180 lines
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180 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.324.5827" ID-GBIF-Dataset="752f7715-2b9f-45a4-b9ee-c759bebf3981" ID-PMC="PMC3760134" ID-Pensoft-Pub="1313-2970-324-1" ID-PubMed="24003317" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-324-1" ModsDocOrigin="ZooKeys 324" ModsDocTitle="Taxonomic revision of the olingos (Bassaricyon), with description of a new species, the Olinguito" checkinTime="1451247026135" checkinUser="pensoft" docAuthor="Helgen, Kristofer M., Pinto, C. Miguel, Kays, Roland, Helgen, Lauren E., Tsuchiya, Mirian T. N., Quinn, Aleta, Wilson, Don E. & Maldonado, Jesus E." docDate="2013" docId="CBFBACD8137B8C41D7C96AF81B5FCE17" docLanguage="en" docName="ZooKeys 324: 1-83" docOrigin="ZooKeys 324" docSource="http://dx.doi.org/10.3897/zookeys.324.5827" docTitle="Bassaricyon neblina subsp. osborni Helgen, Pinto, Kays, Helgen, Tsuchiya, Quinn, Wilson & Maldonado, 2013, subsp. n." docType="treatment" docVersion="5" lastPageNumber="21" masterDocId="FF99321FFFDBFFCCFFA0FFB7FFC59601" masterDocTitle="Taxonomic revision of the olingos (Bassaricyon), with description of a new species, the Olinguito" masterLastPageNumber="83" masterPageNumber="1" pageNumber="19" updateTime="1668156320301" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the olingos (Bassaricyon), with description of a new species, the Olinguito</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Helgen, Kristofer M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pinto, C. Miguel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kays, Roland</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Helgen, Lauren E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tsuchiya, Mirian T. N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Quinn, Aleta</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wilson, Don E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maldonado, Jesus E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>324</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.324.5827</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.324.5827</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-324-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152047246" LSID="urn:lsid:zoobank.org:act:3A1FA881-9742-4F31-9079-F778B12D7C9A" httpUri="http://treatment.plazi.org/id/CBFBACD8137B8C41D7C96AF81B5FCE17" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
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<subSubSection pageId="18" pageNumber="19" type="nomenclature">
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<paragraph pageId="18" pageNumber="19">
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<taxonomicName LSID="http://species-id.net/wiki/Bassaricyon_neblina_osborni" class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon neblina" order="Carnivora" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="neblina" subSpecies="osborni">Bassaricyon neblina osborni</taxonomicName>
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<taxonomicNameLabel pageId="18" pageNumber="19">subsp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="19" type="diagnosis">
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<paragraph pageId="18" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="18" pageNumber="19">
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This is the largest subspecies of
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon neblina" order="Carnivora" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="neblina">Bassaricyon neblina</taxonomicName>
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, with a short rostrum, widely splayed zygomata, wide rostrum and braincase, and very large molars and posterior premolars; the dorsal pelage is of moderate length, tan to orangish-brown in overall color, with prominent black and gold tipping, with a more grayish face and limbs, with the limbs bearing relatively short fur, and a tail usually grizzled with golden-brown fur tipping.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="distribution">
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<paragraph pageId="19" pageNumber="20">
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<pageBreakToken pageId="19" pageNumber="20" start="start">Distribution</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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This is the representative of
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon neblina" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="neblina">Bassaricyon neblina</taxonomicName>
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on the eastern slopes of the Western Andes of Colombia (e.g., Castilla Mountains [AMNH]; Sabanetas [FMNH]; El Tambo [NMS]; the vicinity of Cali [
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<bibRefCitation author="Poglayen-Neuwall, I" journalOrPublisher="Zoologische Beitraege" pageId="54" pageNumber="55" pagination="179 - 233" title="Fortpflanzung, Geburt und Aufzucht, nebst anderen Beobachtungen von Makibaeren (Bassaricyon Allen, 1876)." volume="22" year="1976">Poglayen-Neuwall 1976</bibRefCitation>
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]; El Duende [
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<bibRefCitation author="Saavedra-Rodriguez, CA" journalOrPublisher="Check List" pageId="55" pageNumber="56" pagination="505 - 507" title="Bassaricyon gabbii Allen, 1876 (Carnivora: Procyonidae): new distribution point on western range of Colombian Andes." volume="7" year="2011">
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<normalizedToken originalValue="Saavedra-Rodríguez">Saavedra-Rodriguez</normalizedToken>
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and Velandia-Perilla 2011
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</bibRefCitation>
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]; Gallera: "western slope of most eastern ridge of southern Western Andes" [AMNH,
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<bibRefCitation pageId="19" pageNumber="20">Paynter 1997:222</bibRefCitation>
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]) and the western slopes of the Central Andes of Colombia (Cerro Munchique [FMNH]). One specimen (AMNH 42351, from Santa Elena, Antioquia Department) derives from the eastern slopes of the Central Andes in northern Colombia (habitat described as "deforested, grassy, and bushy (
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<bibRefCitation author="Chapman, FM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="45" pageNumber="46" pagination="1 - 729" title="The distribution of bird-life in Colombia: a contribution to a biological survey of South America." volume="36" year="1917">Chapman 1917:61</bibRefCitation>
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)";
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<bibRefCitation pageId="19" pageNumber="20">Paynter 1997:403</bibRefCitation>
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); this shows that this subspecies also crosses to the eastern slopes of the Central Andes in Antioquia. Further south, in the department of Huila, the smaller subspecies
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon neblina subsp. hershkovitzi" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="neblina" subSpecies="hershkovitzi">Bassaricyon neblina hershkovitzi</taxonomicName>
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subsp. n. (see below) occurs on the eastern slopes of the Central Andes.
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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Records to date of
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon neblina subsp. osborni" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="neblina" subSpecies="osborni">Bassaricyon neblina osborni</taxonomicName>
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are from 1500 to at least 2750 m elevation in Cauca, Valle del Cauca, and Antioquia Departments of Colombia (Figure 16).
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon medius subsp. medius" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="medius" subSpecies="medius">Bassaricyon medius medius</taxonomicName>
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is also recorded from the Cauca Valley (east slopes of Western Andes and western slopes of Central Andes) at elevations up to at least 725 m (UV-3774:
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<bibRefCitation author="Saavedra-Rodriguez, CA" journalOrPublisher="Check List" pageId="55" pageNumber="56" pagination="505 - 507" title="Bassaricyon gabbii Allen, 1876 (Carnivora: Procyonidae): new distribution point on western range of Colombian Andes." volume="7" year="2011">
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<normalizedToken originalValue="Saavedra-Rodríguez">Saavedra-Rodriguez</normalizedToken>
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and Velandia-Perilla 2011
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</bibRefCitation>
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; see account of
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon medius" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="medius">Bassaricyon medius</taxonomicName>
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below), so these two taxa (
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon medius subsp. medius" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="medius" subSpecies="medius">Bassaricyon medius medius</taxonomicName>
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and
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon neblina subsp. osborni" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="neblina" subSpecies="osborni">Bassaricyon neblina osborni</taxonomicName>
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) are presumably regionally sympatric (and probably elevationally stratified) across the range of this Olinguito subspecies on the slopes of the Western and Central Andes.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="etymology">
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<paragraph pageId="19" pageNumber="20">Etymology.</paragraph>
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<paragraph pageId="19" pageNumber="20">
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The name honors Henry Fairfield Osborn (1857-1935), paleontologist, faculty of Princeton and Columbia Universities, and Curator of Vertebrate Paleontology (1891-1909) and President (1909-1933) of the American Museum of Natural History (
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<bibRefCitation author="Gregory, WK" journalOrPublisher="Biographical Memoirs of the National Academy of Sciences of the United States of America" pageId="49" pageNumber="50" pagination="53 - 119" title="Biographical memoir of Henry Fairfield Osborn, 1857 - 1935." volume="19" year="1937">Gregory 1937</bibRefCitation>
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,
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<bibRefCitation pageId="19" pageNumber="20">Colbert 1996</bibRefCitation>
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). "
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon osborni" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="osborni">Bassaricyon osborni</taxonomicName>
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" is a manuscript name (never formally published) associated with a specimen of this taxon (AMNH 32609, with
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<normalizedToken originalValue="“Type”">"Type"</normalizedToken>
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written on the skull), demonstrating a century-old intention, later discarded (probably by J.A. Allen or H.E. Anthony, see below), to name this taxon after Osborn. Here we validate this unpublished name as a newly described subspecies of
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<taxonomicName class="Mammalia" family="Procyonidae" genus="Bassaricyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bassaricyon neblina" order="Carnivora" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="neblina">Bassaricyon neblina</taxonomicName>
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, but we choose a more complete specimen than AMNH 32609, which has a damaged mandible and various broken teeth, as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="holotype">
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<paragraph pageId="19" pageNumber="20">Holotype.</paragraph>
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<paragraph pageId="19" pageNumber="20">FMNH 88476, adult male, skin and skull, Munchique, 2000 m, Cauca Department, Colombia (collected by K. von Sneidern, 3 June 1957).</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="paratypes">
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<paragraph pageId="19" pageNumber="20">Paratypes.</paragraph>
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<paragraph pageId="19" pageNumber="20">
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AMNH 32608, adult female, skin and skull, and AMNH 32609, adult male, skin and skull, Gallera (
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<bibRefCitation author="Chapman, FM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="45" pageNumber="46" pagination="139 - 166" title="Diagnoses of apparently new Colombian birds." volume="31" year="1912">Chapman 1912:155</bibRefCitation>
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; = "La Gallera" of
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<bibRefCitation pageId="19" pageNumber="20">Paynter 1997:222</bibRefCitation>
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), 5000 feet (=1524 m), Cauca Department, Colombia (both collected by L. Miller, 13 July 1911); NMS A59-5083, adult female, skin and skull, El Tambo, 1700 m, Cauca Department, Colombia (collected by K. von Sneidern); FMNH 85818, adult male, skin and skull, Munchique, 2000 m, Cauca Department, Colombia (collected by K. von Sneidern, 19 January 1956); FMNH 89220, adult female, skin and skull, Sabanetas, 2000 m, Cauca Department, Colombia (collected by K. von Sneidern, 26 September 1957); FMNH 90052, adult female, skin and skull, Sabanetas, 1900 m, Cauca Department, Colombia (collected by K. von Sneidern, 12 February 1959).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="referred specimens">
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<paragraph pageId="20" pageNumber="21">
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<pageBreakToken pageId="20" pageNumber="21" start="start">Referred</pageBreakToken>
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specimens.
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</paragraph>
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<paragraph pageId="20" pageNumber="21">
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AMNH 14185, skin (skull not found), adult male, Castilla Mountains ("La Castilla" of
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<bibRefCitation pageId="20" pageNumber="21">Paynter 1997</bibRefCitation>
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), Valle del Cauca Department (collected by J.H. Batty, 9 June 1898); AMNH 42351, adult male, skin and skull, Santa Elena, apparently at 9000 feet (= 2750 m), Antioquia Department, Colombia (collected by H. Niceforo Maria, 10 January 1919) (
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<bibRefCitation pageId="20" pageNumber="21">Paynter 1997:403</bibRefCitation>
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); USNM 598996, adult male, skin, skull, and postcranial skeleton, from Colombia, specific locality unknown (received from Tulane University).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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