treatments-xml/data/03/AE/CB/03AECB0EDC45FFFFFE93378A53C6924E.xml

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<document id="92EBAD27AD92E6EB2971672351086DA5" ID-DOI="10.5281/zenodo.5301059" ID-ISSN="0374-1036" ID-Zenodo-Dep="5301059" ID-ZooBank="EA465CB7-1796-4656-9D14-EF0964899EDAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630099098575" checkinUser="marcus" docAuthor="Makranczy, György" docDate="2014" docId="03AECB0EDC45FFFFFE93378A53C6924E" docLanguage="en" docName="ActaEntMusNatPra.2014.54.2.539-554.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thinodromus chagosanus" docType="treatment" docVersion="4" lastPageNumber="551" masterDocId="FF97B376DC4CFFF3FFF33756520A915D" masterDocTitle="Review of the Thinodromus circulus species group (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="554" masterPageNumber="539" pageNumber="548" updateTime="1699149139555" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8D58F1EA9A843F5B883D3BD16890ABEC">Review of the Thinodromus circulus species group (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="ADCA788C039629F181BA8D0D5A7ADB66">Makranczy, György</mods:namePart>
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<mods:date id="5B21DBA8C603E20B84182F0A75AE310A">2014</mods:date>
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<subSubSection id="C31D2993DC45FFFAFE93378A515891AE" box="[352,850,220,243]" pageId="9" pageNumber="548" type="nomenclature">
<paragraph id="8BB87A18DC45FFFAFE93378A515891AE" blockId="9.[352,850,220,243]" box="[352,850,220,243]" pageId="9" pageNumber="548">
<heading id="D0F0CD74DC45FFFAFE93378A515891AE" bold="true" box="[352,850,220,243]" centered="true" fontSize="8" level="2" pageId="9" pageNumber="548" reason="3">
<taxonomicName id="4C07019BDC45FFFAFE93378A515891AE" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[352,850,220,243]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="chagosanus">
<emphasis id="B973A60ADC45FFFAFE93378A515891AE" bold="true" box="[352,850,220,243]" pageId="9" pageNumber="548">
<emphasis id="B973A60ADC45FFFAFE93378A507491AE" bold="true" box="[352,638,220,243]" italics="true" pageId="9" pageNumber="548">Thinodromus chagosanus</emphasis>
(Bernhauer, 1922)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC45FFFAFE0437AC50B09050" box="[503,698,250,269]" pageId="9" pageNumber="548" type="description">
<paragraph id="8BB87A18DC45FFFAFE0437AC50B09050" blockId="9.[113,1089,250,356]" box="[503,698,250,269]" pageId="9" pageNumber="548">
(
<figureCitation id="133C669DDC45FFFAFE0E37AC503F9050" box="[509,565,250,269]" captionStart="Figs 14" captionStartId="2.[151,188,1526,1545]" captionTargetBox="[163,1115,218,1465]" captionTargetId="figure-10@2.[163,1116,217,1465]" captionTargetPageId="2" captionText="Figs 14. Forebodies of Thinodromus species. 1 T. circulus (Bernhauer, 1922); 2 T. chagosanus (Bernhauer, 1922); 3 T. palustris (Bernhauer, 1922); 4 T. arcitenens (Fauvel, 1905)." figureDoi="http://doi.org/10.5281/zenodo.5301061" httpUri="https://zenodo.org/record/5301061/files/figure.png" pageId="9" pageNumber="548">Figs 2</figureCitation>
,
<figureCitation id="133C669DDC45FFFAFDCD37AC507F9050" box="[574,629,250,269]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="9" pageNumber="548">1314</figureCitation>
,
<figureCitation id="133C669DDC45FFFAFD8D37AC50BF9050" box="[638,693,250,269]" captionStart-0="Figs 2024. 20" captionStart-1="Figs 2530. 2526" captionStartId-0="7.[113,150,1480,1499]" captionStartId-1="10.[151,188,1493,1512]" captionTargetBox-0="[127,1077,222,1463]" captionTargetBox-1="[151,1103,219,1465]" captionTargetId-0="figure-9@7.[125,1078,219,1466]" captionTargetId-1="figure-9@10.[151,1103,219,1466]" captionTargetPageId-0="7" captionTargetPageId-1="10" captionText-0="Figs 2024. 20 Thinodromus velutinus (Sharp, 1887), tergite VIII; 2122 T. circulus (Bernhauer, 1922): 21 tergite VIII, 22 aedeagus; 2324 T. chagosanus (Bernhauer, 1922): 23 male sternite IX, 24 aedeagus. Scale bar: 0.1 mm for 2021; 0.15 mm for 2224." captionText-1="Figs 2530. 2526 Thinodromus chagosanus (Bernhauer, 1922): 25 male sternite VIII, 26 male tergite X. 2728 T. palustris (Bernhauer, 1922): 27 male sternite VIII, 28 male tergite X. 2930 T. arcitenens (Fauvel, 1905): 29 male sternite VIII, 30 male tergite X. Scale bar: 0.07 mm for 30; 0.08 mm for 2829; 0.1 mm for 2627; 0.11 mm for 25." figureDoi-0="http://doi.org/10.5281/zenodo.5301069" figureDoi-1="http://doi.org/10.5281/zenodo.5301073" httpUri-0="https://zenodo.org/record/5301069/files/figure.png" httpUri-1="https://zenodo.org/record/5301073/files/figure.png" pageId="9" pageNumber="548">2326</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC45FFFAFF82364A51969039" pageId="9" pageNumber="548" type="reference_group">
<paragraph id="8BB87A18DC45FFFAFF82364A52FC9017" blockId="9.[113,1089,250,356]" pageId="9" pageNumber="548">
<treatmentCitationGroup id="AB175D36DC45FFFAFF82364A52FC9017" pageId="9" pageNumber="548">
<treatmentCitation id="0AA65C09DC45FFFAFF82364A50929072" author="BERNHAUER M." box="[113,664,284,303]" page="167" pageId="9" pageNumber="548" year="1922">
<taxonomicName id="4C07019BDC45FFFAFF82364A50929072" authority="Bernhauer, 1922 b: 167" authorityName="Bernhauer" authorityPageNumber="167" authorityYear="1922" box="[113,664,284,303]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="chagosanus" subGenus="Carpalimus">
<emphasis id="B973A60ADC45FFFAFF82364A52EC9072" box="[113,230,284,303]" italics="true" pageId="9" pageNumber="548">Trogophloeus</emphasis>
(
<emphasis id="B973A60ADC45FFFAFF07364A53539072" box="[244,345,284,303]" italics="true" pageId="9" pageNumber="548">Carpalimus</emphasis>
)
<emphasis id="B973A60ADC45FFFAFE95364A53C19072" box="[358,459,284,303]" italics="true" pageId="9" pageNumber="548">chagosanus</emphasis>
<bibRefCitation id="EF9607E9DC45FFFAFE21364A50929072" author="BERNHAUER M." box="[466,664,284,303]" pageId="9" pageNumber="548" pagination="165 - 186" refId="ref7011" refString="BERNHAUER M. 1922 b: Coleoptera, Staphylinidae. (Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, M. A., Volume VII.) Transactions of the Linnean Society of London, Series 2, Zoology 18 (1): 165 - 186." type="journal article" year="1922">Bernhauer, 1922b: 167</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description),
<treatmentCitation id="0AA65C09DC45FFFAFC92364A564B9072" author="SCHEERPELTZ O." box="[865,1089,284,303]" page="1081" pageId="9" pageNumber="548" year="1933">
<bibRefCitation id="EF9607E9DC45FFFAFC92364A56079072" author="SCHEERPELTZ O." box="[865,1037,284,303]" pageId="9" pageNumber="548" pagination="989 - 1500" refId="ref7358" refString="SCHEERPELTZ O. 1933: Staphylinidae VII. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 129. [Supplementum I.]. W. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">SCHEERPELTZ (1933)</bibRefCitation>
: 1081
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB87A18DC45FFFAFF82360751969039" blockId="9.[113,1089,250,356]" box="[113,924,337,356]" pageId="9" pageNumber="548">
<treatmentCitationGroup id="AB175D36DC45FFFAFF82360751969039" box="[113,924,337,356]" pageId="9" pageNumber="548">
<treatmentCitation id="0AA65C09DC45FFFAFF823607500C9039" author="HERMAN L. H." box="[113,518,337,356]" page="387" pageId="9" pageNumber="548" year="1970">
<taxonomicName id="4C07019BDC45FFFAFF82360753D49039" authority=": HERMAN (1970)" authorityName="HERMAN" authorityYear="1970" box="[113,478,337,356]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="chagosanus">
<emphasis id="B973A60ADC45FFFAFF82360753479039" box="[113,333,337,356]" italics="true" pageId="9" pageNumber="548">Thinodromus chagosanus</emphasis>
:
<bibRefCitation id="EF9607E9DC45FFFAFEAB360753D49039" author="HERMAN L. H." box="[344,478,337,356]" pageId="9" pageNumber="548" pagination="343 - 454" refId="ref7167" refString="HERMAN L. H. 1970: Phylogeny and reclassi? cation of the genera of the rove-beetle subfamily Oxytelinae of the World (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History 142 (5): 343 - 454." type="journal article" year="1970">HERMAN (1970)</bibRefCitation>
</taxonomicName>
: 387
</treatmentCitation>
(catalogue),
<treatmentCitation id="0AA65C09DC45FFFAFD853607513A9039" author="HERMAN L. H." box="[630,816,337,356]" page="1765" pageId="9" pageNumber="548" year="2001">
<bibRefCitation id="EF9607E9DC45FFFAFD85360750F79039" author="HERMAN L. H." box="[630,765,337,356]" pageId="9" pageNumber="548" pagination="1 - 4218" refId="ref7213" refString="HERMAN L. H. 2001: Catalog of the Staphylinidae (Insecta: Coleoptera). 1785 to the end of the second Millennium. Volume I - VII. Bulletin of the American Museum of Natural History 265: 1 - 4218." type="journal article" year="2001">HERMAN (2001)</bibRefCitation>
: 1765
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC45FFFAFF82362D53259314" pageId="9" pageNumber="548" type="materials_examined">
<paragraph id="8BB87A18DC45FFFAFF82362D50F390D3" blockId="9.[113,1094,379,585]" box="[113,761,379,398]" pageId="9" pageNumber="548">
<emphasis id="B973A60ADC45FFFAFF82362D52E090D3" bold="true" box="[113,234,379,398]" pageId="9" pageNumber="548">
<typeStatus id="54BCC4BADC45FFFAFF82362D529490D3" box="[113,158,379,398]" pageId="9" pageNumber="548">Type</typeStatus>
locality.
</emphasis>
Chagos Archipelago, Diego Garcia, approx. 7°18ƍS, 72°24ƍE.
</paragraph>
<paragraph id="8BB87A18DC45FFFAFF8236C051EF9083" blockId="9.[113,1094,379,585]" pageId="9" pageNumber="548">
<emphasis id="B973A60ADC45FFFAFF8236C0535C90F4" bold="true" box="[113,342,406,425]" pageId="9" pageNumber="548">Type material examined.</emphasis>
<typeStatus id="54BCC4BADC45FFFAFEAF36C053BF90FA" box="[348,437,406,425]" pageId="9" pageNumber="548" type="lectotype">LECTOTYPE</typeStatus>
(here designated): J, Chagos.; Diego Garcia \ 9.VII.[19]05 \ chagosanus; Bernh.; Cotypus. Scot \ Chicago NHMus; M. Bernhauer; Collection \
<typeStatus id="54BCC4BADC45FFFAFD3336E751289099" box="[704,802,433,452]" pageId="9" pageNumber="548" type="lectotype">Lectotypus</typeStatus>
;
<taxonomicName id="4C07019BDC45FFFAFCDA36E751A89099" box="[809,930,433,452]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C07019BDC45FFFAFC5A36E752A99083" authority="Bernhauer" authorityName="Bernhauer" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="chagosanus">chagosanus Bernhauer</taxonomicName>
; des.
<bibRefCitation id="EF9607E9DC45FFFAFF20369D536D9083" author="MAKRANCZY GY" box="[211,359,459,478]" pageId="9" pageNumber="548" pagination="177 - 208" refId="ref7326" refString="MAKRANCZY GY. 2013: Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae). Acta Entomologica Musei Nationalis Pragae 53: 177 - 208." type="journal article" year="2013">Makranczy, 2013</bibRefCitation>
\
<taxonomicName id="4C07019BDC45FFFAFE84369D53E79083" authorityName="Kraatz" authorityYear="1857" box="[375,493,459,478]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
;
<taxonomicName id="4C07019BDC45FFFAFE06369D50CD9083" authority="(Bernhauer)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[501,711,459,478]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="chagosanus">chagosanus (Bernhauer)</taxonomicName>
; det.
<bibRefCitation id="EF9607E9DC45FFFAFD07369D51809083" author="MAKRANCZY GY" box="[756,906,459,478]" pageId="9" pageNumber="548" pagination="177 - 208" refId="ref7326" refString="MAKRANCZY GY. 2013: Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae). Acta Entomologica Musei Nationalis Pragae 53: 177 - 208." type="journal article" year="2013">Makranczy, 2013</bibRefCitation>
(
<collectionCode id="ED16E2DDDC45FFFAFC6A369D51D69083" box="[921,988,459,478]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="548" type="Museum">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8BB87A18DC45FFFAFF8236B053259314" blockId="9.[113,1094,379,585]" pageId="9" pageNumber="548">
<emphasis id="B973A60ADC45FFFAFF8236B0506190A4" bold="true" box="[113,619,486,505]" pageId="9" pageNumber="548">Other material examined. CHAGOS ARCHIPELAGO:</emphasis>
Diego Garcia, Pointe Marianne,
<date id="FFB95CD8DC45FFFAFC7436B051EC90A4" box="[903,998,486,505]" pageId="9" pageNumber="548" value="1971-04-12">12.IV.1971</date>
, leg. A.M. Hutson (
<collectionCode id="ED16E2DDDC45FFFAFF49355752F19349" box="[186,251,513,532]" pageId="9" pageNumber="548">BMNH</collectionCode>
accession number 1971-346), at light (1 microscopic slide,
<collectionCode id="ED16E2DDDC45FFFAFD04355751369349" box="[759,828,513,532]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="548">HNHM</collectionCode>
).
<emphasis id="B973A60ADC45FFFAFCB9355751EA9349" bold="true" box="[842,992,513,532]" pageId="9" pageNumber="548">
<collectingCountry id="F3103A88DC45FFFAFCB9355751D69349" box="[842,988,513,532]" name="Seychelles" pageId="9" pageNumber="548">SEYCHELLES</collectingCountry>
:
</emphasis>
Mahé Sud, Anse à la Mouche,
<date id="FFB95CD8DC45FFFAFEEE354D53A39373" box="[285,425,539,558]" pageId="9" pageNumber="548" value="1972-08-01" valueMax="1972-08-15" valueMin="1972-08-01">1.15.VIII.1972</date>
, leg. P.L.G. Benoit &amp; J.J. van Mol (Miss. zool. belge aux Séchelles) (2 JJ
<collectionCode id="ED16E2DDDC45FFFAFF82356052B89314" box="[113,178,566,585]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="548" type="Museum">MRAC</collectionCode>
, 1 J
<collectionCode id="ED16E2DDDC45FFFAFF133560532F9314" box="[224,293,566,585]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="548">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC45FFFFFF823537503C90CE" lastPageId="12" lastPageNumber="551" pageId="9" pageNumber="548" type="description">
<paragraph id="8BB87A18DC45FFFFFF823537503C90CE" blockId="9.[113,1090,609,1592]" lastBlockId="12.[151,1127,220,787]" lastPageId="12" lastPageNumber="551" pageId="9" pageNumber="548">
<emphasis id="B973A60ADC45FFFAFF823537531D9325" bold="true" box="[113,279,609,632]" pageId="9" pageNumber="548">Redescription.</emphasis>
<emphasis id="B973A60ADC45FFFAFEED353753B39325" box="[286,441,609,632]" italics="true" pageId="9" pageNumber="548">Measurements</emphasis>
(in mm, n = 4): HW = 0.54 (0.510.55); TW = 0.48 (0.45 0.49); PW = 0.53 (0.500.56); SW = 0.655 (0.620.68); AW = 0.735 (0.680.76); HL = 0.35 (0.340.35); EL = 0.21 (0.210.22); TL = 0.04 (0.030.04); PL = 0.41 (0.400.42); SL = 0.665 (0.650.68); SC = 0.62 (0.610.63); FB = 1.43 (1.401.46); BL = 2.65 (2.602.69).
<emphasis id="B973A60ADC45FFFAFF8235B7533B93A5" box="[113,305,737,760]" italics="true" pageId="9" pageNumber="548">Lustre and colour</emphasis>
. Fine punctation on foreparts and shallow microsculpture of body providing a greasy lustre. Head, pronotum and abdomen brownish black, elytra very dark brown, sometimes slightly reddish apical edge and outer posterior corners often a little darker, black. Legs medium brown, apices of femora and middle of tibiae darker, blackish. Mouthparts and antennae dark brown, latter with ¿rst segment often lighter.
<emphasis id="B973A60ADC45FFFAFD2C343751BE9225" box="[735,948,865,888]" italics="true" pageId="9" pageNumber="548">Shape and sculpture</emphasis>
. Forebody as in
<figureCitation id="133C669DDC45FFFAFF7834D752C192C5" box="[139,203,897,920]" captionStart="Figs 14" captionStartId="2.[151,188,1526,1545]" captionTargetBox="[163,1115,218,1465]" captionTargetId="figure-10@2.[163,1116,217,1465]" captionTargetPageId="2" captionText="Figs 14. Forebodies of Thinodromus species. 1 T. circulus (Bernhauer, 1922); 2 T. chagosanus (Bernhauer, 1922); 3 T. palustris (Bernhauer, 1922); 4 T. arcitenens (Fauvel, 1905)." figureDoi="http://doi.org/10.5281/zenodo.5301061" httpUri="https://zenodo.org/record/5301061/files/figure.png" pageId="9" pageNumber="548">Fig. 2</figureCitation>
. Head transverse, eyes very large, occupying sides of head, temples marked but very small, length insigni¿cant compared to that of eye. Neck delineated only by different, alveolate microsculpture, but without transversal groove. Antennae slightly elongate, antennomeres 4 and 5 1.051.08× and 1.311.35× longer than broad, antennomere 9 1.001.04× longer than broad, antennomere 10 approximately as long as broad. Pronotum rather transverse, strongly obtuse-angled anterior corners super¿cially appear somewhat rounded but still marked. Posterior half of pronotal sides quite straight, even feebly concave; posterior corners obtuse-angled and rounded. Horseshoe-shaped impression slightly marked except posterior/median part where rather impressed; slight lateral depressions connected by it. Middle of disc bearing pair of shallow depressions. Slight (thin) marginal bead (mostly lateral) of pronotum observable only in sublateral view. Elytra combined imperceptably broader than long, gently dilated towards apex, with a pair of small, oval, slightly elongate impressions behind scutellum and extending posteriorly in longitudinal impressions, connecting to somewhat depressed centre of elytral disc. Posterior elytral margin (slightly oblique) with very thin bead and in outer 1/3 with small membranous lobe protruding. Apex of abdominal tergite VII with palisade fringe.
<emphasis id="B973A60ADC45FFFAFF4C323750039425" box="[191,521,1377,1400]" italics="true" pageId="9" pageNumber="548">Punctation and microsculpture</emphasis>
. Head and pronotum punctation rather ¿ne, but deep and quite dense, mixed in with colliculate miscrosculpture; interspaces only a fraction of puncture diameters. Epistomal suture marked as a less punctate/sculptured area, with trace of transversal strigulate microsculpture. Punctation less deep and dense on ridges, e.g. outside/along roundish depressions at centre of pronotal disc. Microsculpture more rough and scabrous in depressions and near posterior pronotal corners, on pronotum generally stronger on sides than centre of disc. Elytra with slightly larger punctures, but also larger interspaces with less distinct microsculpture so appearing a little more lustrous.Abdomen with scattered, ¿ne but distinct punctures, slightly imbricate or coriaceous microsculpture with isodiametric cells and a rather greasy lustre.
<emphasis id="B973A60ADC40FFFFFE1B364A506C906E" box="[488,614,284,307]" italics="true" pageId="12" pageNumber="551">Pubescence</emphasis>
. Body setation short, ¿ne, but rather dense, hair sizes vary more on abdomen, apices of segments with longer hairs.
<emphasis id="B973A60ADC40FFFFFC99366A5332902E" italics="true" pageId="12" pageNumber="551">Primary and secondary sexual features</emphasis>
. Male sternite VIII as in
<figureCitation id="133C669DDC40FFFFFDB4360A509F902E" box="[583,661,348,371]" captionStart="Figs 2530. 2526" captionStartId="10.[151,188,1493,1512]" captionTargetBox="[151,1103,219,1465]" captionTargetId="figure-9@10.[151,1103,219,1466]" captionTargetPageId="10" captionText="Figs 2530. 2526 Thinodromus chagosanus (Bernhauer, 1922): 25 male sternite VIII, 26 male tergite X. 2728 T. palustris (Bernhauer, 1922): 27 male sternite VIII, 28 male tergite X. 2930 T. arcitenens (Fauvel, 1905): 29 male sternite VIII, 30 male tergite X. Scale bar: 0.07 mm for 30; 0.08 mm for 2829; 0.1 mm for 2627; 0.11 mm for 25." figureDoi="http://doi.org/10.5281/zenodo.5301073" httpUri="https://zenodo.org/record/5301073/files/figure.png" pageId="12" pageNumber="551">Fig. 25</figureCitation>
, male sternite IX as in
<figureCitation id="133C669DDC40FFFFFC7D360A51D6902E" box="[910,988,348,371]" captionStart="Figs 2024. 20" captionStartId="7.[113,150,1480,1499]" captionTargetBox="[127,1077,222,1463]" captionTargetId="figure-9@7.[125,1078,219,1466]" captionTargetPageId="7" captionText="Figs 2024. 20 Thinodromus velutinus (Sharp, 1887), tergite VIII; 2122 T. circulus (Bernhauer, 1922): 21 tergite VIII, 22 aedeagus; 2324 T. chagosanus (Bernhauer, 1922): 23 male sternite IX, 24 aedeagus. Scale bar: 0.1 mm for 2021; 0.15 mm for 2224." figureDoi="http://doi.org/10.5281/zenodo.5301069" httpUri="https://zenodo.org/record/5301069/files/figure.png" pageId="12" pageNumber="551">Fig. 23</figureCitation>
, male tergite X as in
<figureCitation id="133C669DDC40FFFFFF1A362A533D90CE" box="[233,311,380,403]" captionStart="Figs 2530. 2526" captionStartId="10.[151,188,1493,1512]" captionTargetBox="[151,1103,219,1465]" captionTargetId="figure-9@10.[151,1103,219,1466]" captionTargetPageId="10" captionText="Figs 2530. 2526 Thinodromus chagosanus (Bernhauer, 1922): 25 male sternite VIII, 26 male tergite X. 2728 T. palustris (Bernhauer, 1922): 27 male sternite VIII, 28 male tergite X. 2930 T. arcitenens (Fauvel, 1905): 29 male sternite VIII, 30 male tergite X. Scale bar: 0.07 mm for 30; 0.08 mm for 2829; 0.1 mm for 2627; 0.11 mm for 25." figureDoi="http://doi.org/10.5281/zenodo.5301073" httpUri="https://zenodo.org/record/5301073/files/figure.png" pageId="12" pageNumber="551">Fig. 26</figureCitation>
, aedeagus as in
<figureCitation id="133C669DDC40FFFFFE10362A503890CE" box="[483,562,380,403]" captionStart="Figs 2024. 20" captionStartId="7.[113,150,1480,1499]" captionTargetBox="[127,1077,222,1463]" captionTargetId="figure-9@7.[125,1078,219,1466]" captionTargetPageId="7" captionText="Figs 2024. 20 Thinodromus velutinus (Sharp, 1887), tergite VIII; 2122 T. circulus (Bernhauer, 1922): 21 tergite VIII, 22 aedeagus; 2324 T. chagosanus (Bernhauer, 1922): 23 male sternite IX, 24 aedeagus. Scale bar: 0.1 mm for 2021; 0.15 mm for 2224." figureDoi="http://doi.org/10.5281/zenodo.5301069" httpUri="https://zenodo.org/record/5301069/files/figure.png" pageId="12" pageNumber="551">Fig. 24</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF782A90DC46FFF9FF64328353689765" ID-DOI="http://doi.org/10.5281/zenodo.5301073" ID-Zenodo-Dep="5301073" httpUri="https://zenodo.org/record/5301073/files/figure.png" pageId="10" pageNumber="549" startId="10.[151,188,1493,1512]" targetBox="[151,1103,219,1465]" targetPageId="10">
<paragraph id="8BB87A18DC46FFF9FF64328353689765" blockId="10.[151,1128,1493,1592]" pageId="10" pageNumber="549">
Figs 2530. 2526
<taxonomicName id="4C07019BDC46FFF9FEA0328350DD94B5" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[339,727,1493,1512]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="549" phylum="Arthropoda" rank="species" species="chagosanus">
<emphasis id="B973A60ADC46FFF9FEA03283503A94B5" box="[339,560,1493,1512]" italics="true" pageId="10" pageNumber="549">Thinodromus chagosanus</emphasis>
(Bernhauer, 1922)
</taxonomicName>
: 25 male sternite VIII, 26 male tergite X. 2728
<taxonomicName id="4C07019BDC46FFF9FF1232B953EE975F" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[225,484,1519,1538]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="549" phylum="Arthropoda" rank="species" species="palustris">
<emphasis id="B973A60ADC46FFF9FF1232B9534B975F" box="[225,321,1519,1538]" italics="true" pageId="10" pageNumber="549">T. palustris</emphasis>
(Bernhauer, 1922)
</taxonomicName>
: 27 male sternite VIII, 28 male tergite X. 2930
<taxonomicName id="4C07019BDC46FFF9FC5D32B952C79740" authority="(Fauvel, 1905)" baseAuthorityName="Fauvel" baseAuthorityYear="1905" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="549" phylum="Arthropoda" rank="species" species="arcitenens">
<emphasis id="B973A60ADC46FFF9FC5D32B95611975F" box="[942,1051,1519,1538]" italics="true" pageId="10" pageNumber="549">T. arcitenens</emphasis>
(
<bibRefCitation id="EF9607E9DC46FFF9FBD532B952CF9740" author="FAUVEL A." pageId="10" pageNumber="549" pagination="75 - 86" refId="ref7119" refString="FAUVEL A. 1905: Staphylinides de Java, recueillis par M. le Dr. K. Kraepelin et M. le Dr. [J. C.] Koningsberger en 1904. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 22: 75 - 86." type="journal article" year="1905">Fauvel, 1905</bibRefCitation>
)
</taxonomicName>
: 29 male sternite VIII, 30 male tergite X. Scale bar: 0.07 mm for 30; 0.08 mm for 2829; 0.1 mm for 2627; 0.11 mm for 25.
</paragraph>
</caption>
<caption id="DF782A90DC47FFF8FF8232A151F99762" ID-DOI="http://doi.org/10.5281/zenodo.5301075" ID-Zenodo-Dep="5301075" httpUri="https://zenodo.org/record/5301075/files/figure.png" pageId="11" pageNumber="550" startId="11.[113,150,1527,1546]" targetBox="[114,1064,219,1466]" targetPageId="11">
<paragraph id="8BB87A18DC47FFF8FF8232A151F99762" blockId="11.[113,1089,1527,1599]" pageId="11" pageNumber="550">
Figs 3135. 31
<taxonomicName id="4C07019BDC47FFF8FEF832A150449757" authority="(Sharp, 1887)" authorityName="Makranczy" authorityYear="2014" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[267,590,1527,1546]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="550" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B973A60ADC47FFF8FEF832A153C59757" box="[267,463,1527,1546]" italics="true" pageId="11" pageNumber="550">Thinodromus velutinus</emphasis>
(
<bibRefCitation id="EF9607E9DC47FFF8FE2F32A1504D9757" author="SHARP D. S." box="[476,583,1527,1546]" pageId="11" pageNumber="550" pagination="673 - 824" refId="ref7488" refString="SHARP D. S. 1887: Staphylinidae. Pp. 673 - 824, pls. 17 - 19. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part 2. Taylor &amp; Francis, London, xvi + 824 pp., 19 pls." type="book chapter" year="1887">Sharp, 1887</bibRefCitation>
)
</taxonomicName>
, female ringstructure. 32
<taxonomicName id="4C07019BDC47FFF8FCCD32A156379757" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[830,1085,1527,1546]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="550" phylum="Arthropoda" rank="species" species="circulus">
<emphasis id="B973A60ADC47FFF8FCCD32A151929757" box="[830,920,1527,1546]" italics="true" pageId="11" pageNumber="550">T. circulus</emphasis>
(Bernhauer, 1922)
</taxonomicName>
, female ringstructure. 3334
<taxonomicName id="4C07019BDC47FFF8FE9D314750679779" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[366,621,1553,1572]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="550" phylum="Arthropoda" rank="species" species="palustris">
<emphasis id="B973A60ADC47FFF8FE9D314753C79779" box="[366,461,1553,1572]" italics="true" pageId="11" pageNumber="550">T. palustris</emphasis>
(Bernhauer, 1922)
</taxonomicName>
: 33 male sternite IX, 34 aedeagus. 35
<taxonomicName id="4C07019BDC47FFF8FC26314752FB9762" authority="(Fauvel, 1905)" baseAuthorityName="Fauvel" baseAuthorityYear="1905" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="550" phylum="Arthropoda" rank="species" species="arcitenens">
<emphasis id="B973A60ADC47FFF8FC263147564A9779" box="[981,1088,1553,1572]" italics="true" pageId="11" pageNumber="550">T.arcitenens</emphasis>
(
<bibRefCitation id="EF9607E9DC47FFF8FF84317A52E09762" author="FAUVEL A." box="[119,234,1580,1599]" pageId="11" pageNumber="550" pagination="75 - 86" refId="ref7119" refString="FAUVEL A. 1905: Staphylinides de Java, recueillis par M. le Dr. K. Kraepelin et M. le Dr. [J. C.] Koningsberger en 1904. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 22: 75 - 86." type="journal article" year="1905">Fauvel, 1905</bibRefCitation>
)
</taxonomicName>
, aedeagus. Scale bar: 0.1 mm for 31; 0.12 mm for 32, 35; 0.15 mm for 33; 0.2 mm for 34.
</paragraph>
</caption>
<subSubSection id="C31D2993DC40FFFFFF6436CA5055908E" pageId="12" pageNumber="551" type="diagnosis">
<paragraph id="8BB87A18DC40FFFFFF6436CA5055908E" blockId="12.[151,1127,220,787]" pageId="12" pageNumber="551">
<emphasis id="B973A60ADC40FFFFFF6436CA538690EE" bold="true" box="[151,396,412,435]" pageId="12" pageNumber="551">Differential diagnosis.</emphasis>
<taxonomicName id="4C07019BDC40FFFFFE6336CA509690EE" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[400,668,412,435]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="551" phylum="Arthropoda" rank="species" species="chagosanus">
<emphasis id="B973A60ADC40FFFFFE6336CA509690EE" box="[400,668,412,435]" italics="true" pageId="12" pageNumber="551">Thinodromus chagosanus</emphasis>
</taxonomicName>
appears to be the only species in this species group with small (barely marked) temples.
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC40FFFFFF64368A50EF934E" pageId="12" pageNumber="551" type="distribution">
<paragraph id="8BB87A18DC40FFFFFF64368A50EF934E" blockId="12.[151,1127,220,787]" pageId="12" pageNumber="551">
<emphasis id="B973A60ADC40FFFFFF64368A532090AE" bold="true" box="[151,298,476,499]" pageId="12" pageNumber="551">Distribution.</emphasis>
Known from the
<collectingCountry id="F3103A88DC40FFFFFE1F368A505490AE" box="[492,606,476,499]" name="Seychelles" pageId="12" pageNumber="551">Seychelles</collectingCountry>
and the Chagos Archipelago, therefore likely to be widespread across tropical islands of the Old World.
</paragraph>
</subSubSection>
<subSubSection id="C31D2993DC40FFFFFF64354A53C6924E" pageId="12" pageNumber="551" type="discussion">
<paragraph id="8BB87A18DC40FFFFFF64354A53C6924E" blockId="12.[151,1127,220,787]" pageId="12" pageNumber="551">
<emphasis id="B973A60ADC40FFFFFF64354A530C936E" bold="true" box="[151,262,540,563]" pageId="12" pageNumber="551">Remarks.</emphasis>
The plate-like armature in the hypopharynx illustrated in
<bibRefCitation id="EF9607E9DC40FFFFFC81354A5668936E" author="MAKRANCZY GY" box="[882,1122,540,563]" pageId="12" pageNumber="551" pagination="29 - 119" refId="ref7258" refString="MAKRANCZY GY. 2006: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 98: 29 - 119." type="journal article" year="2006">MAKRANCZY (2006: 60</bibRefCitation>
, Fig. 42) is erroneously captioned as
<taxonomicName id="4C07019BDC40FFFFFDEE356A50BF930E" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[541,693,572,595]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="551" phylum="Arthropoda" rank="species" species="chagosanus">
<emphasis id="B973A60ADC40FFFFFDEE356A50BF930E" box="[541,693,572,595]" italics="true" pageId="12" pageNumber="551">T. chagosanus</emphasis>
</taxonomicName>
, while in fact it belongs to
<taxonomicName id="4C07019BDC40FFFFFC28356A5342932E" authority="Gildenkov, 2000" authorityName="Gildenkov" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="551" phylum="Arthropoda" rank="species" species="thoracicus">
<emphasis id="B973A60ADC40FFFFFC28356A566D930E" box="[987,1127,572,595]" italics="true" pageId="12" pageNumber="551">T. thoracicus</emphasis>
Gildenkov, 2000
</taxonomicName>
(full taxonomic treatment in
<bibRefCitation id="EF9607E9DC40FFFFFD72350A514A932E" author="MAKRANCZY GY" box="[641,832,604,627]" pageId="12" pageNumber="551" pagination="33 - 61" refId="ref7293" refString="MAKRANCZY GY. 2009: The genus Thinodromus Kraatz, 1857 in West-Central Africa (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 101: 33 - 61." type="journal article" year="2009">MAKRANCZY 2009</bibRefCitation>
), the correct one for
<taxonomicName id="4C07019BDC40FFFFFBEE350A52E493CE" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="551" phylum="Arthropoda" rank="species" species="chagosanus">
<emphasis id="B973A60ADC40FFFFFBEE350A52E493CE" italics="true" pageId="12" pageNumber="551">T. chagosanus</emphasis>
</taxonomicName>
is in
<figureCitation id="133C669DDC40FFFFFED8352A537193CE" box="[299,379,636,659]" captionStart="Figs 514. 510" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 514. 510 Thinodromus circulus (Bernhauer, 1922) (5 mesotarsus; 6 male sternite VIII; 7 male tergite IX; 8 male sternite IX; 9 male tergite X; 10 spermatheca); 1112 T. velutinus (Sharp, 1887) (11 tergite X; 12 spermatheca); 1314 T. chagosanus (Bernhauer, 1922) (13 mesotarsus; 14 platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 1012; 0.12 mm for 89; 0.14 mm for 67." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="12" pageNumber="551">Fig. 14</figureCitation>
. It must be noted that the two are of the same general structure but of slightly different shape. It is also remarkable that
<taxonomicName id="4C07019BDC40FFFFFD4F35CA514393EE" authorityName="Gildenkov" authorityYear="2000" box="[700,841,668,691]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="551" phylum="Arthropoda" rank="species" species="thoracicus">
<emphasis id="B973A60ADC40FFFFFD4F35CA514393EE" box="[700,841,668,691]" italics="true" pageId="12" pageNumber="551">T. thoracicus</emphasis>
</taxonomicName>
has rather similar habitus (
<bibRefCitation id="EF9607E9DC40FFFFFF6C35EA5388938E" author="MAKRANCZY GY" box="[159,386,700,723]" pageId="12" pageNumber="551" pagination="29 - 119" refId="ref7258" refString="MAKRANCZY GY. 2006: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici 98: 29 - 119." type="journal article" year="2006">MAKRANCZY 2006: 88</bibRefCitation>
, Fig. 115) to that of
<taxonomicName id="4C07019BDC40FFFFFDBC35EA50DE938E" baseAuthorityName="Fauvel" baseAuthorityYear="1905" box="[591,724,700,723]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="551" phylum="Arthropoda" rank="species" species="arcitenens">
<emphasis id="B973A60ADC40FFFFFDBC35EA50DE938E" box="[591,724,700,723]" italics="true" pageId="12" pageNumber="551">T. arcitenens</emphasis>
</taxonomicName>
(treated below). Of numerous existing
<typeStatus id="54BCC4BADC40FFFFFF64358A52FE93AE" box="[151,244,732,755]" pageId="12" pageNumber="551" type="syntype">syntypes</typeStatus>
in depositories, the
<typeStatus id="54BCC4BADC40FFFFFE3A358A502193AE" box="[457,555,732,755]" pageId="12" pageNumber="551" type="lectotype">lectotype</typeStatus>
is a male with clear genital traits and chosen to ¿x the interpretation of this species.
</paragraph>
</subSubSection>
</treatment>
</document>