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<mods:role id="FC4B1DFCD1C1302738B03DFC66229FA7">
<mods:roleTerm id="DC75CD605CC8CCED1F9E1BC615C739D4">Author</mods:roleTerm>
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<mods:namePart id="87A18358A9317BD413A84FC49A087A6D">Sanborn, Allen F.</mods:namePart>
<mods:affiliation id="01E20CEB99748CF521F8C0B8F488614D">Department of Biology, Barry University, 11300 NE Second Avenue, Miami Shores, FL 33161 - 6695, USA. urn: lsid: zoobank. org: author: 9544 FF 3 F- 9 B 1 F- 457 A- 90 A 2 - 8 A 1 C 0 C 1 F 22 A 0</mods:affiliation>
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<mods:title id="8654542B4E3AA50138570A23A26F8001">Zootaxa</mods:title>
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<mods:part id="633A4B923E595626255D2125E81CA1AE">
<mods:date id="459050B3F7695243B8D5BFFD76599D55">2021</mods:date>
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<mods:number id="4C911529E1C2BC3C133B91A331BA5A9A">2021-02-02</mods:number>
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<mods:number id="3D77B123F51ED6E9D1B61CFAB5ED3D65">4920</mods:number>
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<mods:number id="C76C5548637942DFA8B1A700CCE6095D">4</mods:number>
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<mods:classification id="A3B4E446177FA081F6323687449E75C2">journal article</mods:classification>
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<mods:identifier id="D5639086AC08D3E788A430DFBD474761" type="DOI">10.11646/zootaxa.4920.4.3</mods:identifier>
<mods:identifier id="303F86A5AE5D9ECF3AB980EF2FD91452" type="GBIF-Dataset">3f6c02fc-66e9-4ad6-a759-ef23da595c48</mods:identifier>
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<treatment id="03D787C8EC4EFFC9FFE05881FDE43786" ID-DOI="http://doi.org/10.5281/zenodo.4531852" ID-GBIF-Taxon="177317071" ID-Zenodo-Dep="4531852" LSID="urn:lsid:plazi:treatment:03D787C8EC4EFFC9FFE05881FDE43786" httpUri="http://treatment.plazi.org/id/03D787C8EC4EFFC9FFE05881FDE43786" lastPageId="7" lastPageNumber="516" pageId="4" pageNumber="513">
<subSubSection id="C3646555EC4EFFCAFFE05881FE2237CE" box="[151,504,150,178]" pageId="4" pageNumber="513" type="nomenclature">
<paragraph id="8BC136DEEC4EFFCAFFE05881FE2237CE" blockId="4.[151,504,150,213]" box="[151,504,150,178]" pageId="4" pageNumber="513">
<heading id="D08981B2EC4EFFCAFFE05881FE2237CE" bold="true" box="[151,504,150,178]" fontSize="11" level="1" pageId="4" pageNumber="513" reason="1">
<taxonomicName id="4C7E4D5DEC4EFFCAFFE05881FE2237CE" ID-CoL="6SB43" ID-ENA="2218839" authority="Stal, 1866 b" authorityName="Stal" authorityYear="1866" box="[151,504,150,178]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="513" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B90AEACCEC4EFFCAFFE05881FE2237CE" bold="true" box="[151,504,150,178]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFE05881FEBF37CD" bold="true" box="[151,357,151,178]" italics="true" pageId="4" pageNumber="513">Nosola paradoxa</emphasis>
<bibRefCitation id="EFEF4B2FEC4EFFCAFE1A5880FE2237CE" author="Stal" box="[365,504,150,177]" firstAuthor="Stal" pageId="4" pageNumber="513" pagination="151 - 172" refId="ref10910" refString="Stal, C. (1866 b) Analecta Hemipterologica. Berliner Entomologischer Zeitschrift, 10, 151 - 172. https: // doi. org / 10.1002 / mmnd. 18660100112" type="journal article" year="1866" yearSuffix="b">Stål, 1866b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC4EFFCAFFE058ADFEC837AA" box="[151,274,187,213]" pageId="4" pageNumber="513" type="description">
<paragraph id="8BC136DEEC4EFFCAFFE058ADFEC837AA" blockId="4.[151,504,150,213]" box="[151,274,187,213]" pageId="4" pageNumber="513">
(
<figureCitation id="13452A5BEC4EFFCAFFE858ADFED037AA" box="[159,266,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1031,1055]" captionStartId-1="3.[151,250,1778,1802]" captionStartId-2="5.[151,250,1689,1713]" captionStartId-3="6.[151,250,1898,1922]" captionTargetBox-0="[151,1435,345,1007]" captionTargetBox-1="[151,1435,417,1755]" captionTargetBox-2="[151,1436,186,1665]" captionTargetBox-3="[161,1427,309,1875]" captionTargetId-0="figure-108@1.[151,1435,345,1007]" captionTargetId-1="figure-129@3.[151,1435,417,1755]" captionTargetId-2="figure-19@5.[151,1436,181,1665]" captionTargetId-3="figure-97@6.[161,1427,309,1875]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Nosola paradoxa Stål, 1866, holotype male in dorsal view and labels. Scale bar = 50 mm. [Photograph courtesy of Gunvi Lindberg, Swedish Museum of Natural History]." captionText-1="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." captionText-2="FIGURE 3. Nosola paradoxa Stål, 1866, male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in lateroventral view; D, phallus in left view; E, phallus in right view. Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), basal plate (bp), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). The arrow indicates the bud-like lobe developed between the dorsal crest and the lateral branches. Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." captionText-3="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4491183" figureDoi-1="http://doi.org/10.5281/zenodo.4491187" figureDoi-2="http://doi.org/10.5281/zenodo.4491189" figureDoi-3="http://doi.org/10.5281/zenodo.4491192" httpUri-0="https://zenodo.org/record/4491183/files/figure.png" httpUri-1="https://zenodo.org/record/4491187/files/figure.png" httpUri-2="https://zenodo.org/record/4491189/files/figure.png" httpUri-3="https://zenodo.org/record/4491192/files/figure.png" pageId="4" pageNumber="513">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC4EFFCAFFE058ECFDBA366D" box="[151,608,250,274]" pageId="4" pageNumber="513" type="reference_group">
<paragraph id="8BC136DEEC4EFFCAFFE058ECFDBA366D" blockId="4.[151,608,250,274]" box="[151,608,250,274]" pageId="4" pageNumber="513">
<treatmentCitationGroup id="AB6E11F0EC4EFFCAFFE058ECFDBA366D" box="[151,608,250,274]" pageId="4" pageNumber="513">
<treatmentCitation id="0ADF10CFEC4EFFCAFFE058ECFE2E366D" author="Stal, C." box="[151,500,250,274]" page="171" pageId="4" pageNumber="513" year="1866">
<taxonomicName id="4C7E4D5DEC4EFFCAFFE058ECFE2E366D" ID-CoL="6SB43" authority="Stal, 1866 b: 171" authorityName="Stal" authorityPageNumber="171" authorityYear="1866" box="[151,500,250,274]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="513" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B90AEACCEC4EFFCAFFE058ECFE9C366D" box="[151,326,250,274]" italics="true" pageId="4" pageNumber="513">Nosola paradoxa</emphasis>
<bibRefCitation id="EFEF4B2FEC4EFFCAFE3A58ECFE2E366D" author="Stal" box="[333,500,250,274]" firstAuthor="Stal" page="171" pageId="4" pageNumber="513" pagination="151 - 172" refId="ref10910" refString="Stal, C. (1866 b) Analecta Hemipterologica. Berliner Entomologischer Zeitschrift, 10, 151 - 172. https: // doi. org / 10.1002 / mmnd. 18660100112" type="journal article" year="1866" yearSuffix="b">Stål, 1866b: 171</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(sp. nov.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC4EFFCAFFE0592DFE1C36E2" pageId="4" pageNumber="513" type="materials_examined">
<paragraph id="8BC136DEEC4EFFCAFFE0592DFC123606" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFE0592DFE9E362A" bold="true" box="[151,324,315,341]" pageId="4" pageNumber="513">Type material.</emphasis>
<materialsCitation id="3B163C83EC4EFFCAFE3C592DFA47362A" ID-GBIF-Occurrence="4109705301" box="[331,1437,315,342]" collectionCode="NHRS" country="Boliv" location="Boliv" pageId="4" pageNumber="513" specimenCode="000065262" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C5887CEC4EFFCAFE3C592DFE69362A" box="[331,435,315,341]" pageId="4" pageNumber="513" type="holotype">Holotype</typeStatus>
<specimenCount id="9D78FD57EC4EFFCAFECD592DFE16362A" box="[442,460,315,341]" pageId="4" pageNumber="513" type="male">♂</specimenCount>
(
<figureCitation id="13452A5BEC4EFFCAFEA3592DFDC3362A" box="[468,537,315,341]" captionStart="FIGURE 1" captionStartId="1.[151,250,1031,1055]" captionTargetBox="[151,1435,345,1007]" captionTargetId="figure-108@1.[151,1435,345,1007]" captionTargetPageId="1" captionText="FIGURE 1. Nosola paradoxa Stål, 1866, holotype male in dorsal view and labels. Scale bar = 50 mm. [Photograph courtesy of Gunvi Lindberg, Swedish Museum of Natural History]." figureDoi="http://doi.org/10.5281/zenodo.4491183" httpUri="https://zenodo.org/record/4491183/files/figure.png" pageId="4" pageNumber="513">Fig. 1</figureCitation>
): “
<collectingCountry id="F369764EEC4EFFCAFD4D592DFDA0362A" box="[570,634,315,341]" pageId="4" pageNumber="513">Boliv</collectingCountry>
.// Guérin//
<taxonomicName id="4C7E4D5DEC4EFFCAFD8D592DFC4B3629" authority="Stal" authorityName="Stal" authorityYear="1866" box="[762,913,315,342]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="513" phylum="Arthropoda" rank="species" species="paradoxa">paradoxa Stål</taxonomicName>
// typus// NHRS-GULI /
<specimenCode id="DBD89EA5EC4EFFCAFBD0592DFAF23629" box="[1191,1320,315,342]" pageId="4" pageNumber="513">000065262</specimenCode>
” (
<collectionCode id="ED6FAE1BEC4EFFCAFA37592DFA4B362A" box="[1344,1425,315,341]" collectionName="Sweden, Stockholm, Naturhistoriska riksmuseet" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="4" pageNumber="513">NHRS</collectionCode>
).
</materialsCitation>
The specimen is considered a
<typeStatus id="54C5887CEC4EFFCAFE9F5949FD903606" box="[488,586,351,377]" pageId="4" pageNumber="513" type="holotype">holotype</typeStatus>
by monotypy (ICZN, art. 73.1.2).
</paragraph>
<paragraph id="8BC136DEEC4EFFCAFFB05995FE1C36E2" blockId="4.[151,1437,315,2034]" box="[199,454,387,413]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB05995FEBC36E2" bold="true" box="[199,358,387,413]" pageId="4" pageNumber="513">
<typeStatus id="54C5887CEC4EFFCAFFB05995FED836E2" box="[199,258,387,413]" pageId="4" pageNumber="513">Type</typeStatus>
locality.
</emphasis>
Bolivia.
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC4EFFCAFFB059B1FEEB3699" pageId="4" pageNumber="513" type="discussion">
<paragraph id="8BC136DEEC4EFFCAFFB059B1FEEB3699" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB059B1FEE136BE" bold="true" box="[199,315,423,449]" pageId="4" pageNumber="513">Remarks.</emphasis>
The original description is an abbreviated series of remarks on the coloration of the new species (
<bibRefCitation id="EFEF4B2FEC4EFFCAFFE859DDFEFF3699" author="Stal" box="[159,293,459,486]" firstAuthor="Stal" pageId="4" pageNumber="513" pagination="151 - 172" refId="ref10910" refString="Stal, C. (1866 b) Analecta Hemipterologica. Berliner Entomologischer Zeitschrift, 10, 151 - 172. https: // doi. org / 10.1002 / mmnd. 18660100112" type="journal article" year="1866" yearSuffix="b">Stål, 1866b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC4EFFC8FFB059F9FA8E3662" lastPageId="6" lastPageNumber="515" pageId="4" pageNumber="513" type="description">
<paragraph id="8BC136DEEC4EFFCAFFB059F9FDDB3552" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB059F9FE833576" bold="true" box="[199,345,495,521]" pageId="4" pageNumber="513">Description.</emphasis>
Head, pronotum and mesonotum tawny (castaneous in
<typeStatus id="54C5887CEC4EFFCAFCBF59F9FBF43575" box="[968,1070,495,522]" pageId="4" pageNumber="513" type="holotype">holotype</typeStatus>
), abdomen tawny. Body covered by blond setae and small spines.
</paragraph>
<paragraph id="8BC136DEEC4EFFCAFFB05A21FA9B3431" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB05A21FEDE352E" box="[199,260,567,593]" italics="true" pageId="4" pageNumber="513">Head</emphasis>
(
<figureCitation id="13452A5BEC4EFFCAFE625A21FEB9352E" box="[277,355,567,593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="4" pageNumber="513">Figs. 2</figureCitation>
A–C; 4A). Head about as wide as mesonotum, tawny (castaneous in
<typeStatus id="54C5887CEC4EFFCAFB035A21FB00352D" box="[1140,1242,567,594]" pageId="4" pageNumber="513" type="holotype">holotype</typeStatus>
), triangular, with transverse piceous fascia along supra–antennal plates. Eyes castaneous, ocelli rosaceous. Ocular tubercle marked with piceous. Posterior margin of eyes with a black stain and pile setae. Supra–antennal plates prominent relative to anterior margin of head and not meeting eye.Ventral head light brown with transverse piceous fascia between eye and supra–antennal plate. Postclypeus light brown with ten tawny transverse grooves, marked with piceous along medial transverse grooves on either side of midline, blond setae and small spines, central sulcus narrow. Anteclypeus tawny, posterior margin covered with blond setae. Mentum ochraceous, labium ochraceous striped with castaneous with piceous tip reaching middle abdominal sternite I. Proximal scape ochraceous, pedicel castaneous.
</paragraph>
<paragraph id="8BC136DEEC4EFFCAFFB05B41FAAA337E" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB05B41FEC2340E" box="[199,280,855,881]" italics="true" pageId="4" pageNumber="513">Thorax</emphasis>
(
<figureCitation id="13452A5BEC4EFFCAFE5C5B41FE63340D" box="[299,441,855,882]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="4" pageNumber="513">Figs. 2A, D</figureCitation>
;
<figureCitation id="13452A5BEC4EFFCAFEBF5B41FDCB340D" box="[456,529,855,882]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="4" pageNumber="513">4A, B</figureCitation>
). Pronotum tawny (castaneous in
<typeStatus id="54C5887CEC4EFFCAFCD75B41FBDE340D" box="[928,1028,855,882]" pageId="4" pageNumber="513" type="holotype">holotype</typeStatus>
) covered by blond setae and small spines, two piceous longitudinal fasciae along the median and sub-lateral lobes. Lateral fissures marked with piceous. Pronotal collar tawny, lateral margin convex, lateral angle pronounced and projected laterally, wider than mesonotum lateral margin. Mesonotum tawny (castaneous in
<typeStatus id="54C5887CEC4EFFCAFC235BD5FC6034A1" box="[852,954,963,990]" pageId="4" pageNumber="513" type="holotype">holotype</typeStatus>
), wing groove tawny. Cruciform elevation centrally swollen, the surface of posterolateral projections flat, not covering first abdominal tergite completely.
</paragraph>
<paragraph id="8BC136DEEC4EFFCAFFB05C1DFAEB3311" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB05C1DFEE5335A" box="[199,319,1035,1061]" italics="true" pageId="4" pageNumber="513">Operculum</emphasis>
(
<figureCitation id="13452A5BEC4EFFCAFE3A5C1DFE73335A" box="[333,425,1035,1061]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="4" pageNumber="513">Figs. 2E</figureCitation>
;
<figureCitation id="13452A5BEC4EFFCAFEC45C1DFE0C335A" box="[435,470,1035,1061]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="4" pageNumber="513">4C</figureCitation>
). Male operculum ochraceous semilunar with anterior margin straight and base angled posteromedially, posterior margin slightly convex, lateral margin convex, internal angles separated by swelling of abdominal sternite I. The female operculum ochraceous, with medial margin straight and lateral margin short and convex.
</paragraph>
<paragraph id="8BC136DEEC4EFFCAFFB05C6EFCFC32AA" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB05C6EFED133EE" box="[199,267,1144,1169]" italics="true" pageId="4" pageNumber="513">Wings</emphasis>
(
<figureCitation id="13452A5BEC4EFFCAFE6D5C61FEA533EE" box="[282,383,1143,1169]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="4" pageNumber="513">Figs. 2A</figureCitation>
;
<figureCitation id="13452A5BEC4EFFCAFEFD5C61FE6A33EE" box="[394,432,1143,1169]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="4" pageNumber="513">4A</figureCitation>
). Wings hyaline with semi opaque appearance; costal vein strongly arched and widened at base, the expanded proximal costal margin forming an arc bringing the leading edge of the forewing almost to the apex of the postclypeus.Veins tawny marked with castaneous. Forewings with eight apical cells, nine apical cells in one wing of male, infuscation on radial, radiomedial, medial and mediocubital crossveins and on proximal radius anterior 2, radius posterior distal to junction of radiomedial crossvein, proximal median vein 1 and proximal median vein 2, linear infuscation within apical cells terminating in spot on ambient vein, light infuscation delineating nodal line. Basal cell opaque, subcylindrical longer than broad. Hindwings with six apical cells, cubital cell 1 widening distally to base of apical cell 6, venation tawny with castaneous anal veins 2 and 3, anal vein 3 curved at distal terminus. Jugal fold short, not extending to half length of anal lobe. White along proximal cubitus posterior, proximal anal veins 1 and 2 and along anal vein 3, anal cell 3 white.
</paragraph>
<paragraph id="8BC136DEEC4EFFCAFFB05DF6FB6A31D2" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB05DF6FF263286" box="[199,252,1504,1529]" italics="true" pageId="4" pageNumber="513">Legs</emphasis>
(
<figureCitation id="13452A5BEC4EFFCAFE7C5DC9FE583285" box="[267,386,1503,1530]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="4" pageNumber="513">Fig. 2D, E</figureCitation>
;
<figureCitation id="13452A5BEC4EFFCAFEFA5DC9FE683286" box="[397,434,1503,1529]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="4" pageNumber="513">4C</figureCitation>
). Coxae, trochanters, and femora tawny with blond setae, castaneous at apex. Fore femora with castaneous markings and narrow posterolateral region covered by blond setae towards fore tibia. Fore femora with two spines, primary spine in horizontal position in relation to femur, close to femur, secondary spine largest and darker, a right angle to femoral axis. Tarsi castaneous marked with piceous. Pretarsal claws castaneous with piceous tips. Male meracanthus ochraceous, an elongated triangle, short, not reaching middle of operculum. Female meracanthus ochraceous, an elongated triangle, extending over anterior margin of operculum.
</paragraph>
<paragraph id="8BC136DEEC4EFFCAFFB05EA1FB74308E" blockId="4.[151,1437,315,2034]" pageId="4" pageNumber="513">
<emphasis id="B90AEACCEC4EFFCAFFB05EA1FEEB31AE" box="[199,305,1719,1745]" italics="true" pageId="4" pageNumber="513">Abdomen</emphasis>
(
<figureCitation id="13452A5BEC4EFFCAFE355EA1FE3031AE" box="[322,490,1719,1746]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="4" pageNumber="513">Figs. 2A, F, G</figureCitation>
;
<figureCitation id="13452A5BEC4EFFCAFE8F5EA1FDB931AD" box="[504,611,1719,1746]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="4" pageNumber="513">4A, D, E</figureCitation>
). Abdomen tawny, subcylindrical, anterior margin of tergites with blond setae. Timbal cover flat, dorsolateral margin concave exposing dorsal timbal, apex acutely angled almost reaching lateral metascutellar plate, lateral margin meeting the lateral margin of operculum. Male sternites tawny, sternite II with anterior and posterior margins arched anteriorly with blond setae, sternite VII sub-rectangular longest of male sternites, lateral margins concave in anterior sternites becoming convex in posterior sternites, posterior margin slightly emarginated, sternite VIII ovoid with blond setae at apex. Female tergite 9 with the posterior margin slightly convex in lateral view, ventral margins almost straight. Female sternites tawny covered by blond and white setae, sternite VII with the posterior margin sinuous with an obtuse groove in the middle portion, the dorsal beak shorter than the ovipositor sheath in lateral view. Epipleurites tawny, medial region reflexed dorsally.
</paragraph>
<caption id="DF016656EC4FFFCBFFE05E8FFC17303E" ID-DOI="http://doi.org/10.5281/zenodo.4491189" ID-Zenodo-Dep="4491189" httpUri="https://zenodo.org/record/4491189/files/figure.png" pageId="5" pageNumber="514" startId="5.[151,250,1689,1713]" targetBox="[151,1436,186,1665]" targetPageId="5">
<paragraph id="8BC136DEEC4FFFCBFFE05E8FFC17303E" blockId="5.[151,1436,1689,1857]" pageId="5" pageNumber="514">
<emphasis id="B90AEACCEC4FFFCBFFE05E8FFECE31CE" bold="true" box="[151,276,1689,1713]" pageId="5" pageNumber="514">FIGURE 3.</emphasis>
<taxonomicName id="4C7E4D5DEC4FFFCBFE6C5E8FFE6431CE" box="[283,446,1689,1713]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="paradox">
<emphasis id="B90AEACCEC4FFFCBFE6C5E8FFE6431CE" box="[283,446,1689,1713]" italics="true" pageId="5" pageNumber="514">Nosola paradox</emphasis>
</taxonomicName>
a
<bibRefCitation id="EFEF4B2FEC4FFFCBFEA65E8FFDE031CE" author="Stal, C." box="[465,570,1689,1713]" journalOrPublisher="Hemiptera Africana" pageId="5" pageNumber="514" pagination="1 - 276" part="4" refId="ref10871" refString="Stal, C. (1866 a) Hemiptera Homoptera Latr. Hemiptera Africana, 4, 1 - 276, pl. 1. https: // doi. org / 10.5962 / bhl. title. 8566" title="Hemiptera Homoptera Latr" type="journal article" year="1866">Stål, 1866</bibRefCitation>
, male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in lateroventral view; D, phallus in left view; E, phallus in right view. Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), basal plate (bp), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). The arrow indicates the bud-like lobe developed between the dorsal crest and the lateral branches. Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC136DEEC4FFFC8FFB05F79FA8E3662" blockId="5.[151,1437,1903,2038]" lastBlockId="6.[151,1437,151,285]" lastPageId="6" lastPageNumber="515" pageId="5" pageNumber="514">
<emphasis id="B90AEACCEC4FFFCBFFB05F79FEF030F6" box="[199,298,1903,1929]" italics="true" pageId="5" pageNumber="514">Genitalia</emphasis>
(
<figureCitation id="13452A5BEC4FFFCBFE4D5F79FE5930F5" box="[314,387,1903,1930]" captionStart="FIGURE 3" captionStartId="5.[151,250,1689,1713]" captionTargetBox="[151,1436,186,1665]" captionTargetId="figure-19@5.[151,1436,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Nosola paradoxa Stål, 1866, male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in lateroventral view; D, phallus in left view; E, phallus in right view. Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), basal plate (bp), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). The arrow indicates the bud-like lobe developed between the dorsal crest and the lateral branches. Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4491189" httpUri="https://zenodo.org/record/4491189/files/figure.png" pageId="5" pageNumber="514">Figs. 3</figureCitation>
,
<figureCitation id="13452A5BEC4FFFCBFEE75F66FE7A30F5" box="[400,416,1904,1930]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="5" pageNumber="514">4</figureCitation>
). Pygofer subcylindrical; pygofer distal shoulder anterior margin terminating in acute projection sclerotized at apex; pygofer basal lobe short, not reaching mid-length of pygofer, rounded apex, not clearly delimited from lateral pygofer margin. Uncus slightly longer than basal lobe in lateral view; uncal dorsal crest projected posteriorly fused on midline; median uncus lobe undeveloped but present, bud-like; lateral margins of lateral branches of uncus slightly concave; anterior margins of ventral apophyses straight, lateral margins slightly convex. Theca a long tube with a long vesica and extruded sclerotized cornuti in two rows ventrally. Vesica slender slightly inflating toward apex; apex of vesica with central spine and clustered cornuti almost forming a spiral. Female dorsal beak tawny, shorter than the tip of the ovipositor sheath in lateral view. Gonapophyses IX and ovipositor sheath castaneous, blond setae at apex of ovipositor sheath.
</paragraph>
</subSubSection>
<caption id="DF016656EC4CFFC8FFE05F7CFE0C3091" ID-DOI="http://doi.org/10.5281/zenodo.4491192" ID-Zenodo-Dep="4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="6" pageNumber="515" startId="6.[151,250,1898,1922]" targetBox="[161,1427,309,1875]" targetPageId="6">
<paragraph id="8BC136DEEC4CFFC8FFE05F7CFE0C3091" blockId="6.[151,1436,1898,2030]" pageId="6" pageNumber="515">
<emphasis id="B90AEACCEC4CFFC8FFE05F7CFEC930FD" bold="true" box="[151,275,1898,1922]" pageId="6" pageNumber="515">FIGURE 4.</emphasis>
<taxonomicName id="4C7E4D5DEC4CFFC8FE6E5F7CFE6630FD" box="[281,444,1898,1922]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="515" phylum="Arthropoda" rank="species" species="paradox">
<emphasis id="B90AEACCEC4CFFC8FE6E5F7CFE6630FD" box="[281,444,1898,1922]" italics="true" pageId="6" pageNumber="515">Nosola paradox</emphasis>
</taxonomicName>
a
<bibRefCitation id="EFEF4B2FEC4CFFC8FEB95F7CFDE230FD" author="Stal, C." box="[462,568,1898,1922]" journalOrPublisher="Hemiptera Africana" pageId="6" pageNumber="515" pagination="1 - 276" part="4" refId="ref10871" refString="Stal, C. (1866 a) Hemiptera Homoptera Latr. Hemiptera Africana, 4, 1 - 276, pl. 1. https: // doi. org / 10.5962 / bhl. title. 8566" title="Hemiptera Homoptera Latr" type="journal article" year="1866">Stål, 1866</bibRefCitation>
. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm.
</paragraph>
</caption>
<subSubSection id="C3646555EC4DFFC9FFB05880FE7D37AA" pageId="7" pageNumber="516" type="materials_examined">
<paragraph id="8BC136DEEC4DFFC9FFB05880FE7D37AA" blockId="7.[151,1436,150,249]" pageId="7" pageNumber="516">
<emphasis id="B90AEACCEC4DFFC9FFB05880FE7037CF" bold="true" box="[199,426,150,176]" pageId="7" pageNumber="516">Material examined</emphasis>
:
<materialsCitation id="3B163C83EC4DFFC9FECE5881FE7937AA" ID-GBIF-Occurrence="3033803307" collectionCode="MZUSP" collectorName="Dirings" country="Bolivia" location="Santa Cruz" municipality="Santa Cruz" pageId="7" pageNumber="516" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Santa Cruz">
<specimenCount id="9D78FD57EC4DFFC9FECE5881FE3A37CE" box="[441,480,151,177]" pageId="7" pageNumber="516" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D78FD57EC4DFFC9FD635881FDED37CE" box="[532,567,151,177]" pageId="7" pageNumber="516" type="female">2 ♀</specimenCount>
(
<collectionCode id="ED6FAE1BEC4DFFC9FD375881FD7037CE" box="[576,682,151,177]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="516">MZUSP</collectionCode>
). “
<collectingCountry id="F369764EEC4DFFC9FDB05881FCC637CE" box="[711,796,151,177]" name="Bolivia" pageId="7" pageNumber="516">Bolivia</collectingCountry>
/ oriental /
<collectingMunicipality id="6BA5ACA4EC4DFFC9FCE85881FBC137CE" box="[927,1051,151,177]" pageId="7" pageNumber="516">Santa Cruz</collectingMunicipality>
/ (500 Mtr.) /
<collectorName id="268B5308EC4DFFC9FBCA5881FACB37CE" box="[1213,1297,151,177]" pageId="7" pageNumber="516">Dirings</collectorName>
// Coll. Dirings / S. Paulo / Z-168”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC4DFFC9FFB058C8FDE43786" box="[199,574,222,249]" pageId="7" pageNumber="516" type="distribution">
<paragraph id="8BC136DEEC4DFFC9FFB058C8FDE43786" blockId="7.[151,1436,150,249]" box="[199,574,222,249]" pageId="7" pageNumber="516">
<emphasis id="B90AEACCEC4DFFC9FFB058C8FEBB3787" bold="true" box="[199,353,222,248]" pageId="7" pageNumber="516">Distribution.</emphasis>
Bolivia, Argentina.
</paragraph>
</subSubSection>
</treatment>
</document>
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