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<document id="5CB43FFD55A011F5D2F5F7F8634AE77B" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B4CFFB9FF79F73AFCF1FCCA" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena infoveola Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="43" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="42" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="06089D8613F6BDE5053084F439098AAF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8E04863C6D6448BF3FC863D0B9CB8A5F">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
<mods:name id="2402C5CCF377FAC7DB5C5C3B106E9333" type="personal">
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<mods:namePart id="713961350537B6AF470C03548FC10FD5">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1579E5FBFB35BF5026F43DE4E435B21B">text</mods:typeOfResource>
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<mods:title id="EE3E6677B5BB446CAD55C14E0EF12EBD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9B562AA959B3CDC0EE2D03795A27458B">
<mods:date id="1637188A662507451198F99A1AEF2199">2011</mods:date>
<mods:detail id="2C400FFD240D735ACFBF154D067026C8" type="pubDate">
<mods:number id="B4DBC56D07E456565C33FACF45AAB909">2011-07-08</mods:number>
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<mods:number id="44C0DFC06D0AC59CCE41CC0522BBEE2D">2944</mods:number>
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<mods:number id="842F9E67C4756B280524DFFFF6ED5820">1</mods:number>
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<mods:url id="AB91128216F01D94244830DC78FF9D77">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
</mods:location>
<mods:classification id="89FFDCB7DBC0060966BE3127F16DBFC1">journal article</mods:classification>
<mods:identifier id="D3BAE082BBE3FDE6461EB079190506B9" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="CAFA200D680AA51D474EEE36DB835B48" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="5EACE2DC46805E73F53A15BBD86E91A1" type="Zenodo-Dep">5283420</mods:identifier>
</mods:mods>
<treatment id="03B087E55B4CFFB9FF79F73AFCF1FCCA" ID-DOI="http://doi.org/10.5281/zenodo.5291431" ID-Zenodo-Dep="5291431" LSID="urn:lsid:plazi:treatment:03B087E55B4CFFB9FF79F73AFCF1FCCA" httpUri="http://treatment.plazi.org/id/03B087E55B4CFFB9FF79F73AFCF1FCCA" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<subSubSection id="C30365785B4CFFBAFF79F73AFD93FAA7" box="[151,549,1303,1329]" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="8BA636F35B4CFFBAFF79F73AFD93FAA7" blockId="41.[151,549,1303,1362]" box="[151,549,1303,1329]" pageId="41" pageNumber="42">
<heading id="D0EE819F5B4CFFBAFF79F73AFD93FAA7" bold="true" box="[151,549,1303,1329]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
<emphasis id="B96DEAE15B4CFFBAFF79F73AFD93FAA7" bold="true" box="[151,549,1303,1329]" pageId="41" pageNumber="42">
<taxonomicName id="4C194D705B4CFFBAFF79F73AFE30FAA7" ID-CoL="3N3BJ" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,390,1303,1329]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="infoveola" status="sp. nov.">
<emphasis id="B96DEAE15B4CFFBAFF79F73AFE30FAA7" bold="true" box="[151,390,1303,1329]" italics="true" pageId="41" pageNumber="42">Hydraena infoveola</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B4CFFBAFE7BF73AFD93FAA7" box="[405,549,1303,1329]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4CFFBAFF79F717FED0FAC4" box="[151,358,1338,1362]" pageId="41" pageNumber="42" type="description">
<paragraph id="8BA636F35B4CFFBAFF79F717FED0FAC4" blockId="41.[151,549,1303,1362]" box="[151,358,1338,1362]" pageId="41" pageNumber="42">
(
<figureCitation id="13222A765B4CFFBAFF71F717FF4BFAC4" box="[159,253,1338,1362]" captionStart="FIGURE 54" captionStartId="185.[151,250,1753,1776]" captionTargetBox="[216,1372,209,1718]" captionTargetId="figure-25@185.[198,1388,193,1732]" captionTargetPageId="185" captionText="FIGURE 54. Hydraena infoveola, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283500" httpUri="https://zenodo.org/record/5283500/files/figure.png" pageId="41" pageNumber="42">Figs. 54</figureCitation>
,
<figureCitation id="13222A765B4CFFBAFEE6F717FE90FAC4" box="[264,294,1338,1362]" captionStart="FIGURES 56–57" captionStartId="187.[151,264,1650,1673]" captionTargetBox="[282,1301,215,1623]" captionTargetId="figure-29@187.[263,1323,193,1629]" captionTargetPageId="187" captionText="FIGURES 56–57. Aedeagi of holotypes. 56. H. infoveola. 57. H. otiarca." figureDoi="http://doi.org/10.5281/zenodo.5283506" httpUri="https://zenodo.org/record/5283506/files/figure.png" pageId="41" pageNumber="42">56</figureCitation>
,
<figureCitation id="13222A765B4CFFBAFEDCF717FEE8FAC4" box="[306,350,1338,1362]" captionStart="FIGURES 435–438" captionStartId="384.[151,264,1650,1673]" captionTargetBox="[165,1421,235,1613]" captionTargetId="figure-23@384.[165,1421,193,1629]" captionTargetPageId="384" captionText="FIGURES 435–438. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284069" httpUri="https://zenodo.org/record/5284069/files/figure.png" pageId="41" pageNumber="42">438</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4CFFBAFF79F7AEFE79F990" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8BA636F35B4CFFBAFF79F7AEFE79F990" blockId="41.[151,1436,1410,2011]" pageId="41" pageNumber="42">
Type Material.
<materialsCitation id="3B713CAE5B4CFFBAFEA0F7AEFD3CFA28" collectingDate="2006-11-11" collectionCode="ZSM" collectorName="Balke &amp; Kinibel" country="Papua New Guinea" elevation="1400" latitude="-7.0944" location="Marawaka" longLatPrecision="1" longitude="145.74112" municipality="Mala" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Gulf" typeStatus="holotype">
<typeStatus id="54A288515B4CFFBAFEA0F7AEFE0DFA0D" box="[334,443,1411,1435]" pageId="41" pageNumber="42" type="holotype">
<emphasis id="B96DEAE15B4CFFBAFEA0F7AEFE0DFA0D" bold="true" box="[334,443,1411,1435]" pageId="41" pageNumber="42">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B4CFFBAFDCEF7AFFD66FA0D" bold="true" box="[544,720,1410,1435]" pageId="41" pageNumber="42">
<collectingRegion id="49DDF8115B4CFFBAFDCEF7AFFD7CFA0D" box="[544,714,1410,1435]" country="Papua New Guinea" name="Gulf" pageId="41" pageNumber="42">Gulf Province</collectingRegion>
:
</emphasis>
<location id="8EC660285B4CFFBAFD33F7AFFCE0FA0C" LSID="urn:lsid:plazi:treatment:03B087E55B4CFFB9FF79F73AFCF1FCCA:8EC660285B4CFFBAFD33F7AFFCE0FA0C" box="[733,854,1410,1434]" country="Papua New Guinea" latitude="-7.0944" longLatPrecision="1" longitude="145.74112" municipality="Mala" name="Marawaka" pageId="41" pageNumber="42" stateProvince="Gulf">Marawaka</location>
,
<collectingMunicipality id="6BC2AC895B4CFFBAFC8FF7AFFC2BFA0C" box="[865,925,1410,1434]" pageId="41" pageNumber="42">Mala</collectingMunicipality>
, elev.
<quantity id="4CE19B165B4CFFBAFC0DF7AFFB8CFA0C" box="[995,1082,1410,1434]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="m" value="1400.0">
<elevation id="0034D1C05B4CFFBAFC0DF7AFFB8CFA0C" box="[995,1082,1410,1434]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B4CFFBAFBABF7AFFB77FA0C" box="[1093,1217,1410,1434]" degrees="7" direction="south" minutes="5.664" orientation="latitude" pageId="41" pageNumber="42" precision="1" value="-7.0944">7° 5.664' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B4CFFBAFB23F7AFFAC3FA0C" box="[1229,1397,1410,1434]" degrees="145" direction="east" minutes="44.467" orientation="longitude" pageId="41" pageNumber="42" precision="1" value="145.74112">145° 44.467' E</geoCoordinate>
,
<date id="FFA710335B4CFFBAFA6FF7AFFF46FA28" pageId="41" pageNumber="42" value="2006-11-11">
<collectingDate id="EFE3E9DB5B4CFFBAFA6FF7AFFF46FA28" pageId="41" pageNumber="42" value="2006-11-11">11 xi 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B4CFFBAFF13F78BFE88FA28" box="[253,318,1446,1470]" pageId="41" pageNumber="42">Balke</collectorName>
&amp;
<collectorName id="26EC53255B4CFFBAFE8BF78BFE0FFA28" box="[357,441,1446,1470]" pageId="41" pageNumber="42">Kinibel</collectorName>
(
<collectingCountry id="F30E76635B4CFFBAFE24F78BFDB2FA28" box="[458,516,1446,1470]" name="Papua New Guinea" pageId="41" pageNumber="42">PNG</collectingCountry>
90) (
<collectionCode id="ED08AE365B4CFFBAFDACF78BFD37FA28" box="[578,641,1446,1470]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="41" pageNumber="42" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B4CFFBAFD78F78AFE7DF990" collectingDate="2006-11-10" collectionCode="MCZ, NHM, NMW, ZSM" collectorName="Balke &amp; Kinibel" country="Papua New Guinea" county="Ande" elevation="1000" latitude="-7.0599666" location="Same" longLatPrecision="1" longitude="145.73958" municipality="Marawaka" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Gulf" typeStatus="paratype">
<emphasis id="B96DEAE15B4CFFBAFD78F78AFCB8FA29" bold="true" box="[662,782,1447,1471]" pageId="41" pageNumber="42">
<typeStatus id="54A288515B4CFFBAFD78F78AFCB8FA29" box="[662,782,1447,1471]" pageId="41" pageNumber="42" type="paratype">Paratypes</typeStatus>
</emphasis>
(27):
<emphasis id="B96DEAE15B4CFFBAFCB0F78BFBA6FA29" bold="true" box="[862,1040,1446,1471]" pageId="41" pageNumber="42">
<collectingRegion id="49DDF8115B4CFFBAFCB0F78BFBBCFA29" box="[862,1034,1446,1471]" country="Papua New Guinea" name="Gulf" pageId="41" pageNumber="42">Gulf Province</collectingRegion>
:
</emphasis>
<location id="8EC660285B4CFFBAFBCFF78BFBE9FA28" LSID="urn:lsid:plazi:treatment:03B087E55B4CFFB9FF79F73AFCF1FCCA:8EC660285B4CFFBAFBCFF78BFBE9FA28" box="[1057,1119,1446,1470]" country="Papua New Guinea" county="Ande" latitude="-7.0599666" longLatPrecision="1" longitude="145.73958" municipality="Marawaka" name="Same" pageId="41" pageNumber="42" stateProvince="Gulf">Same</location>
data as holotype (25
<collectionCode id="ED08AE365B4CFFBAFAB6F78BFA21FA28" box="[1368,1431,1446,1470]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
,
<collectionCode id="ED08AE365B4CFFBAFF79F7E6FF6DFA74" box="[151,219,1483,1506]" pageId="41" pageNumber="42">NHM</collectionCode>
,
<collectionCode id="ED08AE365B4CFFBAFF09F7E6FE99FA74" box="[231,303,1483,1506]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
,
<collectingCountry id="F30E76635B4CFFBAFED4F7E7FEC4FA74" box="[314,370,1482,1506]" name="Papua New Guinea" pageId="41" pageNumber="42">PNG</collectingCountry>
,
<collectionCode id="ED08AE365B4CFFBAFE93F7E7FE0DFA74" box="[381,443,1482,1506]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="41" pageNumber="42" type="Museum">ZSM</collectionCode>
);
<collectingMunicipality id="6BC2AC895B4CFFBAFE21F7E7FDFEFA74" box="[463,584,1482,1506]" pageId="41" pageNumber="42">Marawaka</collectingMunicipality>
, near
<collectingCounty id="62C74E7F5B4CFFBAFD65F7E7FD7DFA74" box="[651,715,1482,1506]" pageId="41" pageNumber="42">Ande</collectingCounty>
, elev.
<quantity id="4CE19B165B4CFFBAFCFCF7E7FCDCFA74" box="[786,874,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="41" pageNumber="42" unit="m" value="1000.0">
<elevation id="0034D1C05B4CFFBAFCFCF7E7FCDCFA74" box="[786,874,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="41" pageNumber="42" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B4CFFBAFC98F7E7FC45FA74" box="[886,1011,1482,1506]" degrees="7" direction="south" minutes="3.598" orientation="latitude" pageId="41" pageNumber="42" precision="1" value="-7.0599666">7° 3.598' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B4CFFBAFBEEF7E7FB1CFA74" box="[1024,1194,1482,1506]" degrees="145" direction="east" minutes="44.375" orientation="longitude" pageId="41" pageNumber="42" precision="1" value="145.73958">145° 44.375' E</geoCoordinate>
,
<date id="FFA710335B4CFFBAFB58F7E7FA84FA74" box="[1206,1330,1482,1506]" pageId="41" pageNumber="42" value="2006-11-10">
<collectingDate id="EFE3E9DB5B4CFFBAFB58F7E7FA84FA74" box="[1206,1330,1482,1506]" pageId="41" pageNumber="42" value="2006-11-10">10 xi 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B4CFFBAFAD0F7E7FAC9FA74" box="[1342,1407,1482,1506]" pageId="41" pageNumber="42">Balke</collectorName>
&amp;
<collectorName id="26EC53255B4CFFBAFF79F7C3FF5DF990" box="[151,235,1518,1542]" pageId="41" pageNumber="42">Kinibel</collectorName>
(
<collectingCountry id="F30E76635B4CFFBAFF17F7C3FE85F990" box="[249,307,1518,1542]" name="Papua New Guinea" pageId="41" pageNumber="42">PNG</collectingCountry>
89) (2
<collectionCode id="ED08AE365B4CFFBAFE6BF7C3FE75F990" box="[389,451,1518,1542]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="41" pageNumber="42" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4CFFBAFF29F43FFE76F894" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="8BA636F35B4CFFBAFF29F43FFE76F894" blockId="41.[151,1436,1410,2011]" pageId="41" pageNumber="42">
<emphasis id="B96DEAE15B4CFFBAFF29F43FFE64F9BD" bold="true" box="[199,466,1554,1579]" pageId="41" pageNumber="42">Differential Diagnosis.</emphasis>
Among members of the
<emphasis id="B96DEAE15B4CFFBAFD05F43EFCE4F9BD" box="[747,850,1555,1579]" italics="true" pageId="41" pageNumber="42">Foliobba</emphasis>
group, most similar in dorsal habitus to
<taxonomicName id="4C194D705B4CFFBAFAF4F43EFA2AF9BD" authorityName="Perkins" authorityYear="2011" box="[1306,1436,1555,1579]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="striolata">
<emphasis id="B96DEAE15B4CFFBAFAF4F43EFA98F9BC" box="[1306,1326,1555,1578]" italics="true" pageId="41" pageNumber="42">H</emphasis>
.
<emphasis id="B96DEAE15B4CFFBAFAD3F43EFA2AF9BD" box="[1341,1436,1555,1579]" italics="true" pageId="41" pageNumber="42">striolata</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B4CFFBAFF71F41BFF4BF9D8" box="[159,253,1590,1614]" captionStart="FIGURE 51" captionStartId="183.[151,250,1753,1776]" captionTargetBox="[199,1395,209,1718]" captionTargetId="figure-27@183.[183,1404,193,1732]" captionTargetPageId="183" captionText="FIGURE 51. Hydraena striolata, holotype, dorsal and lateral habitus, venter, and abdominal apex." figureDoi="http://doi.org/10.5281/zenodo.5283494" httpUri="https://zenodo.org/record/5283494/files/figure.png" pageId="41" pageNumber="42">Figs. 51</figureCitation>
,
<figureCitation id="13222A765B4CFFBAFEE4F41BFE9CF9D8" box="[266,298,1590,1614]" captionStart="FIGURE 54" captionStartId="185.[151,250,1753,1776]" captionTargetBox="[216,1372,209,1718]" captionTargetId="figure-25@185.[198,1388,193,1732]" captionTargetPageId="185" captionText="FIGURE 54. Hydraena infoveola, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283500" httpUri="https://zenodo.org/record/5283500/files/figure.png" pageId="41" pageNumber="42">54</figureCitation>
); differing therefrom by the larger size (ca. 1.58 vs.
<quantity id="4CE19B165B4CFFBAFC92F41BFC53F9D8" box="[892,997,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="41" pageNumber="42" unit="mm" value="1.43">1.43 mm</quantity>
), the testaceous color bands in front of and behind the pronotal fascia (brown in
<taxonomicName id="4C194D705B4CFFBAFD83F476FD46F9E5" authorityName="Perkins" authorityYear="2011" box="[621,752,1627,1651]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="striolata">
<emphasis id="B96DEAE15B4CFFBAFD83F476FD37F9E4" box="[621,641,1627,1650]" italics="true" pageId="41" pageNumber="42">H</emphasis>
.
<emphasis id="B96DEAE15B4CFFBAFD7FF476FD46F9E5" box="[657,752,1627,1651]" italics="true" pageId="41" pageNumber="42">striolata</emphasis>
</taxonomicName>
), the proportionally larger pronotal fascia (color band ratios ca. 6/18/3 vs. 6/14/3), the absent, or nearly imperceptible pronotal foveae PF2, and the more widely separated metaventral plaques. Males of the two species are additionally differentiated by differences in the shapes of the tibiae, and by the striolate area on the fifth abdominal ventrite of
<taxonomicName id="4C194D705B4CFFBAFCB9F4EAFC6EF948" authorityName="Perkins" authorityYear="2011" box="[855,984,1734,1758]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="striolata">
<emphasis id="B96DEAE15B4CFFBAFCB9F4EAFCDDF948" box="[855,875,1735,1758]" italics="true" pageId="41" pageNumber="42">H</emphasis>
.
<emphasis id="B96DEAE15B4CFFBAFC97F4EBFC6EF948" box="[889,984,1734,1758]" italics="true" pageId="41" pageNumber="42">striolata</emphasis>
</taxonomicName>
. The aedeagi of the two species distinctively differ (
<figureCitation id="13222A765B4CFFBAFEC4F4C7FE3EF894" box="[298,392,1770,1794]" captionStart="FIGURES 52–53" captionStartId="184.[151,264,1613,1636]" captionTargetBox="[256,1343,211,1564]" captionTargetId="figure-32@184.[226,1360,193,1592]" captionTargetPageId="184" captionText="FIGURES 52–53. Aedeagi of holotypes. 52. H. foliobba. 53. H. striolata." figureDoi="http://doi.org/10.5281/zenodo.5283496" httpUri="https://zenodo.org/record/5283496/files/figure.png" pageId="41" pageNumber="42">Figs. 53</figureCitation>
,
<figureCitation id="13222A765B4CFFBAFE7AF4C7FE02F894" box="[404,436,1770,1794]" captionStart="FIGURES 56–57" captionStartId="187.[151,264,1650,1673]" captionTargetBox="[282,1301,215,1623]" captionTargetId="figure-29@187.[263,1323,193,1629]" captionTargetPageId="187" captionText="FIGURES 56–57. Aedeagi of holotypes. 56. H. infoveola. 57. H. otiarca." figureDoi="http://doi.org/10.5281/zenodo.5283506" httpUri="https://zenodo.org/record/5283506/files/figure.png" pageId="41" pageNumber="42">56</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4CFFB9FF29F523FA3BFD79" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="8BA636F35B4CFFBAFF29F523FDE7F804" blockId="41.[151,1436,1410,2011]" pageId="41" pageNumber="42">
<emphasis id="B96DEAE15B4CFFBAFF29F523FEEEF8B1" bold="true" box="[199,344,1806,1831]" inLineHeading="true" pageId="41" pageNumber="42" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B4CFFBAFE42F523FDB8F8B0" box="[428,526,1806,1830]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.58/0.68; head 0.23/0.38; pronotum 0.38/0.48, PA 0.42, PB 0.42; elytra 1.00/0.68. Dorsum of head piceous; pronotum testaceous in front of and very narrowly behind piceous fascia, ratios of color bands, as measured in midline, ca. 2/3/1; elytra dark brown; legs and maxillary palpi brown.
</paragraph>
<paragraph id="8BA636F35B4CFFB9FF29F5B2FB10FF61" blockId="41.[151,1436,1410,2011]" lastBlockId="42.[151,1437,151,860]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">Frons punctures ca. 1xef; interstices shining, 1–3xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Pronotum cordiform, median 2/3 of anterior margin emarginate; punctures on disc ca. 1xpd those of frons, interstices shining, 2–3xpd, punctures slightly larger and denser at anterior and posterior; PF1 absent or obsolete; PF2 extremely shallow, nearly imperceptible; PF3 moderately deep; PF4 absent.</paragraph>
<paragraph id="8BA636F35B4FFFB9FF29F329FAAEFE11" blockId="42.[151,1437,151,860]" pageId="42" pageNumber="43">Elytra with summit of posterior declivity at or very near midlength; lateral explanate moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B4FFFB9FF29F3B9FA3BFD79" blockId="42.[151,1437,151,860]" pageId="42" pageNumber="43">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 4/2/7/5. P1 laminate; median carina sinuate in profile. P2 raised, l/w ca. 2/1, sides slightly converging toward apex. Plaques moderately wide, converging very slightly anteriorly, weakly raised, slightly more so laterally than medially, located at sides of deep median depression. Metaventrite with short low ridge extended from each mesoventral coxal cavity. AIS width at straight posterior margin ca. 1.5x P2. All legs long and very slender. Profemur (male) with small, sharply pointed tubercle next to trochanter; protibia slightly arcuate, gradually increasing in width from base to apex, with moderately long spine on medial surface at ca. distal 2/3. Mesotibia moderately arcuate, ca. to same degree as protibia, slightly thickened at distal 2/ 3. Metatibia very slightly arcuate, gradually increasing in width from base to apex, medial surface, with few moderately long spines in distal 1/4. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus (
<figureCitation id="13222A765B4FFFB9FA81F099FF02FD79" captionStart="FIGURES 56–57" captionStartId="187.[151,264,1650,1673]" captionTargetBox="[282,1301,215,1623]" captionTargetId="figure-29@187.[263,1323,193,1629]" captionTargetPageId="187" captionText="FIGURES 56–57. Aedeagi of holotypes. 56. H. infoveola. 57. H. otiarca." figureDoi="http://doi.org/10.5281/zenodo.5283506" httpUri="https://zenodo.org/record/5283506/files/figure.png" pageId="42" pageNumber="43">Fig. 56</figureCitation>
) distal piece with sharply pointed, slightly arcuate process on left side, and large, arm-shaped process apically.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4FFFB9FF29F0D1FBD1FC82" box="[199,1127,764,788]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BA636F35B4FFFB9FF29F0D1FBD1FC82" blockId="42.[151,1437,151,860]" box="[199,1127,764,788]" pageId="42" pageNumber="43">
<emphasis id="B96DEAE15B4FFFB9FF29F0D1FEFBFC82" bold="true" box="[199,333,764,788]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Etymology.</emphasis>
Named in reference to the greatly reduced and absent pronotal foveae.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4FFFB9FF29F132FCF1FCCA" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BA636F35B4FFFB9FF29F132FCF1FCCA" blockId="42.[151,1437,151,860]" pageId="42" pageNumber="43">
<emphasis id="B96DEAE15B4FFFB9FF29F132FED6FCAE" bold="true" box="[199,352,799,824]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Distribution.</emphasis>
Currently known only from two very narrowly separated sites near Marawaka, at the eastern extreme of Area 1; elevations
<quantity id="4CE19B165B4FFFB9FE07F169FD88FCCD" box="[489,574,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1000.0">1000 m</quantity>
and
<quantity id="4CE19B165B4FFFB9FD9AF169FD7FFCCD" box="[628,713,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="42" pageNumber="43" unit="m" value="1400.0">1400 m</quantity>
(
<figureCitation id="13222A765B4FFFB9FD36F169FC8CFCCA" box="[728,826,836,860]" captionStart="FIGURES 435–438" captionStartId="384.[151,264,1650,1673]" captionTargetBox="[165,1421,235,1613]" captionTargetId="figure-23@384.[165,1421,193,1629]" captionTargetPageId="384" captionText="FIGURES 435–438. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284069" httpUri="https://zenodo.org/record/5284069/files/figure.png" pageId="42" pageNumber="43">Fig. 438</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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