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<document id="4F10E3C668B88869C13DCEE3B1724F69" ID-DOI="10.1080/00222933.2018.1435833" ID-ISSN="1464-5262" ID-Zenodo-Dep="5174451" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628558511936" checkinUser="felipe" docAuthor="Wagner, Philipp, Greenbaum, Eli, Bauer, Aaron M., Kusamba, Chifundera &amp; Leaché, Adam D." docDate="2018" docId="03BFD058FFBBFF8814C78BDABA3EFD27" docLanguage="en" docName="JNatHist.52.13-16.771-817.pdf" docOrigin="Journal of Natural History 52 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2018.1435833" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Acanthocercus gregorii" docType="treatment" docVersion="4" lastPageNumber="805" masterDocId="FF86A820FF9BFFAB154A8901BE4DFF9E" masterDocTitle="Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)" masterLastPageNumber="817" masterPageNumber="771" pageNumber="802" updateTime="1699046512504" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9E9D5923A63928714D73950F64054703">Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)</mods:title>
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<heading id="D0E1D622FFBBFF8B14C78BDABDDBFD6B" box="[397,918,731,757]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="802" reason="5">
<treatmentCitationGroup id="AB064660FFBBFF8B14C78BDABDDBFD6B" box="[397,918,731,757]" pageId="32" pageNumber="802">
<treatmentCitation id="0AB7475FFFBBFF8B14C78BDABDDBFD6B" author="Gunther A" box="[397,918,731,757]" page="86" pageId="32" pageNumber="802" year="1894">
<taxonomicName id="4C161ACDFFBBFF8B14C78BDABDDBFD6B" authority="(Gunther, 1894: 86)" baseAuthorityName="Gunther" baseAuthorityPageNumber="86" baseAuthorityYear="1894" box="[397,918,731,757]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="802" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFBBFF8B14C78BDABCE5FD6B" bold="true" box="[397,680,731,757]" italics="true" pageId="32" pageNumber="802">Acanthocercus gregorii</emphasis>
(
<bibRefCitation id="EF871CBFFFBBFF8B17F28BDDBDDDFD6B" author="Gunther A" box="[696,912,732,757]" pageId="32" pageNumber="802" pagination="84 - 91" refId="ref20109" refString="Gunther A. 1894. Report on the collection of reptiles and fishes made by Dr. J. W. Gregory during his expedition to Mount Kenia [sic]. Proc Zool Soc London. 1894: 84 - 91." type="journal article" year="1894">Günther, 1894: 86</bibRefCitation>
)
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<paragraph id="8BA9614EFFBBFF8B17098BFEBCACFC86" blockId="32.[397,918,731,792]" box="[579,737,767,792]" pageId="32" pageNumber="802">
(
<figureCitation id="132D7DCBFFBBFF8B17018BFEBCF6FC86" box="[587,699,767,792]" captionStart="Figure 13" captionStartId="32.[160,225,1645,1668]" captionTargetBox="[163,1152,1021,1615]" captionTargetId="figure-204@32.[163,1152,1021,1615]" captionTargetPageId="32" captionText="Figure 13. (a) Lectotype (BMNH 1946.8.28.67) of Acanthocercus gregorii, from Mkonumbi, Kenya. (b) Holotype (SMF 10138) of Acanthocercus atricollis loveridgei, from Kakoma, O-Afrika [=Tanzania], East Africa." figureDoi="http://doi.org/10.5281/zenodo.5174479" httpUri="https://zenodo.org/record/5174479/files/figure.png" pageId="32" pageNumber="802">Figure 13</figureCitation>
(a))
</paragraph>
<paragraph id="8BA9614EFFBBFF8B15EA8A42BC49FC3E" blockId="32.[160,1155,834,963]" pageId="32" pageNumber="802">
1894
<taxonomicName id="4C161ACDFFBBFF8B15948A42BC0FFCC5" authority="Gunther, Report" authorityName="Gunther, Report" authorityYear="1894" box="[222,578,835,859]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="802" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFBBFF8B15948A42BFCAFCC5" box="[222,391,835,859]" italics="true" pageId="32" pageNumber="802">Agama gregorii</emphasis>
Günther, Report
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on the collection of reptiles and fishes made by Dr J. W. Gregory during his expedition to Mount
<collectingCountry id="F30121DEFFBBFF8B17FD8A64BCB8FCE3" box="[695,757,869,893]" name="Kenya" pageId="32" pageNumber="802">Kenia</collectingCountry>
[sic]. Proceedings of the Zoological Society, London 1894: 8491.
</paragraph>
<paragraph id="8BA9614EFFBBFF8A15EA8AAABC83FF72" blockId="32.[160,1155,834,963]" lastBlockId="33.[183,1156,142,236]" lastPageId="33" lastPageNumber="803" pageId="32" pageNumber="802">
1957
<taxonomicName id="4C161ACDFFBBFF8B15A88AAABA40FC5D" authority="Klausewitz, Eidonomische Untersuchungen" authorityName="Klausewitz, Eidonomische Untersuchungen" authorityYear="1957" box="[226,1037,939,963]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="802" phylum="Chordata" rank="subSpecies" species="atricollis" subSpecies="loveridgei">
<emphasis id="B962BD5CFFBBFF8B15A88AAABC42FC5D" box="[226,527,939,963]" italics="true" pageId="32" pageNumber="802">Agama atricollis loveridgei</emphasis>
Klausewitz, Eidonomische Untersuchungen
</taxonomicName>
über die Rassenkreise
<taxonomicName id="4C161ACDFFBAFF8A1408898FBC5FFF38" authorityName="Ruppell" authorityYear="1835" box="[322,530,142,166]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="cyanogaster">
<emphasis id="B962BD5CFFBAFF8A1408898FBC5FFF38" box="[322,530,142,166]" italics="true" pageId="33" pageNumber="803">Agama cyanogaster</emphasis>
</taxonomicName>
und
<taxonomicName id="4C161ACDFFBAFF8A170E898FBCA7FF38" authorityName="Smith, Illustrations" authorityYear="1849" box="[580,746,142,166]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFBAFF8A170E898FBCA7FF38" box="[580,746,142,166]" italics="true" pageId="33" pageNumber="803">Agama atricollis</emphasis>
</taxonomicName>
. 2. Die Unterarten von
<taxonomicName id="4C161ACDFFBAFF8A1692898FBA33FF38" authorityName="Smith, Illustrations" authorityYear="1849" box="[984,1150,142,166]" class="Reptilia" family="Agamidae" genus="Agama" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFBAFF8A1692898FBA33FF38" box="[984,1150,142,166]" italics="true" pageId="33" pageNumber="803">Agama atricollis</emphasis>
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. Senckenbergiana biologica 38: 157174: 163.
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: SMF 10138, from Kakoma, O-Afrika [=
<collectingCountry id="F30121DEFFBAFF8A147889D5BFD2FF72" box="[306,415,212,236]" name="Tanzania" pageId="33" pageNumber="803">Tanzania</collectingCountry>
], East Africa (
<figureCitation id="132D7DCBFFBAFF8A177D89D5BCEEFF72" box="[567,675,212,236]" captionStart="Figure 13" captionStartId="32.[160,225,1645,1668]" captionTargetBox="[163,1152,1021,1615]" captionTargetId="figure-204@32.[163,1152,1021,1615]" captionTargetPageId="32" captionText="Figure 13. (a) Lectotype (BMNH 1946.8.28.67) of Acanthocercus gregorii, from Mkonumbi, Kenya. (b) Holotype (SMF 10138) of Acanthocercus atricollis loveridgei, from Kakoma, O-Afrika [=Tanzania], East Africa." figureDoi="http://doi.org/10.5281/zenodo.5174479" httpUri="https://zenodo.org/record/5174479/files/figure.png" pageId="33" pageNumber="803">Figure 13</figureCitation>
(b)).
</paragraph>
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<emphasis id="B962BD5CFFBBFF8B15EA8F6CBF45F91A" bold="true" box="[160,264,1645,1668]" pageId="32" pageNumber="802">Figure 13.</emphasis>
(a) Lectotype (BMNH 1946.8.28.67) of
<taxonomicName id="4C161ACDFFBBFF8B17C18F6CBD2BF91A" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[651,870,1645,1668]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="802" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFBBFF8B17C18F6CBD2BF91A" box="[651,870,1645,1668]" italics="true" pageId="32" pageNumber="802">Acanthocercus gregorii</emphasis>
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, from Mkonumbi, Kenya. (b) Holotype (SMF 10138) of
<taxonomicName id="4C161ACDFFBBFF8B14D68F8BBC92F93F" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[412,735,1674,1697]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="802" phylum="Chordata" rank="subSpecies" species="atricollis" subSpecies="loveridgei">
<emphasis id="B962BD5CFFBBFF8B14D68F8BBC92F93F" box="[412,735,1674,1697]" italics="true" pageId="32" pageNumber="802">Acanthocercus atricollis loveridgei</emphasis>
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, from
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Kakoma, O-Afrika
<emphasis id="B962BD5CFFBBFF8B169A8F8ABD98F93C" box="[976,981,1675,1698]" italics="true" pageId="32" pageNumber="802"></emphasis>
[=Tanzania], East Africa.
</paragraph>
</caption>
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<typeStatus id="54ADDFECFFBAFF8A15EA8812BF5FFEB2" box="[160,274,275,300]" pageId="33" pageNumber="803" type="lectotype">Lectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA9614EFFBAFF8A15EA8836BF93FEEF" blockId="33.[160,1155,275,369]" pageId="33" pageNumber="803">
<materialsCitation id="3B7E6B13FFBAFF8A15EA8836BF93FEEF" collectionCode="BMNH" country="Kenya" county="East Africa." location="Mkonumbi" municipality="Coast Province" pageId="33" pageNumber="803" specimenCode="BMNH 1946.8" specimenCount="1" typeStatus="lectotype">
<specimenCode id="DBB0C935FFBAFF8A15EA8836BF73FED1" box="[160,318,311,335]" collectionCode="BMNH" pageId="33" pageNumber="803">BMNH 1946.8</specimenCode>
.28.67, from
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, a grassy coast district with salt-swamps,
<collectingMunicipality id="6BCDFB34FFBAFF8A110F8836BF4EFEEF" pageId="33" pageNumber="803">Coast Province</collectingMunicipality>
,
<collectingCountry id="F30121DEFFBAFF8A14458858BF1BFEEF" box="[271,342,345,369]" name="Kenya" pageId="33" pageNumber="803">Kenya</collectingCountry>
,
<collectingCounty id="62C819C2FFBAFF8A142B8858BF93FEEF" box="[353,478,345,369]" pageId="33" pageNumber="803">East Africa.</collectingCounty>
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</paragraph>
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<emphasis id="B962BD5CFFBAFF8A15EA8898BF6BFE2C" bold="true" box="[160,294,409,434]" italics="true" pageId="33" pageNumber="803">Description</emphasis>
</paragraph>
<paragraph id="8BA9614EFFBAFF8A15EA88BDBDDAFB32" blockId="33.[160,1157,409,1439]" pageId="33" pageNumber="803">
The largest species (total length of adult males up to
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[adult males SVL:
<quantity id="4CEECCABFFBAFF8A15EA88DEBF64FE69" box="[160,297,479,503]" metricMagnitude="-1" metricUnit="m" metricValue="1.145" metricValueMax="1.38" metricValueMin="0.9099999999999999" pageId="33" pageNumber="803" unit="mm" value="114.5" valueMax="138.0" valueMin="91.0">91138 mm</quantity>
, x =
<quantity id="4CEECCABFFBAFF8A142B88DEBF99FE69" box="[353,468,479,503]" metricMagnitude="-1" metricUnit="m" metricValue="1.192" pageId="33" pageNumber="803" unit="mm" value="119.2">119.2 mm</quantity>
, n = 23], adult females to
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) of the genus, but similar in body size to
<taxonomicName id="4C161ACDFFBAFF8A14058B03BF8BFD84" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[335,454,514,538]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFBAFF8A14058B03BF8BFD84" box="[335,454,514,538]" italics="true" pageId="33" pageNumber="803">A. atricollis</emphasis>
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; tail a third to a half longer than the SVL, but there is a series of specimens from one locality in the Serengeti that have extremely short tails, shorter than the SVL. Head distinctly broad in males, lacking the occipital scale. Ear opening as large as the eye, tympanum conspicuous. Nasal scale not convex, smooth and slightly below the canthus rostralis. Nuchal crest absent. Distinct from
<taxonomicName id="4C161ACDFFBAFF8A16268B8DBDAEFD3A" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[876,995,652,676]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFBAFF8A16268B8DBDAEFD3A" box="[876,995,652,676]" italics="true" pageId="33" pageNumber="803">A. atricollis</emphasis>
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in possessing many enlarged, white coloured dorsal scales, forming a dotted pattern (=
<taxonomicName id="4C161ACDFFBAFF8A16B18BAEBACEFD59" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[1019,1155,687,711]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="loveridgei">
<emphasis id="B962BD5CFFBAFF8A16B18BAEBACEFD59" box="[1019,1155,687,711]" italics="true" pageId="33" pageNumber="803">A. loveridgei</emphasis>
</taxonomicName>
morphotype) or forming a dotted pattern with up to five transverse scale rows (=
<taxonomicName id="4C161ACDFFBAFF8A15F28BF5BF61FC92" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[184,300,756,780]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFBAFF8A15F28BF5BF61FC92" box="[184,300,756,780]" italics="true" pageId="33" pageNumber="803">A. gregorii</emphasis>
</taxonomicName>
morphotype). Body scalation heterogeneous, dorsal matrix scales small, smaller than ventral scales, irregularly intermixed by enlarged scales, sometimes forming distinct transverse scale rows. Enlarged scales keeled, most concentrated at the vertebrate. Vertebral zone covered by enlarged scales intermixed with smaller matrix scales, not separated by distinct rows of enlarged scales from the lateral parts of the body. Body scales are arranged in 88126 (x = 103.8, n = 23) rows around midbody and 5285 (x = 66.5, n = 23) longitudinal rows along the vertebrae. Ventral scales smooth, smaller than the enlarged dorsal scales, but larger than the matrix scales, in 7792 (x = 83.3, n = 23) longitudinal rows. Gular scales small, smooth to feebly keeled, but erect and about the same size as the ventral scales. Males with one to three continuous rows (x = 2.0, n = 22) with a total number of 737 (x = 19.0, n = 22) precloacal pores, lacking in females. Dorsal scales of the basal part of the tail larger than those on the other parts of the tail, often forming a distinct patch of thick and swollen scales.
</paragraph>
<paragraph id="8BA9614EFFBAFF8A158A8DB6BFF8FA01" blockId="33.[160,1157,409,1439]" pageId="33" pageNumber="803">
The coloration in life is poorly known.
<bibRefCitation id="EF871CBFFFBAFF8A17CA8DB6BD60FB51" author="Gunther A" box="[640,813,1207,1231]" pageId="33" pageNumber="803" pagination="84 - 91" refId="ref20109" refString="Gunther A. 1894. Report on the collection of reptiles and fishes made by Dr. J. W. Gregory during his expedition to Mount Kenia [sic]. Proc Zool Soc London. 1894: 84 - 91." type="journal article" year="1894">Günther (1894)</bibRefCitation>
, in the first description, mentioned the throat and upper parts of the body bluish, but head, enlarged scales and basal portion of the tail yellow, while
<bibRefCitation id="EF871CBFFFBAFF8A17108DFDBD5DFA8A" author="Sternfeld R" box="[602,784,1276,1300]" pageId="33" pageNumber="803" refId="ref21426" refString="Sternfeld R. 1912. Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika Expedition 1907 - 1908, unter Fuhrung A. Friedrichs, Herzogs zu Mecklenburg. Klinkhard und Biermann, Leipzig. IV. Zoologie II. Lfg. Reptilia; p. 197 - 279 [the volume was published in parts from 1911 to 1913 but this part was published in 1912]." type="book" year="1912">Sternfeld (1912)</bibRefCitation>
described the throat, chest and sometimes belly as reticulated blue and the dorsally enlarged body scales yellow. According to
<bibRefCitation id="EF871CBFFFBAFF8A140F8C40BC42FAC7" author="Klausewitz W" box="[325,527,1345,1369]" pageId="33" pageNumber="803" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz (1957)</bibRefCitation>
the throat of adult males is uniform greenish blue, head bluish green, body uniform blue and the tail bi-coloured yellowish at the base and brownish at the tip.
</paragraph>
</subSubSection>
<subSubSection id="C30C32C5FFBAFF8815EA8CD7BA3EFD27" lastPageId="35" lastPageNumber="805" pageId="33" pageNumber="803" type="distribution">
<paragraph id="8BA9614EFFBAFF8A15EA8CD7BF61FA71" blockId="33.[160,1155,1494,1727]" box="[160,300,1494,1519]" pageId="33" pageNumber="803">
<emphasis id="B962BD5CFFBAFF8A15EA8CD7BF61FA71" bold="true" box="[160,300,1494,1519]" italics="true" pageId="33" pageNumber="803">Distribution</emphasis>
</paragraph>
<paragraph id="8BA9614EFFBAFF8815EA8CFBBDB3FE07" blockId="33.[160,1155,1494,1727]" lastBlockId="35.[160,1156,142,409]" lastPageId="35" lastPageNumber="805" pageId="33" pageNumber="803">
<taxonomicName id="4C161ACDFFBAFF8A15EA8CFBBFEFF98C" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[160,418,1530,1554]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFBAFF8A15EA8CFBBFEFF98C" box="[160,418,1530,1554]" italics="true" pageId="33" pageNumber="803">Acanthocercus gregorii</emphasis>
</taxonomicName>
has the widest distribution within the
<taxonomicName id="4C161ACDFFBAFF8A16D18CFBBA56F98C" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[923,1051,1530,1554]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFBAFF8A16D18CFBBA56F98C" box="[923,1051,1530,1554]" italics="true" pageId="33" pageNumber="803">A. atricollis</emphasis>
</taxonomicName>
-complex (
<figureCitation id="132D7DCBFFBAFF8A15ED8F1CBF59F9AB" box="[167,276,1565,1589]" captionStart="Figure 14" captionStartId="34.[160,225,1390,1413]" captionTargetBox="[178,1138,195,1358]" captionTargetId="figure-2@34.[178,1138,195,1359]" captionTargetPageId="34" captionText="Figure 14. Distribution of Acanthocercus gregorii and A. minutus. The numbers refer to the specific localities." figureDoi="http://doi.org/10.5281/zenodo.5174481" httpUri="https://zenodo.org/record/5174481/files/figure.png" pageId="33" pageNumber="803">Figure 14</figureCitation>
). It is known from
<collectingCountry id="F30121DEFFBAFF8A14BF8F1CBC19F9AB" box="[501,596,1565,1589]" name="Ethiopia" pageId="33" pageNumber="803">Ethiopia</collectingCountry>
,
<collectingCountry id="F30121DEFFBAFF8A172A8F1CBCEAF9AB" box="[608,679,1565,1589]" name="Kenya" pageId="33" pageNumber="803">Kenya</collectingCountry>
,
<collectingCountry id="F30121DEFFBAFF8A17FE8F1CBD57F9AB" box="[692,794,1565,1589]" name="Tanzania" pageId="33" pageNumber="803">Tanzania</collectingCountry>
,
<collectingCountry id="F30121DEFFBAFF8A166D8F1CBDCEF9AB" box="[807,899,1565,1589]" name="Rwanda" pageId="33" pageNumber="803">Rwanda</collectingCountry>
,
<collectingCountry id="F30121DEFFBAFF8A16DA8F1CBDA4F9AB" box="[912,1001,1565,1589]" name="Burundi" pageId="33" pageNumber="803">Burundi</collectingCountry>
and extreme northern
<collectingCountry id="F30121DEFFBAFF8A145E8F3EBFE7F9C9" box="[276,426,1599,1623]" name="Mozambique" pageId="33" pageNumber="803">Mozambique</collectingCountry>
.
<bibRefCitation id="EF871CBFFFBAFF8A14F58F3EBCC0F9C9" author="Klausewitz W" box="[447,653,1599,1623]" pageId="33" pageNumber="803" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz (1957)</bibRefCitation>
mentioned specimens from
<collectingCountry id="F30121DEFFBAFF8A16B18F3EBA1EF9C9" box="[1019,1107,1599,1623]" name="Uganda" pageId="33" pageNumber="803">Uganda</collectingCountry>
(as
<taxonomicName id="4C161ACDFFBAFF8A15EA8F63BF6AF9E4" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[160,295,1634,1658]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="loveridgei">
<emphasis id="B962BD5CFFBAFF8A15EA8F63BF6AF9E4" box="[160,295,1634,1658]" italics="true" pageId="33" pageNumber="803">A. loveridgei</emphasis>
</taxonomicName>
) but it was not possible to confirm this and the presence of
<taxonomicName id="4C161ACDFFBAFF8A16B98F63BA29F9E4" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[1011,1124,1634,1658]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFBAFF8A16B98F63BA29F9E4" box="[1011,1124,1634,1658]" italics="true" pageId="33" pageNumber="803">A. gregorii</emphasis>
</taxonomicName>
in
<collectingCountry id="F30121DEFFBAFF8A15EA8F84BEB5F903" box="[160,248,1669,1693]" name="Uganda" pageId="33" pageNumber="803">Uganda</collectingCountry>
is dubious. He (
<bibRefCitation id="EF871CBFFFBAFF8A14838F84BCC5F903" author="Klausewitz W" box="[457,648,1669,1693]" pageId="33" pageNumber="803" pagination="157 - 174" refId="ref20422" refString="Klausewitz W. 1957. Eidonomische Untersuchungen uber die Rassenkreise Agama cyanogaster und Agama atricollis. Die Unterarten von Agama atricollis [Eidonomic studies about the species complex of Agama cyanogaster and Agama atricollis. The subspecies of Agama atricollis]. Senckenb Biol. 38: 157 - 174." type="journal article" year="1957">Klausewitz 1957</bibRefCitation>
) also recognized
<taxonomicName id="4C161ACDFFBAFF8A16208F84BDBAF903" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[874,1015,1669,1693]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="803" phylum="Chordata" rank="species" species="loveridgei">
<emphasis id="B962BD5CFFBAFF8A16208F84BDBAF903" box="[874,1015,1669,1693]" italics="true" pageId="33" pageNumber="803">A. loveridgei</emphasis>
</taxonomicName>
from Lake Bengweulu and the southern margins of the Luangwa Valley in
<collectingCountry id="F30121DEFFBAFF8A16C68FA6BDAEF921" box="[908,995,1703,1727]" name="Zambia" pageId="33" pageNumber="803">Zambia</collectingCountry>
. The previous 1 = Gojam; 2 = Batia; 3 = Lit Marafia; 4 = Didessa [Oromia]; 5 = Addis; 6 = Adda; 7 = Modjo; 8 = Adama [Nazareth]; 9 = Welenchiti; 10 = Akaki; 11 = Mount Zaguala; 12 = Maki River; 13 = Wonji; 14 = Harra, Lake Zuai; 15 = Lake Langano; 16 = Didessa; 17 = without locality; 18 = Lake Awasa; 19 = between Dodola and Adaba; 20 = Dodola; 21 = Mount Gaysay; 22 = Dinshu; 23 = Sheikh Husein; 24 = Sidam Bale bridge; 25 = Kebre Mengist; 26 = Felenguai; 27 = just N of Neghelli; 28 = Neghelli; 29 = Huri Hills; 30 = Moyala; 31 =
<collectingRegion id="49D2AFACFFB9FF8916F28F1BBA4BF9B0" box="[952,1030,1562,1582]" country="Kenya" name="Marsabit" pageId="34" pageNumber="804">Marsabit</collectingRegion>
; 32 = Elgonyi; 33 = Maralal; 34 = Ntumot Lagga; 35 = Gorba Tula; 36 = Guaso Nyiro; 37 =
<collectingRegion id="49D2AFACFFB9FF8916098F34BDC8F9D7" box="[835,901,1589,1609]" country="Kenya" name="Kericho" pageId="34" pageNumber="804">Kericho</collectingRegion>
Valley; 38 = Kirui; 39 = Lake Elmenteita; 40 = Naivasha; 41 = Fort Hall; 42 = Thika; 43 = Nairobi; 44 = Nairobi National Park; 45 = Attui; 46 =
<collectingRegion id="49D2AFACFFB9FF8915928F6BBF43F9E0" box="[216,270,1642,1662]" country="Kenya" name="Narok" pageId="34" pageNumber="804">Narok</collectingRegion>
; 47 = South N
<emphasis id="B962BD5CFFB9FF8914EA8F6BBFE9F9E0" box="[416,420,1642,1662]" italics="true" pageId="34" pageNumber="804"></emphasis>
guassa Mjiro River; 48 = Ngong Hills; 49 = SW of Ngong Hills; 50 =
<collectingRegion id="49D2AFACFFB9FF8911748F6BBACCF9E0" box="[1086,1153,1642,1662]" country="Kenya" name="Kajiado" pageId="34" pageNumber="804">Kajiado</collectingRegion>
; 51 = Sultan Hamud; 52 = Namanga; 53 = Makneni; 54 =
<collectingRegion id="49D2AFACFFB9FF8917D68F84BC80F907" box="[668,717,1669,1689]" country="Kenya" name="Lamu" pageId="34" pageNumber="804">Lamu</collectingRegion>
island; 55 = Mkonumbi; 56 = Gede; 57 =
<collectingCountry id="F30121DEFFB9FF8911088F84BACEF907" box="[1090,1155,1669,1689]" name="Kenya" pageId="34" pageNumber="804">Malindi</collectingCountry>
district; 58 = Mount Sagalla; 59 =
<collectingCountry id="F30121DEFFB9FF89149D8FA1BC62F92A" box="[471,559,1696,1716]" name="Kenya" pageId="34" pageNumber="804">Mombasa</collectingCountry>
; 60 = Mnambe Village; 61 = Tanga; 62 = Maweni; 63 = Korogwe; record is now referred to
<taxonomicName id="4C161ACDFFB8FF8814AA898FBCEEFF38" baseAuthorityName="Falk" baseAuthorityYear="1925" box="[480,675,142,166]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="cyanocephalus">
<emphasis id="B962BD5CFFB8FF8814AA898FBCEEFF38" box="[480,675,142,166]" italics="true" pageId="35" pageNumber="805">A. cyanocephalus</emphasis>
</taxonomicName>
and the latter to
<taxonomicName id="4C161ACDFFB8FF8816C0898FBDB3FF38" authorityName="Wagner, Greenbaum &amp; Bauer, A" authorityYear="2018" box="[906,1022,142,166]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="branchi">
<emphasis id="B962BD5CFFB8FF8816C0898FBDB3FF38" box="[906,1022,142,166]" italics="true" pageId="35" pageNumber="805">A. branchi</emphasis>
</taxonomicName>
. Therefore,
<taxonomicName id="4C161ACDFFB8FF8815EC89B0BF7CFF57" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[166,305,177,201]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="loveridgei">
<emphasis id="B962BD5CFFB8FF8815EC89B0BF7CFF57" box="[166,305,177,201]" italics="true" pageId="35" pageNumber="805">A. loveridgei</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C161ACDFFB8FF88141189B0BF9DFF57" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[347,464,177,201]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFB8FF88141189B0BF9DFF57" box="[347,464,177,201]" italics="true" pageId="35" pageNumber="805">A. gregorii</emphasis>
</taxonomicName>
) does not occur in
<collectingCountry id="F30121DEFFB8FF88178089B0BD6CFF57" box="[714,801,177,201]" name="Zambia" pageId="35" pageNumber="805">Zambia</collectingCountry>
. Specimens of the
<taxonomicName id="4C161ACDFFB8FF88114489B0BACEFF57" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[1038,1155,177,201]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFB8FF88114489B0BACEFF57" box="[1038,1155,177,201]" italics="true" pageId="35" pageNumber="805">A. gregorii</emphasis>
</taxonomicName>
morphotype are known from coastal East Africa between
<collectingRegion id="49D2AFACFFB8FF88160589D5BDC0FF72" box="[847,909,212,236]" country="Kenya" name="Lamu" pageId="35" pageNumber="805">Lamu</collectingRegion>
Island,
<collectingCountry id="F30121DEFFB8FF8816A789D5BA7CFF72" box="[1005,1073,212,236]" name="Kenya" pageId="35" pageNumber="805">Kenya</collectingCountry>
in the north and
<collectingRegion id="49D2AFACFFB8FF88145189F7BF2FFE90" box="[283,354,246,270]" country="Tanzania" name="Tanga" pageId="35" pageNumber="805">Tanga</collectingRegion>
,
<collectingCountry id="F30121DEFFB8FF88142589F7BF9FFE90" box="[367,466,246,270]" name="Tanzania" pageId="35" pageNumber="805">Tanzania</collectingCountry>
in the south, as well as from two localities in the Serengeti National Park (see
<figureCitation id="132D7DCBFFB8FF8814368818BFA5FEAF" box="[380,488,281,305]" captionStart="Figure 15" captionStartId="36.[160,225,1253,1276]" captionTargetBox="[178,1138,142,1222]" captionTargetId="figure-183@36.[178,1138,142,1222]" captionTargetPageId="36" captionText="Figure 15. Holotype (ZMB 23906) of Acanthocercus kiwuensis, from Kissenji, Rwanda." figureDoi="http://doi.org/10.5281/zenodo.5174483" httpUri="https://zenodo.org/record/5174483/files/figure.png" pageId="35" pageNumber="805">Figure 15</figureCitation>
). All other records refer to the
<taxonomicName id="4C161ACDFFB8FF8816148818BDAAFEAF" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[862,999,281,305]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="loveridgei">
<emphasis id="B962BD5CFFB8FF8816148818BDAAFEAF" box="[862,999,281,305]" italics="true" pageId="35" pageNumber="805">A. loveridgei</emphasis>
</taxonomicName>
morphotype.
<taxonomicName id="4C161ACDFFB8FF8815EA883DBFD3FECA" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[160,414,316,340]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFB8FF8815EA883DBFD3FECA" box="[160,414,316,340]" italics="true" pageId="35" pageNumber="805">Acanthocercus gregorii</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="4C161ACDFFB8FF8817C8883DBCB1FECA" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[642,764,316,340]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B962BD5CFFB8FF8817C8883DBCB1FECA" box="[642,764,316,340]" italics="true" pageId="35" pageNumber="805">A. minutus</emphasis>
</taxonomicName>
in
<collectingCountry id="F30121DEFFB8FF881661883DBDC7FECA" box="[811,906,316,340]" name="Ethiopia" pageId="35" pageNumber="805">Ethiopia</collectingCountry>
, with
<taxonomicName id="4C161ACDFFB8FF881692883DBA2CFECA" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[984,1121,316,340]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="kiwuensis">
<emphasis id="B962BD5CFFB8FF881692883DBA2CFECA" box="[984,1121,316,340]" italics="true" pageId="35" pageNumber="805">A. kiwuensis</emphasis>
</taxonomicName>
in
<collectingCountry id="F30121DEFFB8FF8815EA885FBEB7FEE8" box="[160,250,350,374]" name="Rwanda" pageId="35" pageNumber="805">Rwanda</collectingCountry>
and
<collectingCountry id="F30121DEFFB8FF88140C885FBFECFEE8" box="[326,417,350,374]" name="Burundi" pageId="35" pageNumber="805">Burundi</collectingCountry>
, and with
<taxonomicName id="4C161ACDFFB8FF881778885FBCADFEE8" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[562,736,350,374]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="ugandaensis">
<emphasis id="B962BD5CFFB8FF881778885FBCADFEE8" box="[562,736,350,374]" italics="true" pageId="35" pageNumber="805">A. ugandaensis</emphasis>
</taxonomicName>
in
<collectingCountry id="F30121DEFFB8FF88165D885FBD16FEE8" box="[791,859,350,374]" name="Kenya" pageId="35" pageNumber="805">Kenya</collectingCountry>
and northern
<collectingCountry id="F30121DEFFB8FF881151885FBACCFEE8" box="[1051,1153,350,374]" name="Tanzania" pageId="35" pageNumber="805">Tanzania</collectingCountry>
. However, the detailed limits of its distribution are still incompletely known.
</paragraph>
<caption id="DF6931C6FFB9FF8915EA8C6FBEB1FA3C" ID-DOI="http://doi.org/10.5281/zenodo.5174481" ID-Zenodo-Dep="5174481" httpUri="https://zenodo.org/record/5174481/files/figure.png" pageId="34" pageNumber="804" startId="34.[160,225,1390,1413]" targetBox="[178,1138,195,1358]" targetPageId="34">
<paragraph id="8BA9614EFFB9FF8915EA8C6FBEB1FA3C" blockId="34.[160,1155,1389,1442]" pageId="34" pageNumber="804">
<emphasis id="B962BD5CFFB9FF8915EA8C6FBF44FA1B" bold="true" box="[160,265,1390,1413]" pageId="34" pageNumber="804">Figure 14.</emphasis>
Distribution of
<taxonomicName id="4C161ACDFFB9FF8914E08C6CBCCBFA1A" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[426,646,1389,1412]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="804" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFB9FF8914E08C6CBCCBFA1A" box="[426,646,1389,1412]" italics="true" pageId="34" pageNumber="804">Acanthocercus gregorii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C161ACDFFB9FF8917F68C6CBD69FA1A" baseAuthorityName="Klausewitz" baseAuthorityYear="1957" box="[700,804,1389,1412]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="804" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B962BD5CFFB9FF8917F68C6CBD69FA1A" box="[700,804,1389,1412]" italics="true" pageId="34" pageNumber="804">A. minutus</emphasis>
</taxonomicName>
. The numbers refer to the specific localities.
</paragraph>
</caption>
<paragraph id="8BA9614EFFB8FF8815EA88D1BFD8FE77" blockId="35.[160,1156,464,697]" box="[160,405,464,489]" pageId="35" pageNumber="805">
<emphasis id="B962BD5CFFB8FF8815EA88D1BFD8FE77" bold="true" box="[160,405,464,489]" italics="true" pageId="35" pageNumber="805">Habitat and ecology</emphasis>
</paragraph>
<paragraph id="8BA9614EFFB8FF8815EA88F5BA3EFD27" blockId="35.[160,1156,464,697]" pageId="35" pageNumber="805">
Detailed information is sparse and inconsistent.
<bibRefCitation id="EF871CBFFFB8FF8817F588F5BD25FD92" author="Gunther A" box="[703,872,500,524]" pageId="35" pageNumber="805" pagination="84 - 91" refId="ref20109" refString="Gunther A. 1894. Report on the collection of reptiles and fishes made by Dr. J. W. Gregory during his expedition to Mount Kenia [sic]. Proc Zool Soc London. 1894: 84 - 91." type="journal article" year="1894">Günther (1894)</bibRefCitation>
mentioned
<taxonomicName id="4C161ACDFFB8FF8816BE88F5BA2FFD92" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[1012,1122,500,524]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFB8FF8816BE88F5BA2FFD92" box="[1012,1122,500,524]" italics="true" pageId="35" pageNumber="805">A. gregorii</emphasis>
</taxonomicName>
as ground living and described the habitat as a grassy coast-district with salt-swamps. In the Serengeti specimens were observed in rocky areas and on the ground hiding, even in skulls of large mammals, but also on acacia trees (PW, own observation). According to these observations,
<taxonomicName id="4C161ACDFFB8FF8814738B7FBFEBFD08" baseAuthorityName="Gunther" baseAuthorityYear="1894" box="[313,422,638,662]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="gregorii">
<emphasis id="B962BD5CFFB8FF8814738B7FBFEBFD08" box="[313,422,638,662]" italics="true" pageId="35" pageNumber="805">A. gregorii</emphasis>
</taxonomicName>
seems not to be a strictly tree-dwelling species, in contrast to e.g.
<taxonomicName id="4C161ACDFFB8FF8815EA8BA0BF5EFD27" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[160,275,673,697]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="805" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFFB8FF8815EA8BA0BF5EFD27" box="[160,275,673,697]" italics="true" pageId="35" pageNumber="805">A. atricollis</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF871CBFFFB8FF88146A8BA0BF82FD27" author="Sternfeld R" box="[288,463,673,697]" pageId="35" pageNumber="805" refId="ref21426" refString="Sternfeld R. 1912. Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika Expedition 1907 - 1908, unter Fuhrung A. Friedrichs, Herzogs zu Mecklenburg. Klinkhard und Biermann, Leipzig. IV. Zoologie II. Lfg. Reptilia; p. 197 - 279 [the volume was published in parts from 1911 to 1913 but this part was published in 1912]." type="book" year="1912">Sternfeld (1912)</bibRefCitation>
mentioned gravid females in December from Uvira Kasongo.
</paragraph>
</subSubSection>
</treatment>
</document>