<documentid="DB71E605D91948377BF39EAB2041ECDD"ID-CLB-Dataset="48132"ID-DOI="10.5281/zenodo.207073"ID-GBIF-Dataset="a3815507-6100-464e-aaf6-60d6820a1eec"ID-ISSN="1175-5326"ID-Zenodo-Dep="207073"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="existingObjects,plazi"IM.taxonomicNames_approvedBy="felipe"checkinTime="1460228776049"checkinUser="plazi"docAuthor="Torres-Carvajal, Omar, Etheridge, Richard & Queiroz, Kevin De"docDate="2011"docId="038287FDFF9B606669838DD1D44BF84F"docLanguage="en"docName="zt02752p044.pdf"docOrigin="Zootaxa 2752"docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article"docStyleId="890A69B780ED73D6DB8551B71C8AC79E"docStyleName="Zootaxa.2009-2012.journal_article"docStyleVersion="4"docTitle="Morunasaurus annularis O'Shaughnessy 1881"docType="treatment"docVersion="11"lastPageNumber="34"masterDocId="FFBBFF85FFBB604769148B6ED174FF93"masterDocTitle="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)"masterLastPageNumber="44"masterPageNumber="1"pageNumber="33"updateTime="1720202091178"updateUser="ExternalLinkService">
<mods:titleid="D1644CAD776E2E93F1764FC5F915C107">A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)</mods:title>
<bibRefCitationid="EFBA4B1AFF9B606768C38DD1D3A9F94A"author="O'Shaughnessy"box="[471,733,1727,1753]"pageId="32"pageNumber="40"refString="O'Shaughnessy, A. W. E. (1881) An account of the collection of lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245."type="proceedings paper"year="1881">O'Shaughnessy 1881</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraphid="8B9436EBFF9B606769838C68D3D1F88F"blockId="32.[151,713,1798,1851]"box="[151,677,1798,1820]"pageId="32"pageNumber="33">Proposed standard English name: ringed manticores</paragraph>
<paragraphid="8B9436EBFF9B606769838C4BD3BDF8A8"blockId="32.[151,713,1798,1851]"box="[151,713,1829,1851]"pageId="32"pageNumber="33">Proposed standard Spanish name: mantícoras de anillos</paragraph>
<bibRefCitationid="EFBA4B1AFF9B606768938C33D31FF8E7"author="O'Shaughnessy"box="[391,619,1885,1908]"pageId="32"pageNumber="40"refString="O'Shaughnessy, A. W. E. (1881) An account of the collection of lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245."type="proceedings paper"year="1881">O’Shaughnessy (1881</bibRefCitation>
<bibRefCitationid="EFBA4B1AFF9B60676BCF8C15D2CAF802"author="Boulenger"box="[731,958,1915,1937]"pageId="32"pageNumber="38"refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History) II. Taylor and Francis, London, 497 pp."type="book"year="1885">Boulenger (1885:200)</bibRefCitation>
;
<bibRefCitationid="EFBA4B1AFF9B60676ADC8C13D5C4F802"author="Burt"box="[968,1200,1915,1938]"pageId="32"pageNumber="38"refString="Burt, C. E. & Burt, M. D. (1933) A preliminary check list of the lizards of South America. Transactions of the Academy of Sciences of St Louis, 28, 1 - 104."type="journal article"year="1933">Burt & Burt (1933:26)</bibRefCitation>
<bibRefCitationid="EFBA4B1AFF9B6067688E8CF5D34BF823"author="Dunn"box="[410,575,1946,1968]"pageId="32"pageNumber="39"refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79."type="journal article"year="1933">Dunn (1933:76)</bibRefCitation>
</taxonomicName>
;
<bibRefCitationid="EFBA4B1AFF9B60676B5D8CF5D2C9F822"author="Peters"box="[585,957,1947,1969]"pageId="32"pageNumber="40"refString="Peters, J. A. & Donoso-Barros, R. (1970) Catalogue of the neotropical Squamata: Part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293."type="journal article"year="1970">Peters & Donoso-Barros (1970:204)</bibRefCitation>
<bibRefCitationid="EFBA4B1AFF9A60666AF98B8ED508FF64"author="Dunn"box="[1005,1148,223,247]"pageId="33"pageNumber="39"refString="Dunn, E. R. (1933) Amphibians and reptiles from El Valle de Anton, Panama. Occasional Papers of the Boston Society of Natural History, 8, 65 - 79."type="journal article"year="1933">Dunn (1933)</bibRefCitation>
is that the tail is compressed in the former and circular (presumably in cross section) in the latter; however, we find that the tail in both species is nearly circular in cross section.
(character states in parentheses) in having usually two femoral pores on each leg (3–4); four postrostrals (five); two postmentals (four), a smaller body size (maximum SVL
(1) dorsal head scales smooth, not projecting dorsally, some occipitals granular; (2) posterior superciliaries not enlarged relative to adjacent scales; (3) scales on lateral edge of skull roof just posterior to superciliaries not projecting; (4) all pretympanic scales similar in size; (5) gular scales granular or flat and smooth, juxtaposed or slightly imbricate, not projecting ventrally; (6) dorsal neck scales heterogeneous in size, granular or large and conical; lateral neck scales granular, homogeneous in size; (7) some vertebrals between the scapular and pelvic regions larger than adjacent dorsals, forming a distinct but discontinuous middorsal longitudinal row of enlarged, elliptical, smooth, and unraised scales; (8) nuchal region without distinct middorsal longitudinal row of enlarged scales; (9) dorsals smooth and heterogeneous in size, with largest scales more projecting and abundant on posterior half of body; (10) discontinuous longitudinal row of raised, enlarged scales between dorsals and flank scales present; (11) scales on flanks heterogeneous in size, mostly granular with a few scattered enlarged scales; (12) ventrals smooth; (13) fore limb scales smooth or slightly keeled dorsally and ventrally; (14) hind limb scales smooth or slightly keeled dorsally and ventrally; scattered conical, sharply pointed, enlarged scales present dorsally; scales of pes heterogeneous in size; (15) caudals heterogeneous, with scales making up the posteriormost whorl of each segment modified as conspicuous spines (
<figureCitationid="13102A6EFF9A60666A0288E2D22EFC37"box="[790,858,908,932]"captionStart="FIGURE 3"captionStartId="12.[151,250,1517,1540]"captionTargetBox="[280,1305,321,1491]"captionTargetId="figure@12.[274,1314,306,1505]"captionTargetPageId="12"captionText="FIGURE 3. Caudal segments from the anterior third of the tail of ten species of Hoplocercinae in lateral view. (A) Enyalioides cofanorum, (B) E. heterolepis, (C) E. laticeps, (D) E. microlepis, (E) E. oshaughnessyi, (F) E. palpebralis, (G) E. praestabilis, (H) E. rubrigularis, (I) E. touzeti, and (J) Morunasaurus annularis. Scale bars = 5 mm. Illustration by D. Paucar."httpUri="https://zenodo.org/record/207076/files/figure.png"pageId="33"pageNumber="34">Fig. 3</figureCitation>
); (16) tail nearly circular in cross section. Meristic and morphometric characters are presented in
<tableCitationid="C6A90350FF9A60666B8988C1D384FC54"box="[669,752,943,967]"captionStart="TABLE 1"captionStartId="6.[151,235,1304,1325]"captionTargetBox="[151,1436,1377,1892]"captionTargetPageId="6"captionText="TABLE 1. Summary of counts and measurements (mm) for species of Hoplocercinae. Range (first line) and mean ± standard"httpUri="http://table.plazi.org/id/DF546663FFBD604169838E76D4E8FABD"pageId="33"pageNumber="34"tableUuid="DF546663FFBD604169838E76D4E8FABD">Table 1</tableCitation>
<emphasisid="B95FEAF9FF9A606669D188BDD0E4FC7F"bold="true"box="[197,400,979,1004]"pageId="33"pageNumber="34">Coloration in life</emphasis>
(
<figureCitationid="13102A6EFF9A606668B488BAD091FC7F"box="[416,485,980,1004]"captionStart="FIGURE 4"captionStartId="14.[151,250,1897,1920]"captionTargetBox="[204,1383,190,1874]"captionTargetId="figure@14.[204,1384,184,1876]"captionTargetPageId="14"captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae."httpUri="https://zenodo.org/record/207077/files/figure.png"pageId="33"pageNumber="34">Fig. 4</figureCitation>
). Adult males (QCAZ 7819): dorsum dark brown with a series of faint, cream dorsolateral blotches; sides of head yellowish brown; sides of neck orange; cream white vertical bar, bordered with dark brown anteriorly and posteriorly, extends dorsally from anterior insertion of fore limb to scapular region, where it becomes faint; flanks light yellow with a brown reticulate pattern anteriorly, and mostly light brown with scattered yellow and cream scales posteriorly; limbs light brown with scattered yellowish cream scales; chin light brown; gular region extensively covered by black mark that extends posteriorly as a wide midventral stripe that covers most of the ventral surface of body; venter light blue laterally; ventral surface of fore limbs brown with scattered yellow scales; ventral surface of hind limbs and pelvic region light grey; tail light cream ventrally and dark brown dorsally and laterally, with several grey projecting scales; iris copper brown.
Adult females: head dark brown dorsally, light brown laterally, and orange ventrally; cream stripe extends from mandibular angle to tympanum; posterior aspect of neck with a cream band bordered with dark brown, which extends dorsally over shoulder and is continuous middorsally; body cream ventrally and light brown dorsally, with five dark brown transverse bands that are wider middorsally; limbs dark brown dorsally and reddish-cream ventrally, with scattered cream flecks dorsally; tail dark brown dorsally and laterally, reddish brown ventrally (
<bibRefCitationid="EFBA4B1AFF9A60666C598EC6D066FA77"author="Kohler"pageId="33"pageNumber="40"refString="Kohler, G., Seipp, R., Moya, S. & Almendariz, A. (1999) Zur Kenntnis von Morunasaurus annularis (O'Shaughnessy, 1881). Salamandra, 35, 181 - 190."type="journal article"year="1999">
<bibRefCitationid="EFBA4B1AFF9A60666DA28E9ED4E8F99B"box="[1206,1436,1520,1544]"pageId="33"pageNumber="40"refString="Kohler, G., Seipp, R., Moya, S. & Almendariz, A. (1999) Zur Kenntnis von Morunasaurus annularis (O'Shaughnessy, 1881). Salamandra, 35, 181 - 190."type="journal article">
) within a series of nearly 20 holes in a 10-meter high wall of compact soil next to a narrow patch of primary forest surrounded by pasture. Both specimens were in holes about
<figureCitationid="13102A6EFF9A60666D348CCED505F82B"box="[1056,1137,1952,1976]"captionStart="FIGURE 11"captionStartId="34.[151,250,1664,1687]"captionTargetBox="[159,1434,217,1627]"captionTargetId="figure@34.[151,1436,195,1643]"captionTargetPageId="34"captionText="FIGURE 11. Distribution of Morunasaurus annularis (triangles), M. groi (squares), and M. peruvianus (dots)."httpUri="https://zenodo.org/record/207088/files/figure.png"pageId="33"pageNumber="34">Fig. 11</figureCitation>
). This species is known to occur in sympatry with
