treatments-xml/data/03/B0/87/03B087E55B5AFFD3FF79F649FF64FE62.xml

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<document id="2B8F88F525B99D474874437A666783A2" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B5AFFD3FF79F649FF64FE62" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena clarinis Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="65" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="64" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2742C130B462A603269859C90ABED97E">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="5DE806D1829A966293C3E9DA3018BDA3">Perkins, Philip D.</mods:namePart>
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<mods:title id="3808152F4C2B558311B2DED9A9C6579E">Zootaxa</mods:title>
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<mods:date id="FB313E78F2A33FD72DC1DFB9539E5972">2011</mods:date>
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<mods:number id="923C4F1ECA23B9B28EA7CC1FEFD053A1">2944</mods:number>
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<mods:identifier id="4838234C95AF81AEDE8BB9D665156B2D" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<treatment id="03B087E55B5AFFD3FF79F649FF64FE62" ID-DOI="http://doi.org/10.5281/zenodo.5291534" ID-Zenodo-Dep="5291534" LSID="urn:lsid:plazi:treatment:03B087E55B5AFFD3FF79F649FF64FE62" httpUri="http://treatment.plazi.org/id/03B087E55B5AFFD3FF79F649FF64FE62" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<subSubSection id="C30365785B5AFFACFF79F649FDA6FBEB" box="[151,528,1123,1150]" pageId="63" pageNumber="64" type="nomenclature">
<paragraph id="8BA636F35B5AFFACFF79F649FDA6FBEB" blockId="63.[151,528,1123,1182]" box="[151,528,1123,1150]" pageId="63" pageNumber="64">
<heading id="D0EE819F5B5AFFACFF79F649FDA6FBEB" bold="true" box="[151,528,1123,1150]" fontSize="11" level="1" pageId="63" pageNumber="64" reason="1">
<emphasis id="B96DEAE15B5AFFACFF79F649FDA6FBEB" bold="true" box="[151,528,1123,1150]" pageId="63" pageNumber="64">
<taxonomicName id="4C194D705B5AFFACFF79F649FEC7FBE8" ID-CoL="3N37M" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,369,1124,1150]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="clarinis" status="sp. nov.">
<emphasis id="B96DEAE15B5AFFACFF79F649FEC7FBE8" bold="true" box="[151,369,1124,1150]" italics="true" pageId="63" pageNumber="64">Hydraena clarinis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B5AFFACFE6EF64EFDA6FBEB" box="[384,528,1123,1149]" pageId="63" pageNumber="64" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5AFFACFF79F6ABFEC2FB08" box="[151,372,1158,1182]" pageId="63" pageNumber="64" type="description">
<paragraph id="8BA636F35B5AFFACFF79F6ABFEC2FB08" blockId="63.[151,528,1123,1182]" box="[151,372,1158,1182]" pageId="63" pageNumber="64">
(
<figureCitation id="13222A765B5AFFACFF71F6ABFF4BFB08" box="[159,253,1158,1182]" captionStart="FIGURE 99" captionStartId="219.[151,250,1612,1635]" captionTargetBox="[210,1376,215,1581]" captionTargetId="figure-26@219.[186,1401,193,1591]" captionTargetPageId="219" captionText="FIGURE 99. Hydraena clarinis, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283607" httpUri="https://zenodo.org/record/5283607/files/figure.png" pageId="63" pageNumber="64">Figs. 99</figureCitation>
,
<figureCitation id="13222A765B5AFFACFEE6F6ABFE82FB08" box="[264,308,1158,1182]" captionStart="FIGURES 100101" captionStartId="220.[151,264,1650,1673]" captionTargetBox="[309,1276,224,1579]" captionTargetId="figure-32@220.[264,1323,193,1629]" captionTargetPageId="220" captionText="FIGURES 100101. Aedeagi of holotypes. 100. H. nanocolorata. 101. H. clarinis." figureDoi="http://doi.org/10.5281/zenodo.5283609" httpUri="https://zenodo.org/record/5283609/files/figure.png" pageId="63" pageNumber="64">101</figureCitation>
,
<figureCitation id="13222A765B5AFFACFEAEF6ABFEDDFB08" box="[320,363,1158,1182]" captionStart="FIGURES 475478" captionStartId="394.[151,264,1650,1673]" captionTargetBox="[165,1421,259,1607]" captionTargetId="figure-23@394.[165,1421,193,1629]" captionTargetPageId="394" captionText="FIGURES 475478. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284093" httpUri="https://zenodo.org/record/5284093/files/figure.png" pageId="63" pageNumber="64">476</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5AFFACFF79F6E2FDA6FAC4" pageId="63" pageNumber="64" type="materials_examined">
<paragraph id="8BA636F35B5AFFACFF79F6E2FDA6FAC4" blockId="63.[151,1436,1230,2010]" pageId="63" pageNumber="64">
Type Material.
<materialsCitation id="3B713CAE5B5AFFACFEBEF6E2FAA4FAB8" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Koje Creek" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B5AFFACFEBEF6E2FE0BFB71" box="[336,445,1231,1255]" pageId="63" pageNumber="64" type="holotype">
<emphasis id="B96DEAE15B5AFFACFEBEF6E2FE0BFB71" bold="true" box="[336,445,1231,1255]" pageId="63" pageNumber="64">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B5AFFACFDCBF6E2FCB5FB71" bold="true" box="[549,771,1230,1255]" pageId="63" pageNumber="64">
<collectingRegion id="49DDF8115B5AFFACFDCBF6E2FD48FB71" box="[549,766,1230,1255]" country="Papua New Guinea" name="Madang" pageId="63" pageNumber="64">Madang Province</collectingRegion>
:
</emphasis>
<collectingRegion id="49DDF8115B5AFFACFCFAF6E3FCE5FB70" box="[788,851,1230,1254]" country="Papua New Guinea" name="West Sepik" pageId="63" pageNumber="64">Sepik</collectingRegion>
<collectingMunicipality id="6BC2AC895B5AFFACFCB5F6E2FC5FFB70" box="[859,1001,1230,1254]" pageId="63" pageNumber="64">Ramu Basin</collectingMunicipality>
,
<location id="8EC660285B5AFFACFC1BF6E3FBCFFB70" LSID="urn:lsid:plazi:treatment:03B087E55B5AFFD3FF79F649FF64FE62:8EC660285B5AFFACFC1BF6E3FBCFFB70" box="[1013,1145,1230,1254]" latitude="-5.55" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" name="Koje Creek" pageId="63" pageNumber="64" stateProvince="Madang">Kojé Creek</location>
, unshaded creek, small to medium cobbles, gravel and fine sand (the latter in areas of slack water), filamentous algae on stony substrates, small accumulations of leaf litter,
<quantity id="4CE19B165B5AFFACFDFDF73BFDEAFAB8" box="[531,604,1302,1326]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="63" pageNumber="64" unit="m" value="160.0">
<elevation id="0034D1C05B5AFFACFDFDF73BFDEAFAB8" box="[531,604,1302,1326]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="63" pageNumber="64" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B5AFFACFD86F73BFD77FAB8" box="[616,705,1302,1326]" degrees="5" direction="south" minutes="33" orientation="latitude" pageId="63" pageNumber="64" precision="925" value="-5.55">5° 33' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B5AFFACFD23F73BFCF2FAB8" box="[717,836,1302,1326]" degrees="145" direction="east" minutes="23" orientation="longitude" pageId="63" pageNumber="64" precision="925" value="145.38333">145° 23' E</geoCoordinate>
,
<date id="FFA710335B5AFFACFCBEF73BFBAFFAB8" box="[848,1049,1302,1326]" pageId="63" pageNumber="64" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">
<collectingDate id="EFE3E9DB5B5AFFACFCBEF73BFBAFFAB8" box="[848,1049,1302,1326]" pageId="63" pageNumber="64" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">1 vii30 viii 1988</collectingDate>
</date>
,
<collectorName id="26EC53255B5AFFACFBCAF73AFB1BFAB8" box="[1060,1197,1302,1326]" pageId="63" pageNumber="64">D. Dudgeon</collectorName>
(
<collectionCode id="ED08AE365B5AFFACFB52F73BFABCFAB8" box="[1212,1290,1302,1326]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="63" pageNumber="64">NMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B5AFFACFAF2F73AFDBAFAC4" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Koje Creek" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B5AFFACFAF2F73AFA22FAB9" bold="true" box="[1308,1428,1303,1327]" pageId="63" pageNumber="64">
<typeStatus id="54A288515B5AFFACFAF2F73AFA22FAB9" box="[1308,1428,1303,1327]" pageId="63" pageNumber="64" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as holotype (8
<collectionCode id="ED08AE365B5AFFACFE59F717FDB2FAC4" box="[439,516,1338,1362]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="63" pageNumber="64">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5AFFACFF29F773FF67F990" pageId="63" pageNumber="64" type="diagnosis">
<paragraph id="8BA636F35B5AFFACFF29F773FF67F990" blockId="63.[151,1436,1230,2010]" pageId="63" pageNumber="64">
<emphasis id="B96DEAE15B5AFFACFF29F773FE65FAE1" bold="true" box="[199,467,1374,1399]" pageId="63" pageNumber="64">Differential Diagnosis.</emphasis>
Differentiated from other members of the
<emphasis id="B96DEAE15B5AFFACFC54F773FBA9FAE0" box="[954,1055,1374,1398]" italics="true" pageId="63" pageNumber="64">Colorata</emphasis>
group by the small size (ca.
<quantity id="4CE19B165B5AFFACFA85F773FF72FA0C" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="63" pageNumber="64" unit="mm" value="1.13">1.13 mm</quantity>
) and the clear pronotum, which does not have a macula or fascia (
<figureCitation id="13222A765B5AFFACFC42F7AFFBB6FA0C" box="[940,1024,1410,1434]" captionStart="FIGURE 99" captionStartId="219.[151,250,1612,1635]" captionTargetBox="[210,1376,215,1581]" captionTargetId="figure-26@219.[186,1401,193,1591]" captionTargetPageId="219" captionText="FIGURE 99. Hydraena clarinis, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283607" httpUri="https://zenodo.org/record/5283607/files/figure.png" pageId="63" pageNumber="64">Fig. 99</figureCitation>
). The testaceous pronotum is similar in color to that of
<taxonomicName id="4C194D705B5AFFACFE8BF78AFE4AFA28" authorityName="Perkins" authorityYear="2011" box="[357,508,1447,1470]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="ramuensis">
<emphasis id="B96DEAE15B5AFFACFE8BF78AFECFFA28" box="[357,377,1447,1470]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B96DEAE15B5AFFACFE67F78AFE4AFA28" box="[393,508,1447,1470]" italics="true" pageId="63" pageNumber="64">ramuensis</emphasis>
</taxonomicName>
(which see), but otherwise the two species differ distinctively. The male genitalia bear some superficial similarity to that of
<taxonomicName id="4C194D705B5AFFACFD92F7E6FD49FA75" authorityName="Perkins" authorityYear="2011" box="[636,767,1483,1507]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="colorata">
<emphasis id="B96DEAE15B5AFFACFD92F7E6FD26FA74" box="[636,656,1483,1506]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B96DEAE15B5AFFACFD4EF7E6FD49FA75" box="[672,767,1483,1507]" italics="true" pageId="63" pageNumber="64">colorata</emphasis>
</taxonomicName>
, but the two distinctively differ in many details (
<figureCitation id="13222A765B5AFFACFAD9F7E7FA21FA74" box="[1335,1431,1482,1506]" captionStart="FIGURES 8889" captionStartId="211.[151,264,1561,1584]" captionTargetBox="[249,1332,217,1516]" captionTargetId="figure-29@211.[226,1360,193,1540]" captionTargetPageId="211" captionText="FIGURES 8889. Aedeagi of holotypes. 88. H. koma. 89. H. colorata." figureDoi="http://doi.org/10.5281/zenodo.5283585" httpUri="https://zenodo.org/record/5283585/files/figure.png" pageId="63" pageNumber="64">Figs. 89</figureCitation>
,
<figureCitation id="13222A765B5AFFACFF79F7C3FF72F990" box="[151,196,1518,1542]" captionStart="FIGURES 100101" captionStartId="220.[151,264,1650,1673]" captionTargetBox="[309,1276,224,1579]" captionTargetId="figure-32@220.[264,1323,193,1629]" captionTargetPageId="220" captionText="FIGURES 100101. Aedeagi of holotypes. 100. H. nanocolorata. 101. H. clarinis." figureDoi="http://doi.org/10.5281/zenodo.5283609" httpUri="https://zenodo.org/record/5283609/files/figure.png" pageId="63" pageNumber="64">101</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5AFFD3FF29F43FFB41FE11" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" type="description">
<paragraph id="8BA636F35B5AFFACFF29F43FFB4CF9E4" blockId="63.[151,1436,1230,2010]" pageId="63" pageNumber="64">
<emphasis id="B96DEAE15B5AFFACFF29F43FFEEEF9BD" bold="true" box="[199,344,1554,1579]" inLineHeading="true" pageId="63" pageNumber="64" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B5AFFACFE42F43FFDB8F9BC" box="[428,526,1554,1578]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.13/0.50; head 0.18/0.28; pronotum 0.29/0.36, PA 0.31, PB 0.32; elytra 0.67/0.50. Dorsum of head dark brown; pronotum testaceous to light brown, no macula or fascia; elytra brown; legs light brown to testaceous; maxillary palpi testaceous.
</paragraph>
<paragraph id="8BA636F35B5AFFACFF29F452FDA1F8DC" blockId="63.[151,1436,1230,2010]" pageId="63" pageNumber="64">Frons punctures ca. 1xef; interstices shining, 25xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely and very finely punctulate, strongly shining. Genae very slightly raised, shining, without posterior ridge. Pronotum weakly cordiform, anterior margin slightly emarginate; punctures on disc ca. 1xpd those of frons, interstices shining, 13xpd, punctures slightly larger and denser at anterior and posterior; PF1 very shallow, almost obsolete; PF2 very shallow, oval, oblique, very shallowly confluent; PF3 moderately deep; PF4 absent.</paragraph>
<paragraph id="8BA636F35B5AFFACFF29F57BFB7DF84C" blockId="63.[151,1436,1230,2010]" pageId="63" pageNumber="64">Elytra with summit of posterior declivity before midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 13xpd, as are interstices between punctures of a row. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B25FFD3FF29F2BAFB41FE11" blockId="64.[151,1436,151,500]" pageId="64" pageNumber="65">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/1/3/3. P1 laminate; median carina sinuate in profile. P2 very slightly raised, l/w ca. 2/1, sides slightly converging toward apex. Plaques short, carinate, located at sides of deep, inverted V-shaped median depression. Metaventrite with very low midlongitudinal ridge between P2 and apex of median depression. AIS slightly concave, width at straight posterior margin ca. 1.5x P2. Profemur (male) with small, pointed tubercle next to trochanter; protibia slightly arcuate, medial margin in distal 1/2 slightly thickened and bearing row of very short, peg-like spines. Mesotibia straight. Metatibia very slightly, if at all, arcuate. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
<figureCitation id="13222A765B25FFD3FB66F342FB5FFE11" box="[1160,1257,367,391]" captionStart="FIGURES 100101" captionStartId="220.[151,264,1650,1673]" captionTargetBox="[309,1276,224,1579]" captionTargetId="figure-32@220.[264,1323,193,1629]" captionTargetPageId="220" captionText="FIGURES 100101. Aedeagi of holotypes. 100. H. nanocolorata. 101. H. clarinis." figureDoi="http://doi.org/10.5281/zenodo.5283609" httpUri="https://zenodo.org/record/5283609/files/figure.png" pageId="64" pageNumber="65">Fig. 101</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B25FFD3FF29F3B9FC1FFE3D" box="[199,937,404,428]" pageId="64" pageNumber="65" type="etymology">
<paragraph id="8BA636F35B25FFD3FF29F3B9FC1FFE3D" blockId="64.[151,1436,151,500]" box="[199,937,404,428]" pageId="64" pageNumber="65">
<emphasis id="B96DEAE15B25FFD3FF29F3B9FEFBFE3A" bold="true" box="[199,333,404,428]" inLineHeading="true" pageId="64" pageNumber="65" reason="1">Etymology.</emphasis>
Named in reference to the clear and bright pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B25FFD3FF29F39AFF64FE62" pageId="64" pageNumber="65" type="distribution">
<paragraph id="8BA636F35B25FFD3FF29F39AFF64FE62" blockId="64.[151,1436,151,500]" pageId="64" pageNumber="65">
<emphasis id="B96DEAE15B25FFD3FF29F39AFED6FE46" bold="true" box="[199,352,439,464]" inLineHeading="true" pageId="64" pageNumber="65" reason="1">Distribution.</emphasis>
Currently known only from
<typeStatus id="54A288515B25FFD3FD42F395FD6AFE46" box="[684,732,440,464]" pageId="64" pageNumber="65">type</typeStatus>
locality, Kojé Creek in eastern Area 3; elevation
<quantity id="4CE19B165B25FFD3FAFBF39AFAE8FE59" box="[1301,1374,439,463]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="64" pageNumber="65" unit="m" value="160.0">160 m</quantity>
(
<figureCitation id="13222A765B25FFD3FA81F39AFF70FE62" captionStart="FIGURES 475478" captionStartId="394.[151,264,1650,1673]" captionTargetBox="[165,1421,259,1607]" captionTargetId="figure-23@394.[165,1421,193,1629]" captionTargetPageId="394" captionText="FIGURES 475478. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284093" httpUri="https://zenodo.org/record/5284093/files/figure.png" pageId="64" pageNumber="65">Fig. 476</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>