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<document id="AC6F0744F1C4EED9CC57E977631AC328" ID-DOI="10.11646/zootaxa.4048.2.9" ID-GBIF-Dataset="855de62b-0d8a-467a-9760-ecc9e5df04a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="244848" ID-ZooBank="441B7A90-2573-47DB-AED3-C1EDCEAF387D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461347332970" checkinUser="plazi" docAuthor="Watson, Gillian W., Williams, Douglas J. &amp; Miller, Douglass R." docDate="2015" docId="03D02047FFD7FFB2FF1DC8682094FCC0" docLanguage="en" docName="zt04048p300.pdf" docOrigin="Zootaxa 4048 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fiorinia phantasma Cockerell &amp; Robinson 1915" docType="treatment" docVersion="7" lastPageNumber="298" masterDocId="FFE9583FFFD3FFB5FF8ACB74215CFFBF" masterDocTitle="The identity and distribution of Fiorinia phantasma (Cockerell &amp; Robinson) (Hemiptera: Coccomorpha: Diaspididae), with a new synonym" masterLastPageNumber="300" masterPageNumber="291" pageNumber="295" updateTime="1698620766804" updateUser="plazi">
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<mods:title id="ED618D1C2CECA7E812CA2F29973CFAFA">The identity and distribution of Fiorinia phantasma (Cockerell &amp; Robinson) (Hemiptera: Coccomorpha: Diaspididae), with a new synonym</mods:title>
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<mods:namePart id="306799BD80C894F3143122CD62B87AB3">Watson, Gillian W.</mods:namePart>
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<mods:namePart id="D83CCD907A1CE7EA9109B5CB7641A4E3">Williams, Douglas J.</mods:namePart>
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<mods:namePart id="ED778E6D2041AD9AA089492D1259BAA6">Miller, Douglass R.</mods:namePart>
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<mods:date id="FCE5AB2A659E208F128A61B526F2C44F">2015</mods:date>
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<treatment id="03D02047FFD7FFB2FF1DC8682094FCC0" ID-DOI="http://doi.org/10.5281/zenodo.6116572" ID-GBIF-Taxon="119645402" ID-Zenodo-Dep="6116572" LSID="urn:lsid:plazi:treatment:03D02047FFD7FFB2FF1DC8682094FCC0" httpUri="http://treatment.plazi.org/id/03D02047FFD7FFB2FF1DC8682094FCC0" lastPageId="7" lastPageNumber="298" pageId="4" pageNumber="295">
<subSubSection id="C363C2DAFFD7FFB1FF1DC8682056FCE7" pageId="4" pageNumber="295" type="nomenclature">
<paragraph id="8BC69151FFD7FFB1FF1DC868225EFC88" blockId="4.[151,770,796,856]" box="[151,770,796,823]" pageId="4" pageNumber="295">
<heading id="D08E263DFFD7FFB1FF1DC868225EFC88" bold="true" box="[151,770,796,823]" fontSize="11" level="1" pageId="4" pageNumber="295" reason="1">
<taxonomicName id="4C79EAD2FFD7FFB1FF1DC868225EFC88" ID-CoL="6J5BW" authority="Cockerell &amp; Robinson, 1915" authorityName="Cockerell &amp; Robinson" authorityYear="1915" box="[151,770,796,823]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="phantasma">
<emphasis id="B90D4D43FFD7FFB1FF1DC868225EFC88" bold="true" box="[151,770,796,823]" pageId="4" pageNumber="295">
<emphasis id="B90D4D43FFD7FFB1FF1DC86820D4FC89" bold="true" box="[151,392,796,822]" italics="true" pageId="4" pageNumber="295">Fiorinia phantasma</emphasis>
(
<bibRefCitation id="EFE8ECA0FFD7FFB1FE1DC86923A6FC88" author="Cockerell" box="[407,762,797,823]" pageId="4" pageNumber="295" refString="Cockerell, T. D. A. and Robinson, E. (1915) Descriptions and records of Coccidae. Bulletin of the American Museum of Natural History, 34, 105 - 113." type="journal article" year="1915">Cockerell &amp; Robinson, 1915</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC69151FFD7FFB1FF1DC84B2056FCE7" blockId="4.[151,770,796,856]" box="[151,266,831,856]" pageId="4" pageNumber="295">
(
<figureCitation id="13428DD4FFD7FFB1FF15C84B205DFCE7" box="[159,257,831,856]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[151,250,1892,1914]" captionStartId-1="5.[151,250,1253,1275]" captionStartId-2="6.[151,250,1915,1937]" captionTargetBox-0="[217,1369,1225,1868]" captionTargetBox-1="[414,1140,201,1216]" captionTargetBox-2="[203,1398,203,1866]" captionTargetId-0="figure@4.[217,1369,1225,1868]" captionTargetId-1="figure@5.[404,1161,193,1232]" captionTargetId-2="figure@6.[203,1403,203,1882]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Fiorinia phantasma appearance in life: (A) females and immature male scales (white) on Calophyllum neoebudicum leaf in American Samoa, 21. x. 2008; (B) adult females inside second-instar exuviae with some pigmented banding; (C) adult female inside second-instar exuviae with well-developed pigmented banding (photographs by M. Schmaedick); (D) adult female inside second instar exuviae with no pigmented banding on Arundinaria sp., U. K. (© Fera)." captionText-1="FIGURE 2. The original illustrations of Fiorinia phantasma, reproduced from Cockerell &amp; Robinson (1915): “ 6. Fiorinia phantasma n. sp. End of abdomen of adult female. A. Antennae, adult female. 7. Fiorinia phantasma n. sp. End of abdomen of second stage female. ”" captionText-2="FIGURE 3. Fiorinia phantasma (Cockerell &amp; Robinson); original illustration of the holotype of F. coronata from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process." httpUri-0="https://zenodo.org/record/244849/files/figure.png" httpUri-1="https://zenodo.org/record/244850/files/figure.png" httpUri-2="https://zenodo.org/record/244851/files/figure.png" pageId="4" pageNumber="295">Figs 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C363C2DAFFD7FFB1FF1DC8FD23DDFB2D" pageId="4" pageNumber="295" type="reference_group">
<paragraph id="8BC69151FFD7FFB1FF1DC8FD23A7FC03" blockId="4.[151,1435,904,957]" pageId="4" pageNumber="295">
<treatmentCitationGroup id="AB69B67FFFD7FFB1FF1DC8FD23A7FC03" pageId="4" pageNumber="295">
<taxonomicName id="4C79EAD2FFD7FFB1FF1DC8FD23F4FC21" ID-CoL="6J3FB" authority="Williams &amp; Watson, 1988: 114" authorityName="Williams &amp; Watson" authorityPageNumber="114" authorityYear="1988" box="[151,680,904,926]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="coronata">
<emphasis id="B90D4D43FFD7FFB1FF1DC8FD200DFC21" box="[151,337,905,926]" italics="true" pageId="4" pageNumber="295">Fiorinia coronata</emphasis>
<treatmentCitation id="0AD8B740FFD7FFB1FED1C8FC23F4FC21" author="Williams" box="[347,680,904,926]" page="114" pageId="4" pageNumber="295" year="1988">
<bibRefCitation id="EFE8ECA0FFD7FFB1FED1C8FC232EFC21" author="Williams" box="[347,626,904,926]" pageId="4" pageNumber="295" refString="Williams, D. J. &amp; Watson, G. W. (1988) The scale insects of the tropical South Pacific Region. Pt. 1. The armoured scales (Diaspididae). CAB International, Wallingford, 290 pp." type="book" year="1988">Williams &amp; Watson, 1988</bibRefCitation>
: 114
</treatmentCitation>
</taxonomicName>
, 116.
<typeStatus id="54C22FF3FFD7FFB1FD67C8FC2217FC21" box="[749,843,904,926]" pageId="4" pageNumber="295" type="holotype">Holotype</typeStatus>
adult female,
<collectingCountry id="F36ED1C1FFD7FFB1FC63C8FC25C7FC21" box="[1001,1179,904,926]" name="Solomon Islands" pageId="4" pageNumber="295">Solomon Islands</collectingCountry>
: Guadalcanal, on
<taxonomicName id="4C79EAD2FFD7FFB1FAD6C8FD2047FC03" ID-CoL="WP6H" class="Liliopsida" family="Arecaceae" genus="Cocos" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="295" phylum="Tracheophyta" rank="species" species="nucifera">
<emphasis id="B90D4D43FFD7FFB1FAD6C8FD2047FC03" italics="true" pageId="4" pageNumber="295">Cocos nucifera</emphasis>
</taxonomicName>
, coll. E.S. Brown,?/06/1954 (BMNH).
<emphasis id="B90D4D43FFD7FFB1FD38C8D223A7FC03" bold="true" box="[690,763,934,957]" pageId="4" pageNumber="295">Syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC69151FFD7FFB1FF1DC89D23DDFB2D" blockId="4.[151,1436,1001,1170]" pageId="4" pageNumber="295">
<emphasis id="B90D4D43FFD7FFB1FF1DC89D2025FBBD" bold="true" box="[151,377,1001,1026]" pageId="4" pageNumber="295">Appearance in life</emphasis>
is often quite distinctive, with some or all of the elongate second-instar females / exuviae containing pupillarial adult females having a median series of red-brown spots or transverse red-brown stripes on a yellow background (
<figureCitation id="13428DD4FFD7FFB1FEF4CF45209DFBF5" box="[382,449,1073,1098]" captionStart="FIGURE 1" captionStartId="4.[151,250,1892,1914]" captionTargetBox="[217,1369,1225,1868]" captionTargetId="figure@4.[217,1369,1225,1868]" captionTargetPageId="4" captionText="FIGURE 1. Fiorinia phantasma appearance in life: (A) females and immature male scales (white) on Calophyllum neoebudicum leaf in American Samoa, 21. x. 2008; (B) adult females inside second-instar exuviae with some pigmented banding; (C) adult female inside second-instar exuviae with well-developed pigmented banding (photographs by M. Schmaedick); (D) adult female inside second instar exuviae with no pigmented banding on Arundinaria sp., U. K. (© Fera)." httpUri="https://zenodo.org/record/244849/files/figure.png" pageId="4" pageNumber="295">Fig. 1</figureCitation>
and
<bibRefCitation id="EFE8ECA0FFD7FFB1FE72CF4523D9FBF5" author="Garcia" box="[504,645,1073,1098]" pageId="4" pageNumber="295" refString="Garcia, J. N. (2011) Fiorinia phantasma Cockerell and Robinson (Hemiptera: Diaspididae), an armored scale pest new to Hawaii. Proceedings of the Hawaiian Entomological Society, 43, 59 - 61. Available from: http: // scholarspace. manoa. hawaii. edu / bitstream / handle / 10125 / 21682 / 43 _ 59 - 61. pdf? sequence = 1 (accessed 24 May 2015)" type="journal article" year="2011">Garcia 2011</bibRefCitation>
); however, sometimes there may be no striped individuals present in a colony. HDOA (2011), Anon. (2012) and
<bibRefCitation id="EFE8ECA0FFD7FFB1FDE7CF212252FBD1" author="Bethke" box="[621,782,1109,1134]" pageId="4" pageNumber="295" refString="Bethke, J. A. (2012) Insect hot topics: new scale insect in Hawaii. UCNFA News. University of California, Division of Agriculture and Natural Sciences. Available from: http: // ucanr. edu / sites / UCNFAnews / Insect _ Hot _ Topics / INSECT _ HOT _ TOPICS __ New _ scale _ insect _ in _ Hawaii / (accessed 14 July 2015)" type="journal article" year="2012">Bethke (2012)</bibRefCitation>
mention a colour form that is entirely red, but this has not been present in the samples studied by GW.
</paragraph>
</subSubSection>
<caption id="DF06C1D9FFD7FFB1FF1DCC102424F869" httpUri="https://zenodo.org/record/244849/files/figure.png" pageId="4" pageNumber="295" targetBox="[217,1369,1225,1868]" targetPageId="4">
<paragraph id="8BC69151FFD7FFB1FF1DCC102424F869" blockId="4.[151,1436,1892,2006]" pageId="4" pageNumber="295">
<emphasis id="B90D4D43FFD7FFB1FF1DCC102048F8C6" bold="true" box="[151,276,1892,1914]" pageId="4" pageNumber="295">FIGURE 1</emphasis>
.
<taxonomicName id="4C79EAD2FFD7FFB1FEA2CC1120A6F8C5" box="[296,506,1892,1914]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="phantasma">
<emphasis id="B90D4D43FFD7FFB1FEA2CC1120A6F8C5" box="[296,506,1892,1914]" italics="true" pageId="4" pageNumber="295">Fiorinia phantasma</emphasis>
</taxonomicName>
appearance in life: (A) females and immature male scales (white) on
<taxonomicName id="4C79EAD2FFD7FFB1FA93CC102042F826" class="Magnoliopsida" family="Calophyllaceae" genus="Calophyllum" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="295" phylum="Tracheophyta" rank="species" species="neoebudicum">
<emphasis id="B90D4D43FFD7FFB1FA93CC102042F826" italics="true" pageId="4" pageNumber="295">Calophyllum neoebudicum</emphasis>
</taxonomicName>
leaf in American Samoa, 21.x.2008; (B) adult females inside second-instar exuviae with some pigmented banding; (C) adult female inside second-instar exuviae with well-developed pigmented banding (photographs by M. Schmaedick); (D) adult female inside second instar exuviae with no pigmented banding on
<taxonomicName id="4C79EAD2FFD7FFB1FBBECCB425EDF869" box="[1076,1201,1984,2006]" class="Liliopsida" family="Poaceae" genus="Arundinaria" kingdom="Plantae" order="Poales" pageId="4" pageNumber="295" phylum="Tracheophyta" rank="genus">
<emphasis id="B90D4D43FFD7FFB1FBBECCB425EDF869" box="[1076,1201,1984,2006]" italics="true" pageId="4" pageNumber="295">Arundinaria</emphasis>
</taxonomicName>
sp., U.K. (© Fera).
</paragraph>
</caption>
<caption id="DF06C1D9FFD6FFB0FF1DCF912028FA87" httpUri="https://zenodo.org/record/244850/files/figure.png" pageId="5" pageNumber="296" targetBox="[414,1140,201,1216]" targetPageId="5">
<paragraph id="8BC69151FFD6FFB0FF1DCF912028FA87" blockId="5.[151,1436,1253,1336]" pageId="5" pageNumber="296">
<emphasis id="B90D4D43FFD6FFB0FF1DCF91204CFB45" bold="true" box="[151,272,1253,1275]" pageId="5" pageNumber="296">FIGURE 2</emphasis>
. The original illustrations of
<taxonomicName id="4C79EAD2FFD6FFB0FDCDCF922248FB44" box="[583,788,1253,1275]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="phantasma">
<emphasis id="B90D4D43FFD6FFB0FDCDCF922248FB44" box="[583,788,1253,1275]" italics="true" pageId="5" pageNumber="296">Fiorinia phantasma</emphasis>
</taxonomicName>
, reproduced from Cockerell &amp; Robinson (1915): “6.
<taxonomicName id="4C79EAD2FFD6FFB0FACDCF92205AFAA5" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="phantasma" status="sp. nov.">
<emphasis id="B90D4D43FFD6FFB0FACDCF92205AFAA5" italics="true" pageId="5" pageNumber="296">Fiorinia phantasma</emphasis>
</taxonomicName>
<emphasis id="B90D4D43FFD6FFB0FE86CE702019FAA6" bold="true" box="[268,325,1284,1305]" pageId="5" pageNumber="296">
<taxonomicNameLabel id="A23EF038FFD6FFB0FE86CE702019FAA6" box="[268,325,1284,1305]" pageId="5" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
End of abdomen of adult female. A. Antennae, adult female. 7.
<taxonomicName id="4C79EAD2FFD6FFB0FC47CE7125CBFAA5" box="[973,1175,1284,1306]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="phantasma" status="sp. nov.">
<emphasis id="B90D4D43FFD6FFB0FC47CE7125CBFAA5" box="[973,1175,1284,1306]" italics="true" pageId="5" pageNumber="296">Fiorinia phantasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23EF038FFD6FFB0FB17CE712588FAA5" box="[1181,1236,1285,1306]" pageId="5" pageNumber="296" rank="species">n. sp.</taxonomicNameLabel>
End of abdomen of second stage female.”
</paragraph>
</caption>
<subSubSection id="C363C2DAFFD6FFB2FF4DCE112094FCC0" lastPageId="7" lastPageNumber="298" pageId="5" pageNumber="296" type="diagnosis">
<paragraph id="8BC69151FFD6FFB0FF4DCE112224FA55" blockId="5.[151,1436,1381,2018]" pageId="5" pageNumber="296">
<emphasis id="B90D4D43FFD6FFB0FF4DCE112066FAC1" bold="true" box="[199,314,1381,1406]" pageId="5" pageNumber="296">Diagnosis</emphasis>
(after
<bibRefCitation id="EFE8ECA0FFD6FFB0FE0ECE1123F6FAC1" author="Williams" box="[388,682,1381,1406]" pageId="5" pageNumber="296" refString="Williams, D. J. &amp; Watson, G. W. (1988) The scale insects of the tropical South Pacific Region. Pt. 1. The armoured scales (Diaspididae). CAB International, Wallingford, 290 pp." type="book" year="1988">Williams &amp; Watson, 1988</bibRefCitation>
): Adult female on slide (
<figureCitation id="13428DD4FFD6FFB0FC36CE1122A1FAC1" box="[956,1021,1381,1406]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[203,1398,203,1866]" captionTargetId="figure@6.[203,1403,203,1882]" captionTargetPageId="6" captionText="FIGURE 3. Fiorinia phantasma (Cockerell &amp; Robinson); original illustration of the holotype of F. coronata from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process." httpUri="https://zenodo.org/record/244851/files/figure.png" pageId="5" pageNumber="296">Fig. 3</figureCitation>
A) about 0.65 mm long, broadly subrectangular, widest at about second abdominal segment, then narrowing abruptly to a triangular pygidium; body membranous except for lightly sclerotized pygidium; interantennal process (
<figureCitation id="13428DD4FFD6FFB0FC63CED92576FA79" box="[1001,1066,1453,1478]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[203,1398,203,1866]" captionTargetId="figure@6.[203,1403,203,1882]" captionTargetPageId="6" captionText="FIGURE 3. Fiorinia phantasma (Cockerell &amp; Robinson); original illustration of the holotype of F. coronata from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process." httpUri="https://zenodo.org/record/244851/files/figure.png" pageId="5" pageNumber="296">Fig. 3</figureCitation>
E) rounded, bearing short to long spicules, sometimes reduced in size and with only 1 or 2 spicules.
</paragraph>
<paragraph id="8BC69151FFD6FFB0FF4DCE81235FF97D" blockId="5.[151,1436,1381,2018]" pageId="5" pageNumber="296">
Pygidium (
<figureCitation id="13428DD4FFD6FFB0FEC9CE8120D8F9B1" box="[323,388,1525,1550]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[203,1398,203,1866]" captionTargetId="figure@6.[203,1403,203,1882]" captionTargetPageId="6" captionText="FIGURE 3. Fiorinia phantasma (Cockerell &amp; Robinson); original illustration of the holotype of F. coronata from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process." httpUri="https://zenodo.org/record/244851/files/figure.png" pageId="5" pageNumber="296">Fig. 3</figureCitation>
B) with notch at apex formed by parallel median lobes separated by about the width of 1 lobe; median lobes yoked by internal sclerosis, each lobe with pointed apex and 1 or 2 distal notches. Second lobes each bilobed, with inner lobule narrower than median lobe, with rounded apex approximately level with tip of median lobe; outer lobule smaller and shorter than inner lobule. Gland spines apparently absent. Long setae present between median and second lobes, and lateral to second lobes. Marginal ducts narrow, each about
<date id="FFC7B791FFD6FFB0FA8ECDF1247DF921" box="[1284,1313,1669,1694]" pageId="5" pageNumber="296">6x</date>
as long as wide, numbering 4 on each side.
</paragraph>
<paragraph id="8BC69151FFD6FFB0FF4DCDB9228DF8C9" blockId="5.[151,1436,1381,2018]" pageId="5" pageNumber="296">
Ventral surface with perivulvar pores present in 5 elongate groups. Microducts present around submargins of pygidium, forward to about abdominal segment III; others present in median areas of free abdominal segments, and lateral to labium. Duct tubercles present in single, interrupted marginal row between mesothorax to abdominal segment II. Antennae set close together on margin of head, each bearing 1 long seta (
<figureCitation id="13428DD4FFD6FFB0FBD5CC4D25FEF8ED" box="[1119,1186,1849,1874]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[203,1398,203,1866]" captionTargetId="figure@6.[203,1403,203,1882]" captionTargetPageId="6" captionText="FIGURE 3. Fiorinia phantasma (Cockerell &amp; Robinson); original illustration of the holotype of F. coronata from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process." httpUri="https://zenodo.org/record/244851/files/figure.png" pageId="5" pageNumber="296">Fig. 3</figureCitation>
E). Antennal spiracles each associated with 2 disc pores, each pore containing 3 loculi (
<figureCitation id="13428DD4FFD6FFB0FCE6CC2922F3F8C9" box="[876,943,1885,1910]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[203,1398,203,1866]" captionTargetId="figure@6.[203,1403,203,1882]" captionTargetPageId="6" captionText="FIGURE 3. Fiorinia phantasma (Cockerell &amp; Robinson); original illustration of the holotype of F. coronata from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process." httpUri="https://zenodo.org/record/244851/files/figure.png" pageId="5" pageNumber="296">Fig. 3</figureCitation>
C).
</paragraph>
<paragraph id="8BC69151FFD6FFB0FF4DCCF5226FF85D" blockId="5.[151,1436,1381,2018]" pageId="5" pageNumber="296">
Second-instar exuviae moderately sclerotized, with a small median patch of thinner cuticle near the anterior margin located approximately over interantennal process of adult female. Pygidial margin with 5 marginal macroducts (
<figureCitation id="13428DD4FFD6FFB0FEADCCBD2035F85D" box="[295,361,1993,2018]" captionStart="FIGURE 3" captionStartId="6.[151,250,1915,1937]" captionTargetBox="[203,1398,203,1866]" captionTargetId="figure@6.[203,1403,203,1882]" captionTargetPageId="6" captionText="FIGURE 3. Fiorinia phantasma (Cockerell &amp; Robinson); original illustration of the holotype of F. coronata from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process." httpUri="https://zenodo.org/record/244851/files/figure.png" pageId="5" pageNumber="296">Fig. 3</figureCitation>
D), each duct about
<date id="FFC7B791FFD6FFB0FDC6CCBE2334F85D" box="[588,616,1994,2018]" pageId="5" pageNumber="296">2x</date>
longer than wide.
</paragraph>
<caption id="DF06C1D9FFD5FFB3FF1DCC0F2020F853" httpUri="https://zenodo.org/record/244851/files/figure.png" pageId="6" pageNumber="297" targetBox="[203,1398,203,1866]" targetPageId="6">
<paragraph id="8BC69151FFD5FFB3FF1DCC0F2020F853" blockId="6.[151,1436,1915,2028]" pageId="6" pageNumber="297">
<emphasis id="B90D4D43FFD5FFB3FF1DCC0F2051F82F" bold="true" box="[151,269,1915,1937]" pageId="6" pageNumber="297">FIGURE 3</emphasis>
.
<taxonomicName id="4C79EAD2FFD5FFB3FE90CC0823BFF82E" authority="Cockerell &amp; Robinson" authorityName="Cockerell &amp; Robinson" box="[282,739,1915,1937]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="297" phylum="Arthropoda" rank="species" species="phantasma">
<emphasis id="B90D4D43FFD5FFB3FE90CC0820B8F82E" box="[282,484,1915,1937]" italics="true" pageId="6" pageNumber="297">Fiorinia phantasma</emphasis>
(Cockerell &amp; Robinson)
</taxonomicName>
; original illustration of the holotype of
<taxonomicName id="4C79EAD2FFD5FFB3FBFECC0825BBF82E" box="[1140,1255,1916,1937]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="297" phylum="Arthropoda" rank="species" species="coronata">
<emphasis id="B90D4D43FFD5FFB3FBFECC0825BBF82E" box="[1140,1255,1916,1937]" italics="true" pageId="6" pageNumber="297">F. coronata</emphasis>
</taxonomicName>
from Williams &amp; Watson (1988), with dorsal features shown on the left side and ventral characteristics on the right. A = body of adult female; B = detail of pygidium; C = detail of anterior spiracle; D = Dorsum of second-instar female pygidium, E = detail of antennae and inter-antennal process.
</paragraph>
</caption>
<paragraph id="8BC69151FFD4FFB2FF4DCBEC236AFF47" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="298">
An identification key to adult females of the better-documented species of
<taxonomicName id="4C79EAD2FFD4FFB2FBC7CBED25F6FF0F" box="[1101,1194,153,176]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="genus">
<emphasis id="B90D4D43FFD4FFB2FBC7CBED25F6FF0F" box="[1101,1194,153,176]" italics="true" pageId="7" pageNumber="298">Fiorinia</emphasis>
</taxonomicName>
in the Australasian, Nearctic and Neotropical Regions is provided. It has not been possible to include the following four species in the key, for the reasons discussed below:
</paragraph>
<paragraph id="8BC69151FFD4FFB2FF4DCA702527FE37" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="298">
(1)
<taxonomicName id="4C79EAD2FFD4FFB2FF7BCA7123E6FEA3" authority="Lindinger, 1911" authorityName="Lindinger" authorityYear="1911" box="[241,698,260,285]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="species" species="neocaledonica">
<emphasis id="B90D4D43FFD4FFB2FF7BCA7120A0FEA2" box="[241,508,260,285]" italics="true" pageId="7" pageNumber="298">Fiorinia neocaledonica</emphasis>
<bibRefCitation id="EFE8ECA0FFD4FFB2FD8FCA7023E6FEA3" author="Lindinger" box="[517,698,260,285]" pageId="7" pageNumber="298" refString="Lindinger, L. (1911) Beitrage zur Kenntnis der Schildlause und ihre Verbreitung II. Zeitschrift fur Wissenschaftliche Insektenbiologie, 7, 9 - 12, 86 - 90, 126 - 130, 172 - 177." type="journal article" year="1911">Lindinger, 1911</bibRefCitation>
</taxonomicName>
on
<taxonomicName id="4C79EAD2FFD4FFB2FD63CA702517FEA3" box="[745,1099,260,285]" class="Magnoliopsida" family="Myrtaceae" genus="Baeckia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="298" phylum="Magnoliophyta" rank="species" species="pinifoliae">
<emphasis id="B90D4D43FFD4FFB2FD63CA7022E5FEA2" box="[745,953,260,285]" italics="true" pageId="7" pageNumber="298">Baeckia pinifoliae</emphasis>
(Myrtaceae)
</taxonomicName>
in
<collectingCountry id="F36ED1C1FFD4FFB2FBFECA712474FEA2" box="[1140,1320,260,285]" name="New Caledonia" pageId="7" pageNumber="298">New Caledonia</collectingCountry>
, a species with black second instar exuviae coated with white secretion (
<bibRefCitation id="EFE8ECA0FFD4FFB2FCDFCA532554FEFF" author="Lindinger" box="[853,1032,295,320]" pageId="7" pageNumber="298" refString="Lindinger, L. (1911) Beitrage zur Kenntnis der Schildlause und ihre Verbreitung II. Zeitschrift fur Wissenschaftliche Insektenbiologie, 7, 9 - 12, 86 - 90, 126 - 130, 172 - 177." type="journal article" year="1911">Lindinger 1911</bibRefCitation>
). Mille
<emphasis id="B90D4D43FFD4FFB2FBEACA5D25C6FEFF" box="[1120,1178,295,320]" italics="true" pageId="7" pageNumber="298">et al.</emphasis>
(in press) say that it is an endemic species found only in natural habitats in
<collectingCountry id="F36ED1C1FFD4FFB2FC88CA3922EBFEDA" box="[770,951,332,357]" name="New Caledonia" pageId="7" pageNumber="298">New Caledonia</collectingCountry>
, but unfortunately do not comment on its morphology or appearance in life. No material of this species has been seen in this study.
</paragraph>
<paragraph id="8BC69151FFD4FFB2FF4DCAE02393FDA7" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="298">
(2)
<taxonomicName id="4C79EAD2FFD4FFB2FF73CAE123D3FE13" authority="Leonardi, 1907" authorityName="Leonardi" authorityYear="1907" box="[249,655,404,429]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="species" species="iavanica">
<emphasis id="B90D4D43FFD4FFB2FF73CAE12095FE13" box="[249,457,405,428]" italics="true" pageId="7" pageNumber="298">Fiorinia iavanica</emphasis>
Leonardi, 1907
</taxonomicName>
on
<taxonomicName id="4C79EAD2FFD4FFB2FD44CAE023A4FE12" box="[718,760,404,429]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="7" pageNumber="298" phylum="Tracheophyta" rank="genus">
<emphasis id="B90D4D43FFD4FFB2FD44CAE023A4FE12" box="[718,760,404,429]" italics="true" pageId="7" pageNumber="298">Ilex</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C79EAD2FFD4FFB2FCC8CAE022BFFE12" box="[834,995,404,429]" class="Magnoliopsida" family="Aquifoliaceae" kingdom="Plantae" order="Aquifoliales" pageId="7" pageNumber="298" phylum="Tracheophyta" rank="family">Aquifoliaceae</taxonomicName>
) in
<collectingCountry id="F36ED1C1FFD4FFB2FBA9CAE025CCFE12" box="[1059,1168,404,429]" name="Indonesia" pageId="7" pageNumber="298">Indonesia</collectingCountry>
(Java). According to
<bibRefCitation id="EFE8ECA0FFD4FFB2FF1DCAC320D4FE6F" author="MacGillivray" box="[151,392,439,464]" pageId="7" pageNumber="298" refString="MacGillivray, A. D. (1921) The Coccidae. Tables for the Identification of the subfamilies and some of the more important genera and species, together with discussions … Scarab, Urbana, Illinois, 502 pp." type="book" year="1921">MacGillivray (1921)</bibRefCitation>
, median lobes rounded, without notches, divergent; second lobes rounded, not notched; perivulvar pores in 3 groups, numbering 33–35; and with 7–10 pores associated with each anterior spiracle. No material of this species has been seen in this study.
</paragraph>
<paragraph id="8BC69151FFD4FFB2FF4DC95022C7FCAA" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="298">
(3)
<taxonomicName id="4C79EAD2FFD4FFB2FF79C9512328FD83" authority="Froggatt, 1914" authorityName="Froggatt" authorityYear="1914" box="[243,628,548,573]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="species" species="geijeriae">
<emphasis id="B90D4D43FFD4FFB2FF79C951209CFD83" box="[243,448,549,572]" italics="true" pageId="7" pageNumber="298">Fiorinia geijeriae</emphasis>
<bibRefCitation id="EFE8ECA0FFD4FFB2FE40C9512328FD83" author="Froggatt" box="[458,628,548,573]" pageId="7" pageNumber="298" refString="Froggatt, W. W. (1914) Descriptive catalogue of the scale insects (&quot; Coccidae &quot;) of Australia (Part I). The Agricultural Gazette of New South Wales, 25, 127 - 136." type="journal article" year="1914">Froggatt, 1914</bibRefCitation>
</taxonomicName>
on
<taxonomicName id="4C79EAD2FFD4FFB2FD2CC95122A0FD83" box="[678,1020,548,573]" class="Magnoliopsida" family="Rutaceae" genus="Geijera" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="298" phylum="Tracheophyta" rank="species" species="salicifolia">
<emphasis id="B90D4D43FFD4FFB2FD2CC951222BFD82" box="[678,887,548,573]" italics="true" pageId="7" pageNumber="298">Geijera salicifolia</emphasis>
(Rutaceae)
</taxonomicName>
in
<collectingCountry id="F36ED1C1FFD4FFB2FBA2C95025D3FD82" box="[1064,1167,548,573]" name="Australia" pageId="7" pageNumber="298">Australia</collectingCountry>
(New South
<collectingCountry id="F36ED1C1FFD4FFB2FABAC9502424FD82" box="[1328,1400,548,573]" name="United Kingdom" pageId="7" pageNumber="298">Wales</collectingCountry>
), a species with second-instar exuviae black except for a narrow brown margin. Examination of a slide-mounted
<typeStatus id="54C22FF3FFD4FFB2FF1DC91921B2FD3A" box="[151,238,621,645]" pageId="7" pageNumber="298" type="syntype">syntype</typeStatus>
specimen and three adult females newly mounted from the original sample (BMNH), and non-type slidemounted specimens (USNM) indicate that this species is not a member of the genus
<taxonomicName id="4C79EAD2FFD4FFB2FBC8C9E525C3FD17" box="[1090,1183,657,680]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="genus">
<emphasis id="B90D4D43FFD4FFB2FBC8C9E525C3FD17" box="[1090,1183,657,680]" italics="true" pageId="7" pageNumber="298">Fiorinia</emphasis>
</taxonomicName>
. The median lobes are fused together and there is a pair of clavate, paraphysis-like internal sclerotizations on the pygidial margin. Examination of dry specimens from the original sample (BMNH) indicates that
<taxonomicName id="4C79EAD2FFD4FFB2FBA9C9AD25FCFD4F" box="[1059,1184,729,752]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="subSpecies" species="gei" subSpecies="jeriae">
<emphasis id="B90D4D43FFD4FFB2FBA9C9AD25FCFD4F" box="[1059,1184,729,752]" italics="true" pageId="7" pageNumber="298">F. gei jeriae</emphasis>
</taxonomicName>
is not pupillarial, and the females form their scale covers beneath the epidermis of the leaf.
</paragraph>
<paragraph id="8BC69151FFD4FFB2FF4DC8542094FCC0" blockId="7.[151,1437,151,896]" pageId="7" pageNumber="298">
(4)
<taxonomicName id="4C79EAD2FFD4FFB2FF78C8552368FC87" authority="Maskell" authorityName="Maskell" box="[242,564,799,824]" class="Insecta" family="Diaspididae" genus="Fiorinia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="species" species="drimydis">
<emphasis id="B90D4D43FFD4FFB2FF78C85520E1FC87" box="[242,445,799,824]" italics="true" pageId="7" pageNumber="298">Fiorinia drimydis</emphasis>
(Maskell)
</taxonomicName>
was regarded as a
<emphasis id="B90D4D43FFD4FFB2FC96C85522E2FC87" box="[796,958,799,824]" italics="true" pageId="7" pageNumber="298">nomen nudum</emphasis>
by Deitz &amp; Tocker (1979) and a
<emphasis id="B90D4D43FFD4FFB2FADBC85521B7FCE2" italics="true" pageId="7" pageNumber="298">nomen dubium</emphasis>
by
<bibRefCitation id="EFE8ECA0FFD4FFB2FE9DC83020B9FCE3" author="Henderson" box="[279,485,836,861]" pageId="7" pageNumber="298" refString="Henderson, R. C. (2011) Diaspididae (Insecta: Hemiptera: Coccoidea). Fauna of New Zealand, 66, 1 - 275." type="journal article" year="2011">Henderson (2011)</bibRefCitation>
, the only Maskell-collected material available to her being determined as immature specimens of
<taxonomicName id="4C79EAD2FFD4FFB2FEBBC81D20FDFC3F" box="[305,417,873,896]" class="Insecta" family="Diaspididae" genus="Leucaspis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="298" phylum="Arthropoda" rank="genus">
<emphasis id="B90D4D43FFD4FFB2FEBBC81D20FDFC3F" box="[305,417,873,896]" italics="true" pageId="7" pageNumber="298">Leucaspis</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
</treatment>
</document>
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