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156 lines
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<document id="A8F0A29BB7CE6FFF710BC951DDC11BAE" ID-CLB-Dataset="95147" ID-DOI="10.11646/zootaxa.5139.1.1" ID-GBIF-Dataset="8ab25605-ec09-4d57-9dbd-7da2b032e5e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="6572023" ID-ZooBank="145CB6F5-2EA8-40B0-9CCA-3E942AA4A5B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653291328780" checkinUser="plazi" docAuthor="Just, Jean" docDate="2022" docId="03BF0417FFC4FF04FF1A8D10E4AAFC49" docLanguage="en" docName="zootaxa.5139.1.1.pdf" docOrigin="Zootaxa 5139 (1)" docSource="http://zoobank.org/145cb6f5-2ea8-40b0-9cca-3e942aa4a5b9" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tironella pervicax Just 2022, comb. nov." docType="treatment" docVersion="6" lastPageNumber="22" masterDocId="FF867C6FFFD1FF11FF8D8F32E74AFFA1" masterDocTitle="Tirons of the world: a review of ‘ tironid’ amphipods, description of new genera and species, and establishment of a new subfamily Tironinae Stebbing, 1906 stat. nov. (Crustacea, Synopiidae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="22" updateTime="1699337963343" updateUser="ExternalLinkService">
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<mods:title id="504EAB143E4414EEF6A2C14F006CCF67">Tirons of the world: a review of ‘ tironid’ amphipods, description of new genera and species, and establishment of a new subfamily Tironinae Stebbing, 1906 stat. nov. (Crustacea, Synopiidae)</mods:title>
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<mods:namePart id="9020E71F753BCD73480EC3193A1D4E50">Just, Jean</mods:namePart>
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<mods:title id="EE43D82BDF885D69A9CA82CA1A7DCEA0">Zootaxa</mods:title>
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<mods:date id="E3A4D97BDFC567A766035EA8670118B9">2022</mods:date>
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<mods:number id="A6A3B664AB1EE638743997A2454DBD66">2022-05-23</mods:number>
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<mods:identifier id="6D198D896F276DAEA337EC384E17EA57" type="DOI">10.11646/zootaxa.5139.1.1</mods:identifier>
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<treatment id="03BF0417FFC4FF04FF1A8D10E4AAFC49" ID-DOI="http://doi.org/10.5281/zenodo.6572109" ID-GBIF-Taxon="197509217" ID-Zenodo-Dep="6572109" LSID="urn:lsid:plazi:treatment:03BF0417FFC4FF04FF1A8D10E4AAFC49" httpUri="http://treatment.plazi.org/id/03BF0417FFC4FF04FF1A8D10E4AAFC49" lastPageNumber="22" pageId="21" pageNumber="22">
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<subSubSection id="C30CE68AFFC4FF04FF1A8D10E440FD9C" box="[151,778,546,573]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8BA9B501FFC4FF04FF1A8D10E440FD9C" blockId="21.[151,778,546,610]" box="[151,778,546,573]" pageId="21" pageNumber="22">
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<heading id="D0E1026DFFC4FF04FF1A8D10E440FD9C" bold="true" box="[151,778,546,573]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
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<emphasis id="B9626913FFC4FF04FF1A8D10E440FD9C" bold="true" box="[151,778,546,573]" pageId="21" pageNumber="22">
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<taxonomicName id="4C16CE82FFC4FF04FF1A8D10E536FD9D" ID-CoL="bd24ce36-6427-48df-95ca-0e36ce2ececf" authority="(J. L. Barnard, 1967)" authorityName="Just" authorityYear="2022" baseAuthorityName="J. L. Barnard" baseAuthorityYear="1967" box="[151,636,546,573]" class="Malacostraca" family="Synopiidae" genus="Tironella" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pervicax" status="comb. nov.">
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<emphasis id="B9626913FFC4FF04FF1A8D10E63FFD9C" bold="true" box="[151,373,546,573]" italics="true" pageId="21" pageNumber="22">Tironella pervicax</emphasis>
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(J.L.
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<bibRefCitation id="EF87C8F0FFC4FF04FE338D11E53EFD9D" author="Barnard, J. L." box="[446,628,546,573]" pageId="21" pageNumber="22" pagination="1 - 205" refId="ref48389" refString="Barnard, J. L. (1967) Bathyal and Abyssal Gammaridean Amphipoda of Cedros Trench, Baja California. United States National Museum Bulletin, 260, 1 - 205. https: // doi. org / 10.5479 / si. 03629236.260.1" type="journal article" year="1967">Barnard, 1967</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A251D468FFC4FF04FD098D11E440FD9C" box="[644,778,547,573]" pageId="21" pageNumber="22" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30CE68AFFC4FF04FF1A8D75E6D6FDC3" box="[151,412,583,610]" pageId="21" pageNumber="22" type="description">
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<paragraph id="8BA9B501FFC4FF04FF1A8D75E6D6FDC3" blockId="21.[151,778,546,610]" box="[151,412,583,610]" pageId="21" pageNumber="22">
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(
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<figureCitation id="132DA984FFC4FF04FF128D75E7ACFDC0" box="[159,230,583,609]" captionStart="FIGURE 1" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[160,1423,179,1796]" captionTargetId="figure-15@9.[151,1436,179,1806]" captionTargetPageId="9" captionText="FIGURE 1. Representative habitus illustrations of tironin genera. Tiron spiniferus (from Sars, 1895 pl. 140, ♀ 8 mm); Pseudotiron miratus sp. nov. (this paper, ♀, 4.5 mm); Tironella pervicax comb. nov. (from J.L. Barnard, 1967 fig. 84, ♂ 4.7 mm); Glandulotiron pilocaputis sp. nov. (this paper, ♀, 6.3 mm); Metatiron tropakis (from J.L. Barnard, 1967 fig. 45, ♀, 4.9 mm); Minitiron bellairsi comb. nov. (from Just, 1981 fig. 1, ♀ 2.4 mm). a and b, dactylus and unguis of pereopods 3–7: a, stubby; b, pre-stubby." figureDoi="http://doi.org/10.5281/zenodo.6572025" httpUri="https://zenodo.org/record/6572025/files/figure.png" pageId="21" pageNumber="22">Figs 1</figureCitation>
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,
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<figureCitation id="132DA984FFC4FF04FF798D75E648FDC0" box="[244,258,583,609]" captionStart="FIGURE 8" captionStartId="19.[151,250,1857,1881]" captionTargetBox="[151,1431,189,1817]" captionTargetId="figure-15@19.[151,1436,181,1833]" captionTargetPageId="19" captionText="FIGURE 8. Tironella bathyalis gen. et sp. nov. Holotype, ♀, except P, Pseudotiron pervicax. clv, head lateral view; mx1, maxilla 1; P clv, lateral view of head of P. pervicax (from J.L. Barnard, 1967, fig. 84 k); tel, telson; us, urosome." figureDoi="http://doi.org/10.5281/zenodo.6572039" httpUri="https://zenodo.org/record/6572039/files/figure.png" pageId="21" pageNumber="22">8</figureCitation>
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Pclv, 9 Pdac)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30CE68AFFC4FF04FF1A8DB9E35FFD02" box="[151,1045,651,675]" pageId="21" pageNumber="22" type="reference_group">
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<paragraph id="8BA9B501FFC4FF04FF1A8DB9E35FFD02" blockId="21.[151,1045,651,675]" box="[151,1045,651,675]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="AB06922FFFC4FF04FF1A8DB9E35FFD02" box="[151,1045,651,675]" pageId="21" pageNumber="22">
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<taxonomicName id="4C16CE82FFC4FF04FF1A8DB9E57CFD02" ID-CoL="78HXX" authority="J. L. Barnard, 1967" authorityName="J. L. Barnard" authorityYear="1967" box="[151,566,651,675]" class="Malacostraca" family="Synopiidae" genus="Pseudotiron" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pervicax">
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<emphasis id="B9626913FFC4FF04FF1A8DB9E625FD02" box="[151,367,651,675]" italics="true" pageId="21" pageNumber="22">Pseudotiron pervicax</emphasis>
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J.L.
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<bibRefCitation id="EF87C8F0FFC4FF04FE2F8DB9E57CFD02" author="Barnard, J. L." box="[418,566,651,675]" pageId="21" pageNumber="22" pagination="1 - 205" refId="ref48389" refString="Barnard, J. L. (1967) Bathyal and Abyssal Gammaridean Amphipoda of Cedros Trench, Baja California. United States National Museum Bulletin, 260, 1 - 205. https: // doi. org / 10.5479 / si. 03629236.260.1" type="journal article" year="1967">Barnard, 1967</bibRefCitation>
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</taxonomicName>
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, 170–172, fig. 84.—J.L.
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<treatmentCitation id="0AB79310FFC4FF04FCBE8DB9E343FD02" author="Barnard, J. L." box="[819,1033,651,675]" page="46" pageId="21" pageNumber="22" year="1972">
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<bibRefCitation id="EF87C8F0FFC4FF04FCBE8DB9E483FD02" author="Barnard, J. L." box="[819,969,651,675]" pageId="21" pageNumber="22" pagination="1 - 94" refId="ref48434" refString="Barnard, J. L. (1972) A Review of the Family Synopiidae (= Tironidae), Mainly Distributed in the Deep Sea (Crustacea: Amphipoda). Smithsonian Contributions to Zoology, 124 (1 - 4), 1 - 94. https: // doi. org / 10.5479 / si. 00810282.124" type="journal article" year="1972">Barnard (1972</bibRefCitation>
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, p. 46
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</treatmentCitation>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30CE68AFFC4FF04FF1A8DE3E33DFD4D" box="[151,1143,721,748]" pageId="21" pageNumber="22" type="materials_examined">
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<paragraph id="8BA9B501FFC4FF04FF1A8DE3E33DFD4D" blockId="21.[151,1436,721,1000]" box="[151,1143,721,748]" pageId="21" pageNumber="22">
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<emphasis id="B9626913FFC4FF04FF1A8DE3E676FD4A" bold="true" box="[151,316,721,747]" pageId="21" pageNumber="22">Type fixation.</emphasis>
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<materialsCitation id="3B7EBF5CFFC4FF04FECE8DE3E33DFD4D" ID-GBIF-Occurrence="3862594407" box="[323,1143,721,748]" collectionCode="AHF" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54AD0BA3FFC4FF04FECE8DE3E6E4FD4A" box="[323,430,721,747]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="9D107E88FFC4FF04FE348DE0E69AFD4D" box="[441,464,722,748]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
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,
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<quantity id="4CEE18E4FFC4FF04FE518DE3E579FD4A" box="[476,563,721,747]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="21" pageNumber="22" unit="mm" value="4.7">4.7 mm</quantity>
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.—Original designation.
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<collectionCode id="ED072DC4FFC4FF04FCCB8DE0E434FD4D" box="[838,894,722,748]" country="USA" httpUri="http://grbio.org/cool/95c7-rrrz" name="Allan Hancock Foundation, University of Southern California" pageId="21" pageNumber="22">AHF</collectionCode>
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No. 6115. [Not seen].
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30CE68AFFC4FF04FF4A8DC7E558FC65" pageId="21" pageNumber="22" type="discussion">
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<paragraph id="8BA9B501FFC4FF04FF4A8DC7E7BAFCDA" blockId="21.[151,1436,721,1000]" pageId="21" pageNumber="22">
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<emphasis id="B9626913FFC4FF04FF4A8DC7E67EFCAE" bold="true" box="[199,308,757,783]" pageId="21" pageNumber="22">Remarks</emphasis>
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.
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<taxonomicName id="4C16CE82FFC4FF04FEC88DC7E57CFCAE" authorityName="J. L. Barnard" authorityYear="1967" box="[325,566,757,783]" class="Malacostraca" family="Synopiidae" genus="Pseudotiron" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pervicax">
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<emphasis id="B9626913FFC4FF04FEC88DC7E57CFCAE" box="[325,566,757,783]" italics="true" pageId="21" pageNumber="22">Pseudotiron pervicax</emphasis>
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</taxonomicName>
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was illustrated and described by J.L.
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<bibRefCitation id="EF87C8F0FFC4FF04FC638DC7E3D4FCB1" author="Barnard, J. L." box="[1006,1182,757,784]" pageId="21" pageNumber="22" pagination="1 - 205" refId="ref48389" refString="Barnard, J. L. (1967) Bathyal and Abyssal Gammaridean Amphipoda of Cedros Trench, Baja California. United States National Museum Bulletin, 260, 1 - 205. https: // doi. org / 10.5479 / si. 03629236.260.1" type="journal article" year="1967">Barnard (1967)</bibRefCitation>
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, who had some doubt about its placement in
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<taxonomicName id="4C16CE82FFC4FF04FE008C2BE55FFC92" authorityName="Chevreux" authorityYear="1895" box="[397,533,793,819]" class="Malacostraca" family="Synopiidae" genus="Pseudotiron" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B9626913FFC4FF04FE008C2BE55FFC92" box="[397,533,793,819]" italics="true" pageId="21" pageNumber="22">Pseudotiron</emphasis>
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</taxonomicName>
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. He outlined many differences from the genotype
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<taxonomicName id="4C16CE82FFC4FF04FBB48C2BE785FCF9" authority="Chevreux, 1895" authorityName="Chevreux" authorityYear="1895" class="Malacostraca" family="Synopiidae" genus="Pseudotiron" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bouvieri">
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<emphasis id="B9626913FFC4FF04FBB48C2BE269FC92" box="[1081,1315,793,819]" italics="true" pageId="21" pageNumber="22">Pseudotiron bouvieri</emphasis>
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<bibRefCitation id="EF87C8F0FFC4FF04FAA58C2BE785FCF9" author="Chevreux, E." pageId="21" pageNumber="22" pagination="165 - 170" refId="ref48758" refString="Chevreux, E. (1895) Sur un Amphipode, Pseudotiron bouvieri, nov. gen. et sp., de la Famille des Syrrhoidae nouvelle pour la Fauna Mediterranenne. Bulletin de la Societe Zoologique de France, 20, 165 - 170. https: // doi. org / 10.5962 / bhl. part. 29691" type="journal article" year="1895">Chevreux, 1895</bibRefCitation>
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</taxonomicName>
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and the more widely known
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<taxonomicName id="4C16CE82FFC4FF04FDAF8C0CE43BFCF6" authority="Pirlot, 1934" authorityName="Pirlot" authorityYear="1934" box="[546,881,829,855]" class="Malacostraca" family="Synopiidae" genus="Pseudotiron" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="longicaudatus">
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<emphasis id="B9626913FFC4FF04FDAF8C0CE595FCF6" box="[546,735,829,855]" italics="true" pageId="21" pageNumber="22">P. longicaudatus</emphasis>
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<bibRefCitation id="EF87C8F0FFC4FF04FD658C0FE43BFCF6" author="Pirlot, J. M." box="[744,881,829,855]" pageId="21" pageNumber="22" pagination="167 - 235" refId="ref50065" refString="Pirlot, J. M. (1934) Les Amphipodes de l'Expedition du Siboga. Les Amphipodes Gammarides II. Les Amphipodes de la Mer Profonde. 2. Hyperiopsidae, Pardaliscidae, Astyridae nov. fam., Tironidae, Calliopiidae, Paramphithoidae, Amathillopsidae nov. fam., Eusiridae, Gammaridae, Aoridae, Photidae, Ampithoidae, Jassidae (Deuxieme Partie). E. J. Brill, Leiden, Leiden, 167 - 235." type="book chapter" year="1934">Pirlot, 1934</bibRefCitation>
|
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</taxonomicName>
|
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, but chose not to create a separate genus for the species.
|
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</paragraph>
|
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|
<paragraph id="8BA9B501FFC4FF04FF4A8CB7E558FC65" blockId="21.[151,1436,721,1000]" pageId="21" pageNumber="22">
|
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|
|
<taxonomicName id="4C16CE82FFC4FF04FF4A8CB7E6F2FC3E" authorityName="J. L. Barnard" authorityYear="1967" box="[199,440,901,927]" class="Malacostraca" family="Synopiidae" genus="Pseudotiron" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pervicax">
|
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|
|
<emphasis id="B9626913FFC4FF04FF4A8CB7E6F2FC3E" box="[199,440,901,927]" italics="true" pageId="21" pageNumber="22">Pseudotiron pervicax</emphasis>
|
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|
</taxonomicName>
|
|||
|
|
, being very close to
|
|||
|
|
<taxonomicName id="4C16CE82FFC4FF04FD218CB7E4CBFC3E" authorityName="Just" authorityYear="2022" box="[684,897,901,927]" class="Malacostraca" family="Synopiidae" genus="Tironella" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bathyalis">
|
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|
|
<emphasis id="B9626913FFC4FF04FD218CB7E4CBFC3E" box="[684,897,901,927]" italics="true" pageId="21" pageNumber="22">Tironella bathyalis</emphasis>
|
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|
</taxonomicName>
|
|||
|
|
<emphasis id="B9626913FFC4FF04FC078CB7E37DFC3E" bold="true" box="[906,1079,901,927]" pageId="21" pageNumber="22">gen. et sp nov.</emphasis>
|
|||
|
|
described above, is here transferred to the new genus
|
|||
|
|
<taxonomicName id="4C16CE82FFC4FF04FE288C9BE541FC62" authorityName="Just" authorityYear="2022" box="[421,523,937,963]" class="Malacostraca" family="Synopiidae" genus="Tironella" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9626913FFC4FF04FE288C9BE541FC62" box="[421,523,937,963]" italics="true" pageId="21" pageNumber="22">Tironella</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30CE68AFFC4FF04FF4A8CFFE4AAFC49" box="[199,992,973,1000]" pageId="21" pageNumber="22" type="distribution">
|
|||
|
|
<paragraph id="8BA9B501FFC4FF04FF4A8CFFE4AAFC49" blockId="21.[151,1436,721,1000]" box="[199,992,973,1000]" pageId="21" pageNumber="22">
|
|||
|
|
<emphasis id="B9626913FFC4FF04FF4A8CFFE613FC46" bold="true" box="[199,345,973,999]" pageId="21" pageNumber="22">Distribution</emphasis>
|
|||
|
|
. West coast
|
|||
|
|
<collectingCountry id="F301F591FFC4FF04FE6B8CFFE556FC46" box="[486,540,973,999]" name="United States of America" pageId="21" pageNumber="22">USA</collectingCountry>
|
|||
|
|
south to Baja
|
|||
|
|
<collectingRegion id="49D27BE3FFC4FF04FD338CFFE478FC46" box="[702,818,973,999]" country="United States of America" name="California" pageId="21" pageNumber="22">California</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<quantity id="4CEE18E4FFC4FF04FCB38CFFE496FC49" box="[830,988,973,1000]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.205" metricValueMin="1.095" pageId="21" pageNumber="22" unit="m" value="1150.0" valueMax="1205.0" valueMin="1095.0">1095–1205 m</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|