142 lines
15 KiB
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142 lines
15 KiB
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<document id="6C32F883973CB24054F17B1D6BE49AF3" ID-DOI="10.5281/zenodo.208497" ID-GBIF-Dataset="2334b3de-88fe-4fa0-a3ba-63b04b1b4df7" ID-ISSN="1175-5326" ID-Zenodo-Dep="208497" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460471856943" checkinUser="plazi" docAuthor="Heiss, Ernst & Poinar, George O." docDate="2012" docId="F56B3675FFC1FF9EFF78E283FBCDFC95" docLanguage="en" docName="zt03268p054.pdf" docOrigin="Zootaxa 3268" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Carventus bechlyi Heiss & Poinar, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="09524E0DFFC2FF9AFFEFE63EFFA2FFC9" masterDocTitle="The first Carventinae species in Miocene Dominican Amber (Hemiptera: Heteroptera: Aradidae)" masterLastPageNumber="54" masterPageNumber="47" pageNumber="50" updateTime="1698306026984" updateUser="plazi">
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<mods:titleInfo id="A7F1DA0014CAAFEFAA9E5F81426E779C">
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<mods:title id="B304508F0E2FFE5EA44206956E573E0E">The first Carventinae species in Miocene Dominican Amber (Hemiptera: Heteroptera: Aradidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C4AD6D871DEA1F6863B19135642A097E" type="personal">
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<mods:namePart id="C83F2E1AE4189701BDBC4E194771891E">Heiss, Ernst</mods:namePart>
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</mods:name>
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<mods:name id="35A2A54BFE5049BB40BCA83BD4946814" type="personal">
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</mods:role>
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<mods:namePart id="8E14711B1F54D117C8383C4C0E1850F4">Poinar, George O.</mods:namePart>
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<mods:typeOfResource id="40B01551768E0043D3980550446D9B4A">text</mods:typeOfResource>
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<mods:titleInfo id="D4E754E6D4D2889EEC518EA5D0D5CD31">
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<mods:title id="FD434ADBCD9E5B587A40C7A0D8FF8A36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0C59DBC2C4B049E6AB4DFAB071EA7DC4">
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<mods:date id="200B2B58DC6B6E4E41592F7134FE621B">2012</mods:date>
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<mods:detail id="681C32920BFC7FDC7E7B65D01F16D100" type="volume">
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<mods:number id="5CCCE63E0623E80A656F6E73E04884DF">3268</mods:number>
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<mods:classification id="48C92E983CEDED2350EF58FD5D844C7E">journal article</mods:classification>
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<mods:identifier id="7F95553BD15D5AB2D041C902FBF36350" type="DOI">10.5281/zenodo.208497</mods:identifier>
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<mods:identifier id="88C3A38E605C4AAAC91482764ED3BFEB" type="GBIF-Dataset">2334b3de-88fe-4fa0-a3ba-63b04b1b4df7</mods:identifier>
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<mods:identifier id="FF75FEA624FB6A4EFB93BAB21ABB8E0F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2E44677E934B1D6C58F25C4338C6517E" type="Zenodo-Dep">208497</mods:identifier>
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</mods:mods>
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<treatment id="F56B3675FFC1FF9EFF78E283FBCDFC95" ID-DOI="http://doi.org/10.5281/zenodo.6178532" ID-GBIF-Taxon="119515235" ID-Zenodo-Dep="6178532" LSID="urn:lsid:plazi:treatment:F56B3675FFC1FF9EFF78E283FBCDFC95" httpUri="http://treatment.plazi.org/id/F56B3675FFC1FF9EFF78E283FBCDFC95" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
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<subSubSection id="35D8D4E8FFC1FF99FF78E283FE22FB3E" pageId="3" pageNumber="50" type="nomenclature">
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<paragraph id="7D7D8763FFC1FF99FF78E283FDA6FB1F" blockId="3.[151,516,1212,1271]" box="[151,516,1212,1239]" pageId="3" pageNumber="50">
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<heading id="2635300FFFC1FF99FF78E283FDA6FB1F" bold="true" box="[151,516,1212,1239]" fontSize="11" level="1" pageId="3" pageNumber="50" reason="1">
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<emphasis id="4FB65B71FFC1FF99FF78E283FDA6FB1F" bold="true" box="[151,516,1212,1239]" pageId="3" pageNumber="50">
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<taxonomicName id="BAC2FCE0FFC1FF99FF78E283FED2FB1E" box="[151,368,1213,1239]" class="Insecta" family="Aradidae" genus="Carventus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="bechlyi" status="sp. nov.">
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<emphasis id="4FB65B71FFC1FF99FF78E283FED2FB1E" bold="true" box="[151,368,1213,1239]" italics="true" pageId="3" pageNumber="50">Carventus bechlyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5485E60AFFC1FF99FE98E282FE18FB1F" box="[375,442,1212,1238]" pageId="3" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
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Heiss
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7D7D8763FFC1FF99FF78E2E1FE22FB3E" blockId="3.[151,516,1212,1271]" box="[151,384,1247,1271]" pageId="3" pageNumber="50">
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(Photo 3,4,10,
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<figureCitation id="E5F99BE6FFC1FF99FED2E2E1FED5FB3E" box="[317,375,1247,1271]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,684,707]" captionTargetBox="[350,1211,215,642]" captionTargetId="figure@3.[334,1234,193,655]" captionTargetPageId="3" captionText="FIGURES 1 – 3 Reconstruction of head and pronotum. 1 — Neoproxius latocanus n. sp.; 2 — Carventus bechlyi n. sp.; 3 — Proneoproxius cornutus n. gen., n. sp." httpUri="https://zenodo.org/record/208498/files/figure.png" pageId="3" pageNumber="50">Fig.2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="35D8D4E8FFC1FF99FF78E316FDC2FA06" pageId="3" pageNumber="50" type="materials_examined">
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<paragraph id="7D7D8763FFC1FF99FF78E316FDC2FA06" blockId="3.[151,1436,1319,1992]" pageId="3" pageNumber="50">
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<emphasis id="4FB65B71FFC1FF99FF78E316FEAFFA89" bold="true" box="[151,269,1320,1344]" pageId="3" pageNumber="50">
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<typeStatus id="A27939C1FFC1FF99FF78E316FEAAFA89" box="[151,264,1320,1344]" pageId="3" pageNumber="50" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Macropterous male in a brownish transparent piece of cabochon shaped Dominican amber (15x
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<date id="097CA1A3FFC1FF99FF27E375FF47FAAA" box="[200,229,1355,1379]" pageId="3" pageNumber="50">9x</date>
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<quantity id="BA3A2A86FFC1FF99FF0AE375FE86FAAA" box="[229,292,1355,1379]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="50" unit="mm" value="5.0">5mm</quantity>
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); view of dorsal surface is partly obscured by the round surface of the stone, the head at middle by a piece of detritus, the ventral side is clearly visible; antennal segments II–IV cut and lacking on both sides. This specimen is designated as
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<typeStatus id="A27939C1FFC1FF99FE52E3ADFDBDFA62" box="[445,543,1427,1451]" pageId="3" pageNumber="50" type="holotype">holotype</typeStatus>
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and deposited in the collection of the Staatliches Museum für Naturkunde, Stuttgart,
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<collectingCountry id="05D5C7F3FFC1FF99FF37E389FEE2FA06" box="[216,320,1463,1487]" name="Germany" pageId="3" pageNumber="50">Germany</collectingCountry>
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(
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<collectionCode id="1BD31FA6FFC1FF99FEA0E389FE3CFA06" box="[335,414,1463,1487]" pageId="3" pageNumber="50">SMSG</collectionCode>
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) no. Do-4940-H.
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</paragraph>
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</subSubSection>
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<subSubSection id="35D8D4E8FFC1FF99FF28E3E5FD96F996" pageId="3" pageNumber="50" type="diagnosis">
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<paragraph id="7D7D8763FFC1FF99FF28E3E5FD96F996" blockId="3.[151,1436,1319,1992]" pageId="3" pageNumber="50">
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<emphasis id="4FB65B71FFC1FF99FF28E3E5FEE3FA3D" bold="true" box="[199,321,1499,1524]" pageId="3" pageNumber="50">Diagnosis.</emphasis>
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This fossil species resembles the extant
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<taxonomicName id="BAC2FCE0FFC1FF99FCFBE3E2FC0FFA3D" box="[788,941,1500,1524]" class="Insecta" family="Aradidae" genus="Carventus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="mexicanus">
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<emphasis id="4FB65B71FFC1FF99FCFBE3E2FC0FFA3D" box="[788,941,1500,1524]" italics="true" pageId="3" pageNumber="50">C. mexicanus</emphasis>
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</taxonomicName>
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and shares the ventral position of spiracles II–IV ventral and V–VIII lateral, which are visible from above. However, it is distinguished by a smaller size (
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<quantity id="BA3A2A86FFC1FF99FA8CE3C1FF57F9F2" metricMagnitude="-3" metricUnit="m" metricValue="4.025" metricValueMax="4.45" metricValueMin="3.6" pageId="3" pageNumber="50" unit="mm" value="4.025" valueMax="4.45" valueMin="3.6">3.6 – 4.45mm</quantity>
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) and by the different structure of the pronotum with lateral and anterolateral margins not strongly concave and with reduced anterolateral lobes.
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</paragraph>
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</subSubSection>
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<subSubSection id="35D8D4E8FFC1FF9EFF28E055FECFFD26" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="description">
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<paragraph id="7D7D8763FFC1FF99FF28E055FC80F8DA" blockId="3.[151,1436,1319,1992]" pageId="3" pageNumber="50">
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<emphasis id="4FB65B71FFC1FF99FF28E055FEFBF94D" bold="true" box="[199,345,1643,1668]" pageId="3" pageNumber="50">Description.</emphasis>
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Head. Wider across postocular lobes than long (26/24); genae flattened, contiguous and enlarged anteriorly, longer than clypeus but shorter than antennal segment I; antenniferous lobes with blunt apices; antennal segment I club-shaped, the following ones missing; eyes oval inserted in head; postocular lobes triangularly produced laterally, their posterior margin converging straight toward constricted collar; rostrum arising from a slit-like opening, shorter than head, rostral groove flat but distinct.
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</paragraph>
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<paragraph id="7D7D8763FFC1FF99FF28E11EFC1FF801" blockId="3.[151,1436,1319,1992]" pageId="3" pageNumber="50">
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<emphasis id="4FB65B71FFC1FF99FF28E11EFEE2F8F1" bold="true" box="[199,320,1824,1848]" pageId="3" pageNumber="50">Pronotum</emphasis>
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. Twice as wide as long (40/20), lateral margins slightly concave at middle, anterolateral angles nearly rectangular, anterolateral margins converging to concave anterior margin; anterior lobe of disk with large sublateral incrustate elevations depressed at middle with a lower transverse ridge posteriorly; posterior lobe of disk separated from anterior one by a transverse depression followed by a raised transverse carina connecting the elevated humeri; posterior margin incised and convex at base of scutellum.
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</paragraph>
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<paragraph id="7D7D8763FFC6FF9EFF28E6A9FBC8FF1D" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="51">
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<emphasis id="4FB65B71FFC6FF9EFF28E6A9FE9DFF79" bold="true" box="[199,319,151,176]" pageId="4" pageNumber="51">Scutellum</emphasis>
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. Triangular, twice as long as wide (25/12), lateral margins sinuate, apex narrowly rounded, surface with 2 (1+1) sublateral curved incrustations with a triangular depression between them.
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</paragraph>
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<paragraph id="7D7D8763FFC6FF9EFF28E6DEFF41FEF6" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="51">
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<emphasis id="4FB65B71FFC6FF9EFF28E6DEFEE4FF31" bold="true" box="[199,326,224,248]" pageId="4" pageNumber="51">Hemelytra</emphasis>
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. Corium shorter than scutellum, its anterolateral margins expanded and carinate; membrane fully developed covering of tergite VII, surface with distinct longitudinal veins anteriorly, irregularly wrinkled elsewhere.
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</paragraph>
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<paragraph id="7D7D8763FFC6FF9EFF28E775FED3FE06" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="51">
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<emphasis id="4FB65B71FFC6FF9EFF28E775FE9EFEAD" bold="true" box="[199,316,331,356]" pageId="4" pageNumber="51">Abdomen</emphasis>
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. Lateral margins with slightly produced posterolateral angles on deltg III–V, these angularly produced on deltg VI and VII; deltg II+III fused to a triangular sclerite, its surface with a longitudinal ridge; deltg III–VII separated by distinct sutures; tergal plate consisting of mtg III–VI laterally delimited from deltg II–VII by a longitudinal carina.
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</paragraph>
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<paragraph id="7D7D8763FFC6FF9EFF28E7E2FD45FD96" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="51">
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<emphasis id="4FB65B71FFC6FF9EFF28E7E2FEB6FE3D" bold="true" box="[199,276,476,500]" pageId="4" pageNumber="51">Venter</emphasis>
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. Prosternum flat with 2 (1+1) ovate smooth depressions anterolaterally; meso-and metasternum flat at middle, separated from each other and from prosternum and fused sternites II+III by a transverse suture; sternites III–VII with 2:2:1 pattern of apodemal impressions (glabrous spots sensu
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<bibRefCitation id="1953FA92FFC6FF9EFC1DE41AFA81FDF5" author="Usinger" box="[1010,1315,548,572]" pageId="4" pageNumber="51" refString="Usinger, R. L. & Matsuda, R. (1959) Classification of the Aradidae. London, The British Museum, 410 pp." type="book" year="1959">Usinger & Matsuda, 1959</bibRefCitation>
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); spiracles II–IV ventral, V–VIII lateral and visible from above.
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</paragraph>
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<paragraph id="7D7D8763FFC6FF9EFF28E452FBFCFD6E" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="51">
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<emphasis id="4FB65B71FFC6FF9EFF28E452FF5DFD4D" bold="true" box="[199,255,620,644]" pageId="4" pageNumber="51">Legs</emphasis>
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. Femora thickened at middle tapering toward base and apex, fused to trochanters but fusion line distinct; tibiae cylindrical, slightly curved basally, tarsi bi-segmented, claws with large pulvilli.
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</paragraph>
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<paragraph id="7D7D8763FFC6FF9EFF28E48AFECFFD26" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="51">
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<emphasis id="4FB65B71FFC6FF9EFF28E48AFED4FD05" bold="true" box="[199,374,692,716]" pageId="4" pageNumber="51">Measurements</emphasis>
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. Length
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<quantity id="BA3A2A86FFC6FF9EFE34E48AFD97FD05" box="[475,565,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="51" unit="mm" value="3.6">3.6 mm</quantity>
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; width of abdomen across tergite IV
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<quantity id="BA3A2A86FFC6FF9EFC39E48AFB85FD05" box="[982,1063,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="51" unit="mm" value="1.4">1.4mm</quantity>
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, across tergite V
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<quantity id="BA3A2A86FFC6FF9EFB04E48AFAE9FD05" box="[1259,1355,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="4" pageNumber="51" unit="mm" value="1.42">1.42mm</quantity>
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, across tergite VI
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<quantity id="BA3A2A86FFC6FF9EFEE6E4E9FEC8FD26" box="[265,362,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="51" unit="mm" value="1.35">1.35mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="35D8D4E8FFC6FF9EFF28E4C2FBCDFC95" pageId="4" pageNumber="51" type="etymology">
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<paragraph id="7D7D8763FFC6FF9EFF28E4C2FBCDFC95" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="51">
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<emphasis id="4FB65B71FFC6FF9EFF28E4C2FEE4FCDD" bold="true" box="[199,326,764,788]" pageId="4" pageNumber="51">Etymology</emphasis>
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. It is a pleasure to dedicate this species to Dr. Günter Bechly, scientific researcher and head of the Amber section in the Department of Palaeontology at the Staatliches Museum für Naturkunde in Stuttgart,
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<collectingCountry id="05D5C7F3FFC6FF9EFA85E521FF75FC94" name="Germany" pageId="4" pageNumber="51">Germany</collectingCountry>
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, who made available this and other Dominican Amber aradid inclusions for study.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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