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<document id="91D8905FF99645A0445C855C39B0A0DA" ID-CLB-Dataset="30136" ID-DOI="10.11646/zootaxa.4416.1.1" ID-GBIF-Dataset="aa238a39-adf8-4b94-8808-c788670d0455" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242263" ID-ZooBank="3341A80B-7E54-4C6B-8B46-1336094952E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525601053009" checkinUser="plazi" docAuthor="Barbosa, Diego N. &amp; Azevedo, Celso O." docDate="2018" docId="039D87C45C57FFF348F6FC70FDE8F86E" docLanguage="en" docName="zootaxa.4416.1.1.pdf" docOrigin="Zootaxa 4416 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anisepyris ecthelion Barbosa &amp; Azevedo 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="73" masterDocId="FFA4FFBC5C1FFFBB4861FFDFFFCEFFFD" masterDocTitle="Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species" masterLastPageNumber="258" masterPageNumber="1" pageNumber="73" updateTime="1698542699957" updateUser="ExternalLinkService">
<mods:mods id="7D8FBE324CE28C8695071A24F3E8F992" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BEE740A73E5EEA0693E548BD8B2C6248">
<mods:title id="ADB4E63EBC2F54A247E4F2F5AB0DD75C">Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species</mods:title>
</mods:titleInfo>
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<mods:role id="8FAF938BF8BC0A44B903E5581C2AC971">
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<mods:namePart id="28679DC276C9D361601A8B87FDEFD6F5">Barbosa, Diego N.</mods:namePart>
</mods:name>
<mods:name id="F726043D0B8C7D82750A2527C14DCDFC" type="personal">
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<mods:namePart id="38169E7B224B528C100F839A6D8B6F18">Azevedo, Celso O.</mods:namePart>
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<mods:typeOfResource id="A303CF2709A540936FA41648656D5CB7">text</mods:typeOfResource>
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<mods:title id="223B901CDA116A5122940F413AEAAD5C">Zootaxa</mods:title>
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<mods:part id="927072C1F738FCDE8A61ADF162476CB6">
<mods:date id="337A0179F12C46BDE24B946849E991AC">2018</mods:date>
<mods:detail id="1038EA699DB57E82F59F6D479898F557" type="volume">
<mods:number id="63A97354884F76E036E84E8979B59C1F">4416</mods:number>
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<mods:classification id="B09F9206A235796BF5ADED9BBFA7D187">journal article</mods:classification>
<mods:identifier id="7E9156EDD4E23E35CEF6BE34ACA63F0F" type="CLB-Dataset">30136</mods:identifier>
<mods:identifier id="767A8E70A39C8A462DC2FD0B8943427E" type="DOI">10.11646/zootaxa.4416.1.1</mods:identifier>
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<mods:identifier id="5B7167424E4AC53195734A72332971A7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039D87C45C57FFF348F6FC70FDE8F86E" ID-DOI="http://doi.org/10.5281/zenodo.5961275" ID-GBIF-Taxon="143811702" ID-Zenodo-Dep="5961275" LSID="urn:lsid:plazi:treatment:039D87C45C57FFF348F6FC70FDE8F86E" httpUri="http://treatment.plazi.org/id/039D87C45C57FFF348F6FC70FDE8F86E" lastPageNumber="73" pageId="72" pageNumber="73">
<subSubSection id="C32E65595C57FFF348F6FC70FE03FC16" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="8B8B36D25C57FFF348F6FC70FD36FC37" blockId="72.[151,760,943,1003]" box="[151,760,943,970]" pageId="72" pageNumber="73">
<heading id="D0C381BE5C57FFF348F6FC70FD36FC37" bold="true" box="[151,760,943,970]" fontSize="11" level="1" pageId="72" pageNumber="73" reason="1">
<emphasis id="B940EAC05C57FFF348F6FC70FD36FC37" bold="true" box="[151,760,943,970]" pageId="72" pageNumber="73">
<taxonomicName id="4C344D515C57FFF348F6FC70FD40FC37" authority="Barbosa &amp; Azevedo" authorityName="Barbosa &amp; Azevedo" authorityYear="2018" box="[151,654,943,970]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="ecthelion" status="sp. nov.">
<emphasis id="B940EAC05C57FFF348F6FC70FE42FC34" bold="true" box="[151,396,943,969]" italics="true" pageId="72" pageNumber="73">Anisepyris ecthelion</emphasis>
Barbosa &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A27357BB5C57FFF34AFAFC6FFD36FC37" box="[667,760,944,970]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36D25C57FFF348F6FC0DFE03FC16" blockId="72.[151,760,943,1003]" box="[151,461,978,1003]" pageId="72" pageNumber="73">
(
<figureCitation id="130F2A575C57FFF348FEFC0DFEDEFC16" box="[159,272,978,1003]" captionStart="FIGURE 10" captionStartId="67.[151,250,1918,1940]" captionTargetBox="[233,1351,233,1853]" captionTargetId="figure@67.[192,1395,195,1896]" captionTargetPageId="67" captionText="FIGURE 10. Anisepyris: bifidus species-group.A–D.Head in dorsal profile.A. A.breviramus Santos &amp; Azevedo, 2000; B. A. divisus Santos, 2002; C. A. ramosus Santos &amp; Azevedo, 2008; D. A. trinitatis Evans, 1966; E–F. Female mandible in frontal view; G.Mesosomain dorsal profile; H–I. Mesopleuron in frontal view; bogotensis species-group.J–M.Head in dorsal profile. J. A.bogotensis (Kieffer, 1910); K. A. borlachi sp. nov.; L. A.denethori sp. nov.; M.A. ecthelion sp. nov." httpUri="https://zenodo.org/record/1242283/files/figure.png" pageId="72" pageNumber="73">Figs 10M</figureCitation>
; 11A; 11C; 11D)
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C57FFF348F6FBC5FAD6F9FA" pageId="72" pageNumber="73" type="description">
<paragraph id="8B8B36D25C57FFF348F6FBC5FAD6F9FA" blockId="72.[151,1437,1050,1939]" pageId="72" pageNumber="73">
<emphasis id="B940EAC05C57FFF348F6FBC5FE42FBCE" bold="true" box="[151,396,1050,1075]" pageId="72" pageNumber="73">Description, female.</emphasis>
Body length
<quantity id="4CCC9B375C57FFF34A59FBC4FD6BFBCF" box="[568,677,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="4.77" pageId="72" pageNumber="73" unit="mm" value="4.77">4.77 mm</quantity>
. Head and mesosoma black; wings subhyaline; metasoma dark castaneous. Head wide. Mandible with five distal teeth, narrow, curved, and with teeth equally wide. Median clypeal lobe angulate, very short. Frons
<taxonomicName id="4C344D515C57FFF34A00FBBDFD13FB86" authorityName="Kieffer" authorityYear="1908" box="[609,733,1122,1147]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="coriaceous">coriaceous</taxonomicName>
; antennal scrobe ecarinate. Eye large. Frontal angle of ocellar triangle acute; ocelli large. Vertex straight. Pronotal disc as long as wide, strongly
<taxonomicName id="4C344D515C57FFF34C5FFB59FB77FB62" authorityName="Kieffer" authorityYear="1908" box="[1086,1209,1158,1183]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="coriaceous">coriaceous</taxonomicName>
; transverse pronotal carina absent; posterior margin without posterior pronotal sulcus. Notaulus wide. Mesoscutellar disc with mesoscutellar sulcus narrow. Metapectal-propodeal disc long; metapostnotal median carina complete; longitudinal ridges absent; metapostnotal-propodeal suture fused with first metapostnotal carina; paraspiracular carina present and inconspicuous; metapleural carina present and conspicuous; metapectal-propodeal disc with short striae; propodeal declivity strigate. Mesopleuron with anterior fovea closed; mesopleural fovea closed; lower fovea closed; posterior fovea absent; mesopleural suture closed. Forewing with stigma developed; Rs&amp;M vein distinguished. Mesotibia not spinose; claws bifid. Metasomal segments sparsely setose.
<emphasis id="B940EAC05C57FFF34C0AFA5DFB2FFA66" bold="true" box="[1131,1249,1410,1435]" pageId="72" pageNumber="73">Genitalia.</emphasis>
T9 with anterior arm long and overlapping 2vf; 1vv as long as genitalia body; 2vv as long as 1vv length, smoothly narrowing apicad; 3vv short; 1vf strongly curved forward; 2vf slender and with dorsal lap strongly swollen posteriorly; 1rv slightly curved forward, ventral process large, and posterior process short; 2rv slightly curved forward.
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C57FFF348A6F9CDFEBEF96A" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="8B8B36D25C57FFF348A6F9CDFEBEF96A" blockId="72.[151,1437,1050,1939]" pageId="72" pageNumber="73">
<materialsCitation id="3B5C3C8F5C57FFF348A6F9CDFD27F9D6" ID-GBIF-Occurrence="1841141907" box="[199,745,1554,1579]" country="Paraguay" location="Material" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B940EAC05C57FFF348A6F9CDFE7FF9D6" bold="true" box="[199,433,1554,1579]" pageId="72" pageNumber="73">
<location id="8EEB60095C57FFF348A6F9CDFEE1F9D6" LSID="urn:lsid:plazi:treatment:039D87C45C57FFF348F6FC70FDE8F86E:8EEB60095C57FFF348A6F9CDFEE1F9D6" box="[199,303,1554,1579]" country="Paraguay" name="Material" pageId="72" pageNumber="73">Material</location>
examined.
</emphasis>
<typeStatus id="548F88705C57FFF349D9F9CDFDEDF9D6" box="[440,547,1554,1579]" pageId="72" pageNumber="73" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D32FD5B5C57FFF34A4FF9CCFD8DF9D7" box="[558,579,1555,1578]" pageId="72" pageNumber="73" type="female">♀</specimenCount>
,
<collectingCountry id="F32376425C57FFF34A33F9CCFD27F9D6" box="[594,745,1555,1579]" name="Paraguay" pageId="72" pageNumber="73">PARAGUAY</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B5C3C8F5C57FFF34A92F9CBFB45F9B2" ID-GBIF-Occurrence="1841141940" collectingDate="2000-11-21" collectingDateMax="2000-11-25" collectingDateMin="2000-11-21" collectionCode="FIT, CNCI" collectorName="Z. H. Falin" country="Paraguay" elevation="100" latitude="-26.638056" location="San Rafael Res" longLatPrecision="20" longitude="-55.66389" municipality="Yatai Hostettler" pageId="72" pageNumber="73" specimenCount="1" stateProvince="Itapua" typeStatus="paratype">
<emphasis id="B940EAC05C57FFF34A92F9CBFC8DF9D6" box="[755,835,1556,1579]" italics="true" pageId="72" pageNumber="73">
<collectingRegion id="49F0F8305C57FFF34A92F9CBFCF1F9D6" box="[755,831,1556,1579]" country="Paraguay" name="Itapua" pageId="72" pageNumber="73">Itapua</collectingRegion>
,
</emphasis>
<collectingMunicipality id="6BEFACA85C57FFF34B2AF9CDFC34F9D6" box="[843,1018,1554,1579]" pageId="72" pageNumber="73">Yatai Hostettler</collectingMunicipality>
family,
<location id="8EEB60095C57FFF34C39F9CCFAC9F9D7" LSID="urn:lsid:plazi:treatment:039D87C45C57FFF348F6FC70FDE8F86E:8EEB60095C57FFF34C39F9CCFAC9F9D7" box="[1112,1287,1554,1579]" country="Paraguay" latitude="-26.638056" longLatPrecision="20" longitude="-55.66389" municipality="Yatai Hostettler" name="San Rafael Res" pageId="72" pageNumber="73" stateProvince="Itapua">San Rafael Res</location>
[erve],
<elevation id="0019D1E15C57FFF34D34F9CCFA56F9D6" box="[1365,1432,1555,1579]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="m" value="100.0">
<quantity id="4CCC9B375C57FFF34D34F9CCFA56F9D6" box="[1365,1432,1555,1579]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="m" value="100.0">100m</quantity>
</elevation>
,
<geoCoordinate id="EE0050155C57FFF348F6F9E9FED9F9B2" box="[151,279,1590,1615]" direction="south" orientation="latitude" pageId="72" pageNumber="73" precision="15" value="-26.638056">26°38'17”S</geoCoordinate>
<geoCoordinate id="EE0050155C57FFF3497FF9E9FE66F9B2" box="[286,424,1590,1615]" direction="west" orientation="longitude" pageId="72" pageNumber="73" precision="15" value="-55.66389">55°39'50&quot;W</geoCoordinate>
,
<collectingDate id="EFCEE9FA5C57FFF349D5F9E9FD96F9B2" box="[436,600,1590,1615]" pageId="72" pageNumber="73" value="2000-11-21" valueMax="2000-11-25" valueMin="2000-11-21">
<date id="FF8A10125C57FFF349D5F9E9FD96F9B2" box="[436,600,1590,1615]" pageId="72" pageNumber="73" value="2000-11-21" valueMax="2000-11-25" valueMin="2000-11-21">21–25.xi.2000</date>
</collectingDate>
,
<collectionCode id="ED25AE175C57FFF34A02F9E8FD40F9B3" box="[611,654,1591,1614]" pageId="72" pageNumber="73">FIT</collectionCode>
,
<collectorName id="26C153045C57FFF34AF8F9E8FCC0F9B2" box="[665,782,1590,1615]" pageId="72" pageNumber="73">Z.H. Falin</collectorName>
[collector] (
<collectionCode id="ED25AE175C57FFF34BF9F9E8FC2FF9B2" box="[920,993,1591,1615]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="72" pageNumber="73">CNCI</collectionCode>
).
<typeStatus id="548F88705C57FFF34B95F9E8FBAFF9B2" box="[1012,1121,1591,1615]" pageId="72" pageNumber="73" type="paratype">Paratypes</typeStatus>
(4)
</materialsCitation>
.
<materialsCitation id="3B5C3C8F5C57FFF34CF7F9E8FEBEF96A" ID-GBIF-Occurrence="1841141922" collectingDate="2000-11-21" collectingDateMax="2000-11-25" collectingDateMin="2000-11-21" collectionCode="FIT, CNCI" collectorName="Z. H. Falin" country="Paraguay" elevation="100" latitude="-26.638056" location="San Rafael Res" longLatPrecision="20" longitude="-55.66389" municipality="Yatai Hostettler" pageId="72" pageNumber="73" specimenCount="4" specimenCount-female="4" stateProvince="Itapua" typeStatus="paratype">
<collectingCountry id="F32376425C57FFF34CF7F9E8FAE2F9B2" box="[1174,1324,1591,1615]" name="Paraguay" pageId="72" pageNumber="73">PARAGUAY</collectingCountry>
,
<collectingRegion id="49F0F8305C57FFF34D56F9E8FA4EF9B3" box="[1335,1408,1591,1614]" country="Paraguay" name="Itapua" pageId="72" pageNumber="73">
<emphasis id="B940EAC05C57FFF34D56F9E8FA4EF9B3" box="[1335,1408,1591,1614]" italics="true" pageId="72" pageNumber="73">Itapua</emphasis>
</collectingRegion>
,
<specimenCount id="9D32FD5B5C57FFF34DECF9E8FF62F98F" pageId="72" pageNumber="73" type="female">4 ♀</specimenCount>
,
<collectingMunicipality id="6BEFACA85C57FFF348DDF985FEA2F98E" box="[188,364,1626,1651]" pageId="72" pageNumber="73">Yatai Hostettler</collectingMunicipality>
family,
<location id="8EEB60095C57FFF349ACF984FDB0F98F" LSID="urn:lsid:plazi:treatment:039D87C45C57FFF348F6FC70FDE8F86E:8EEB60095C57FFF349ACF984FDB0F98F" box="[461,638,1626,1651]" country="Paraguay" latitude="-26.638056" longLatPrecision="20" longitude="-55.66389" municipality="Yatai Hostettler" name="San Rafael Res" pageId="72" pageNumber="73" stateProvince="Itapua">San Rafael Res</location>
[erve],
<elevation id="0019D1E15C57FFF34AAFF984FCDFF98E" box="[718,785,1627,1651]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="m" value="100.0">
<quantity id="4CCC9B375C57FFF34AAFF984FCDFF98E" box="[718,785,1627,1651]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="m" value="100.0">100m</quantity>
</elevation>
,
<geoCoordinate id="EE0050155C57FFF34B7FF985FC53F98E" box="[798,925,1626,1651]" direction="south" orientation="latitude" pageId="72" pageNumber="73" precision="15" value="-26.638056">26°38'17&quot;S</geoCoordinate>
<geoCoordinate id="EE0050155C57FFF34BC7F985FBE1F98E" box="[934,1071,1626,1651]" direction="west" orientation="longitude" pageId="72" pageNumber="73" precision="15" value="-55.66389">55°39'50&quot;W</geoCoordinate>
,
<collectingDate id="EFCEE9FA5C57FFF34C5CF985FB2CF98E" box="[1085,1250,1626,1651]" pageId="72" pageNumber="73" value="2000-11-21" valueMax="2000-11-25" valueMin="2000-11-21">
<date id="FF8A10125C57FFF34C5CF985FB2CF98E" box="[1085,1250,1626,1651]" pageId="72" pageNumber="73" value="2000-11-21" valueMax="2000-11-25" valueMin="2000-11-21">21–25.xi.2000</date>
</collectingDate>
,
<collectionCode id="ED25AE175C57FFF34C8EF984FAD4F98F" box="[1263,1306,1627,1650]" pageId="72" pageNumber="73">FIT</collectionCode>
,
<collectorName id="26C153045C57FFF34D47F984FA53F98E" box="[1318,1437,1626,1651]" pageId="72" pageNumber="73">Z.H. Falin</collectorName>
[collector] (
<collectionCode id="ED25AE175C57FFF3497BF9A0FEADF96A" box="[282,355,1663,1687]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="72" pageNumber="73">CNCI</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C57FFF348A6F97DFE18F946" box="[199,470,1698,1723]" pageId="72" pageNumber="73" type="distribution">
<paragraph id="8B8B36D25C57FFF348A6F97DFE18F946" blockId="72.[151,1437,1050,1939]" box="[199,470,1698,1723]" pageId="72" pageNumber="73">
<emphasis id="B940EAC05C57FFF348A6F97DFEAEF946" bold="true" box="[199,352,1698,1723]" pageId="72" pageNumber="73">Distribution.</emphasis>
<collectingRegion id="49F0F8305C57FFF34906F97CFE1CF946" box="[359,466,1699,1723]" country="Paraguay" name="Asuncion" pageId="72" pageNumber="73">
<collectingCountry id="F32376425C57FFF34906F97CFE1CF946" box="[359,466,1699,1723]" name="Paraguay" pageId="72" pageNumber="73">Paraguay</collectingCountry>
</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C57FFF348A6F919FDE8F86E" pageId="72" pageNumber="73" type="discussion">
<paragraph id="8B8B36D25C57FFF348A6F919FDE8F86E" blockId="72.[151,1437,1050,1939]" pageId="72" pageNumber="73">
<emphasis id="B940EAC05C57FFF348A6F919FEF5F922" bold="true" box="[199,315,1734,1759]" pageId="72" pageNumber="73">Remarks.</emphasis>
This new species differs from
<taxonomicName id="4C344D515C57FFF34AFFF918FC24F922" authority="Kieffer, 1908" authorityName="Kieffer" authorityYear="1908" box="[670,1002,1734,1759]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rufosignatus">
<emphasis id="B940EAC05C57FFF34AFFF918FC84F922" box="[670,842,1735,1759]" italics="true" pageId="72" pageNumber="73">A. rufosignatus</emphasis>
Kieffer, 1908
</taxonomicName>
by having the clypeus with the apical margin rounded; the antennal scrobe ecarinate; the frontal angle of the ocellae forming a right angle; the pronotal disc without a transverse pronotal carina; and the mesopleuron with the lower fovea closed and not divided, whereas
<taxonomicName id="4C344D515C57FFF3489AF8ECFE69F8B6" authorityName="Kieffer" authorityYear="1908" box="[251,423,1843,1867]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rufosignatus">
<emphasis id="B940EAC05C57FFF3489AF8ECFE69F8B6" box="[251,423,1843,1867]" italics="true" pageId="72" pageNumber="73">A. rufosignatus</emphasis>
</taxonomicName>
has the clypeus with the apical margin angulate; the antennal scrobe carinate; the frontal angle of the ocellar triangle obtuse; the pronotal disc with a transverse pronotal carina; and the mesopleuron with the lower fovea closed and divided.
</paragraph>
</subSubSection>
</treatment>
</document>
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