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<document id="0B9FA16E5BA9836E2E8126ED80D45B57" ID-CLB-Dataset="4210" ID-DOI="10.5852/ejt.2021.767.1495" ID-GBIF-Dataset="67bc6a73-4137-4de3-9dc4-ce87972f742d" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514839" ID-ZooBank="9F019E02-95E7-4B6A-B5BC-01F8E26F3466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631927261920" checkinUser="felipe" docAuthor="Nahok, Benchawan, Tumpeesuwan, Sakboworn &amp; Tumpeesuwan, Chanidaporn" docDate="2021" docId="03AC87E7C329FFC89F35FCF39747BBA4" docLanguage="en" docName="ejt-767_142-166.pdf" docOrigin="European Journal of Taxonomy 767 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.767.1495" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria Godwin-Austen 1918" docType="treatment" docVersion="8" lastPageNumber="149" masterDocId="FF95FF9FC32EFFCF9D1FFF9B931BBE42" masterDocTitle="Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)" masterLastPageNumber="166" masterPageNumber="142" pageNumber="149" updateTime="1699449512379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="82A8A60C8771FBB9E051F03F26ED4AA3">Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)</mods:title>
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<mods:namePart id="3777C9393AFD3581355BA66CDCDDF971">Nahok, Benchawan</mods:namePart>
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<mods:affiliation id="DBE037A9CE6B9F7AFCE82F93BC8BED3F">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. &amp; Educational Research Development and Demonstration Institute, Srinakharinwirot University, Ongkharak District, Nakhon Nayok, 26120 Thailand.</mods:affiliation>
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<mods:namePart id="C613CA174439C52B3A67C2F29B820D3C">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="A2CA9B71DE125637E0D4B7B49241C28D">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. &amp; Palaeontological Research and Education Centre, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:namePart id="80D75F87DAB5D45AAA1AEDD403F1C4C5">Tumpeesuwan, Chanidaporn</mods:namePart>
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<mods:affiliation id="31B6B9714E5FB73AA444F2F0710EA06B">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<treatment id="03AC87E7C329FFC89F35FCF39747BBA4" ID-DOI="http://doi.org/10.5281/zenodo.5514723" ID-GBIF-Taxon="188498234" ID-Zenodo-Dep="5514723" LSID="urn:lsid:plazi:treatment:03AC87E7C329FFC89F35FCF39747BBA4" httpUri="http://treatment.plazi.org/id/03AC87E7C329FFC89F35FCF39747BBA4" lastPageNumber="149" pageId="7" pageNumber="149">
<subSubSection id="C31F657AC329FFC89F35FCF39711BDC0" box="[554,1034,871,898]" pageId="7" pageNumber="149" type="nomenclature">
<paragraph id="8BBA36F1C329FFC89F35FCF39711BDC0" blockId="7.[554,1034,871,898]" box="[554,1034,871,898]" pageId="7" pageNumber="149">
<heading id="D0F2819DC329FFC89F35FCF39711BDC0" box="[554,1034,871,898]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="149" reason="2">
Genus
<taxonomicName id="4C054D72C329FFC89F63FCFC9711BDC0" ID-CoL="7P6XG" authority="Godwin-Austen, 1918" authorityName="Godwin-Austen" authorityYear="1918" box="[636,1034,871,898]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="genus">
<emphasis id="B971EAE3C329FFC89F63FCFC91E6BDC0" bold="true" box="[636,765,871,898]" italics="true" pageId="7" pageNumber="149">Landouria</emphasis>
Godwin-Austen, 1918
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</heading>
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</subSubSection>
<subSubSection id="C31F657AC329FFC89DA2FC329257BA57" pageId="7" pageNumber="149" type="type_taxon">
<paragraph id="8BBA36F1C329FFC89DA2FC329243BD81" blockId="7.[189,344,937,963]" box="[189,344,937,963]" pageId="7" pageNumber="149">
<heading id="D0F2819DC329FFC89DA2FC329243BD81" bold="true" box="[189,344,937,963]" fontSize="11" level="3" pageId="7" pageNumber="149" reason="3">
<emphasis id="B971EAE3C329FFC89DA2FC329243BD81" bold="true" box="[189,344,937,963]" pageId="7" pageNumber="149">
<typeStatus id="54BE8853C329FFC89DA2FC3293E1BD81" box="[189,250,937,963]" pageId="7" pageNumber="149">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBA36F1C329FFC89DA2FC439257BA57" blockId="7.[189,1398,984,1045]" pageId="7" pageNumber="149">
<taxonomicName id="4C054D72C329FFC89DA2FC43912DBDB0" authority="L. Pfeiffer, 1842" authorityName="L. Pfeiffer" authorityYear="1842" box="[189,566,984,1010]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="species" species="huttonii">
<emphasis id="B971EAE3C329FFC89DA2FC439279BDB0" box="[189,354,984,1010]" italics="true" pageId="7" pageNumber="149">Helix huttonii</emphasis>
L. Pfeiffer, 1842
</taxonomicName>
(new name for
<taxonomicName id="4C054D72C329FFC89FE2FC4397C0BDB0" authority="Hutton &amp; Benson, 1838" authorityName="Hutton &amp; Benson" authorityYear="1838" box="[765,1243,984,1010]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="species" species="orbicula">
<emphasis id="B971EAE3C329FFC89FE2FC4390B1BDB0" box="[765,938,984,1010]" italics="true" pageId="7" pageNumber="149">Helix orbicula</emphasis>
<bibRefCitation id="EF944B00C329FFC89EAAFC4397C0BDB0" author="Hutton T. &amp; Benson W. H." box="[949,1243,984,1010]" pageId="7" pageNumber="149" pagination="211 - 218" refId="ref10346" refString="Hutton T. &amp; Benson W. H. 1838. On the land and fresh-water shells of the western Himalaya. Journal of the Asiatic Society of Bengal 7: 211 - 218." type="journal article" year="1838">Hutton &amp; Benson, 1838</bibRefCitation>
</taxonomicName>
), by original designation.
</paragraph>
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<paragraph id="8BBA36F1C329FFC89DA2FBA69246BA14" blockId="7.[189,349,1084,1111]" box="[189,349,1084,1111]" pageId="7" pageNumber="149">
<heading id="D0F2819DC329FFC89DA2FBA69246BA14" bold="true" box="[189,349,1084,1111]" fontSize="11" level="3" pageId="7" pageNumber="149" reason="3">
<emphasis id="B971EAE3C329FFC89DA2FBA69246BA14" bold="true" box="[189,349,1084,1111]" pageId="7" pageNumber="149">
<typeStatus id="54BE8853C329FFC89DA2FBA693E1BA15" box="[189,250,1085,1111]" pageId="7" pageNumber="149">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBA36F1C329FFC89DA2FBF09712BAC7" blockId="7.[189,1033,1131,1157]" box="[189,1033,1131,1157]" pageId="7" pageNumber="149">
Himalaya near Simla, Mahasu, northern
<collectingCountry id="F3127661C329FFC89F85FBF091CDBAC7" box="[666,726,1131,1157]" name="India" pageId="7" pageNumber="149">India</collectingCountry>
(
<bibRefCitation id="EF944B00C329FFC89FFAFBF090E7BAC7" author="Hutton T. &amp; Benson W. H." box="[741,1020,1131,1157]" pageId="7" pageNumber="149" pagination="211 - 218" refId="ref10346" refString="Hutton T. &amp; Benson W. H. 1838. On the land and fresh-water shells of the western Himalaya. Journal of the Asiatic Society of Bengal 7: 211 - 218." type="journal article" year="1838">Hutton &amp; Benson 1838</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31F657AC329FFC89DA2FB379747BBA4" pageId="7" pageNumber="149" type="distribution">
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<emphasis id="B971EAE3C329FFC89DA2FB37924DBA84" bold="true" box="[189,342,1196,1222]" pageId="7" pageNumber="149">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBA36F1C329FFC89DA2FB409250BB7E" blockId="7.[188,1399,1243,1340]" pageId="7" pageNumber="149">
The genus is also reported from northeast
<collectingCountry id="F3127661C329FFC89FBCFB4091F9BAB7" box="[675,738,1243,1269]" name="India" pageId="7" pageNumber="149">India</collectingCountry>
, west Himalaya,
<collectingCountry id="F3127661C329FFC89EB7FB4090EABAB7" box="[936,1009,1243,1269]" name="Nepal" pageId="7" pageNumber="149">Nepal</collectingCountry>
,
<collectingRegion id="49C1F813C329FFC89EE4FB40976BBAB7" box="[1019,1136,1243,1269]" country="Myanmar" name="Mandalay" pageId="7" pageNumber="149">Mandalay</collectingRegion>
and
<collectingRegion id="49C1F813C329FFC899B8FB4097F9BAB7" box="[1191,1250,1243,1269]" country="Myanmar" name="Shan" pageId="7" pageNumber="149">Shan</collectingRegion>
in
<collectingCountry id="F3127661C329FFC8981BFB40966DBAB7" box="[1284,1398,1243,1269]" name="Myanmar" pageId="7" pageNumber="149">Myanmar</collectingCountry>
(
<collectingCountry id="F3127661C329FFC89DDBFB659201BB5A" box="[196,282,1278,1304]" name="Myanmar" pageId="7" pageNumber="149">Burma</collectingCountry>
),
<collectingRegion id="49C1F813C329FFC89C32FB64929CBB5B" box="[301,391,1279,1305]" country="China" name="Yunnan" pageId="7" pageNumber="149">Yunnan</collectingRegion>
in
<collectingCountry id="F3127661C329FFC89CB4FB6592EABB5A" box="[427,497,1278,1304]" name="China" pageId="7" pageNumber="149">China</collectingCountry>
(
<bibRefCitation id="EF944B00C329FFC89F1EFB65919DBB5A" author="Zilch A." box="[513,646,1278,1304]" pageId="7" pageNumber="149" pagination="293 - 319" refId="ref11455" refString="Zilch A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae (5). Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch 1966</bibRefCitation>
;
<bibRefCitation id="EF944B00C329FFC89F8DFB6590F4BB5A" author="Schileyko A. A. &amp; Kuznetsov A. G." box="[658,1007,1278,1305]" pageId="7" pageNumber="149" pagination="43 - 54" refId="ref11134" refString="Schileyko A. A. &amp; Kuznetsov A. G. 1998. Land snails of the genus Landouria Godwin-Austen, 1918 and some other Bradybaenidae of Nepal (Gastropoda, Pulmonata). Ruthenica 8: 43 - 54." type="journal article" year="1998">Schileyko &amp; Kuznetsov 1998</bibRefCitation>
;
<bibRefCitation id="EF944B00C329FFC89EE4FB659605BB5A" author="Ramakrishna &amp; Mitra S. C. &amp; Dey A." box="[1019,1310,1278,1304]" pageId="7" pageNumber="149" pagination="1 - 359" refId="ref10904" refString="Ramakrishna, Mitra S. C. &amp; Dey A. 2010. Annotated checklist of Indian land molluscs. Zoological Survey of India 306: 1 - 359." type="journal article" year="2010">
Ramakrishna
<emphasis id="B971EAE3C329FFC89981FB6497C2BB5A" box="[1182,1241,1278,1304]" italics="true" pageId="7" pageNumber="149">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EF944B00C329FFC89835FB659224BB7E" author="Budha P. B. &amp; Naggs F. &amp; Backeljau T." pageId="7" pageNumber="149" pagination="1 - 48" refId="ref9911" refString="Budha P. B., Naggs F. &amp; Backeljau T. 2015. Annotated checklist of the terrestrial gastropods of Nepal. ZooKeys 492: 1 - 48. https: // doi. org / 10.3897 / zookeys. 492.9175" type="journal article" year="2015">
Budha
<emphasis id="B971EAE3C329FFC89DA2FAB993E3BB79" box="[189,248,1313,1339]" italics="true" pageId="7" pageNumber="149">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
<paragraph id="8BBA36F1C329FFC89DA2FAF89747BBA4" blockId="7.[188,1398,1379,1511]" pageId="7" pageNumber="149">
<bibRefCitation id="EF944B00C329FFC89DA2FAF89137BB3F" author="Schileyko A. A. &amp; Kuznetsov A. G." box="[189,556,1379,1405]" pageId="7" pageNumber="149" pagination="43 - 54" refId="ref11134" refString="Schileyko A. A. &amp; Kuznetsov A. G. 1998. Land snails of the genus Landouria Godwin-Austen, 1918 and some other Bradybaenidae of Nepal (Gastropoda, Pulmonata). Ruthenica 8: 43 - 54." type="journal article" year="1998">Schileyko &amp; Kuznetsov (1998)</bibRefCitation>
tentatively argued that
<taxonomicName id="4C054D72C329FFC89E5EFAF890A6BB3F" authorityName="Godwin-Austen" authorityYear="1918" box="[833,957,1379,1405]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="genus">
<emphasis id="B971EAE3C329FFC89E5EFAF890A6BB3F" box="[833,957,1379,1405]" italics="true" pageId="7" pageNumber="149">Landouria</emphasis>
</taxonomicName>
is restricted to northern
<collectingCountry id="F3127661C329FFC899C1FAF89606BB3F" box="[1246,1309,1379,1405]" name="India" pageId="7" pageNumber="149">India</collectingCountry>
,
<collectingCountry id="F3127661C329FFC89836FAF89669BB3F" box="[1321,1394,1379,1405]" name="Nepal" pageId="7" pageNumber="149">Nepal</collectingCountry>
, northern
<collectingCountry id="F3127661C329FFC89C36FA1D9286BBE2" box="[297,413,1414,1440]" name="Myanmar" pageId="7" pageNumber="149">Myanmar</collectingCountry>
, and
<collectingCountry id="F3127661C329FFC89CC2FA1D914FBBE2" box="[477,596,1414,1440]" name="Sri Lanka" pageId="7" pageNumber="149">Sri Lanka</collectingCountry>
. Yet, recently, species of
<taxonomicName id="4C054D72C329FFC89E9DFA1D90E5BBE2" authorityName="Godwin-Austen" authorityYear="1918" box="[898,1022,1414,1440]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="149" phylum="Mollusca" rank="genus">
<emphasis id="B971EAE3C329FFC89E9DFA1D90E5BBE2" box="[898,1022,1414,1440]" italics="true" pageId="7" pageNumber="149">Landouria</emphasis>
</taxonomicName>
have been reported from
<collectingCountry id="F3127661C329FFC8982FFA1D966DBBE2" box="[1328,1398,1414,1440]" name="China" pageId="7" pageNumber="149">China</collectingCountry>
(
<bibRefCitation id="EF944B00C329FFC89DD9FA3292FBBB81" author="Pall-Gergely B. &amp; Hunyadi A. &amp; Asami T." box="[198,480,1449,1475]" pageId="7" pageNumber="149" pagination="130 - 134" refId="ref10714" refString="Pall-Gergely B., Hunyadi A. &amp; Asami T. 2013. A peculiar new species in the genus Landouria Godwin- Austen, 1918 from China (Gastropoda: Heterobranchia: Stylommatophora: Camaenidae). Molluscan Research 33: 130 - 134. https: // doi. org / 10.1080 / 13235818.2013.782794" type="journal article" year="2013">
Páll-Gergely
<emphasis id="B971EAE3C329FFC89C7DFA319287BB81" box="[354,412,1449,1475]" italics="true" pageId="7" pageNumber="149">et al.</emphasis>
2013
</bibRefCitation>
),
<collectingCountry id="F3127661C329FFC89CECFA319137BB86" box="[499,556,1450,1476]" name="Laos" pageId="7" pageNumber="149">Laos</collectingCountry>
(
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Hirano
<emphasis id="B971EAE3C329FFC89F8DFA3191D7BB81" box="[658,716,1449,1475]" italics="true" pageId="7" pageNumber="149">et al.</emphasis>
2014
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),
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(
<bibRefCitation id="EF944B00C329FFC89EDCFA329780BB81" author="Kohler F. &amp; Shea M. &amp; Kessner V." box="[963,1179,1449,1475]" pageId="7" pageNumber="149" pagination="253 - 264" refId="ref10381" refString="Kohler F., Shea M. &amp; Kessner V. 2018. Two new species of Landouria Godwin-Austen, 1918 from Timor-Leste (Stylommatophora, Camaenidae). Molluscan Research 39 (3): 253 - 264. https: // doi. org / 10.1080 / 13235818.2018.1557780" type="journal article" year="2018">
Köhler
<emphasis id="B971EAE3C329FFC89904FA31974FBB81" box="[1051,1108,1449,1475]" italics="true" pageId="7" pageNumber="149">et al.</emphasis>
2018
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),
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in
<collectingCountry id="F3127661C329FFC8981BFA32966DBB81" box="[1284,1398,1449,1475]" name="Indonesia" pageId="7" pageNumber="149">Indonesia</collectingCountry>
(
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Nurinsiyah
<emphasis id="B971EAE3C329FFC89C4FFA569290BBA4" box="[336,395,1484,1510]" italics="true" pageId="7" pageNumber="149">et al.</emphasis>
2019
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), and
<collectingCountry id="F3127661C329FFC89F08FA579164BBA4" box="[535,639,1484,1510]" name="Thailand" pageId="7" pageNumber="149">Thailand</collectingCountry>
(
<bibRefCitation id="EF944B00C329FFC89F91FA57974BBBA4" author="Tumpeesuwan C. &amp; Tumpeesuwan S." box="[654,1104,1484,1511]" pageId="7" pageNumber="149" pagination="298 - 305" refId="ref11349" refString="Tumpeesuwan C. &amp; Tumpeesuwan S. 2019. First verified record of the genus Landouria Godwin- Austen, 1918 from Thailand (Gastropoda: Stylommatophora: Camaenidae) with description of a new species. Raffles Bulletin of Zoology 67: 298 - 305. https: // doi. org / 10.26107 / RBZ- 2019 - 0021" type="journal article" year="2019">Tumpeesuwan &amp; Tumpeesuwan 2019</bibRefCitation>
).
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